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A life cycle database for parasitic acanthocephalans, cestodes, and nematodes

Parasitologists have worked out many complex life cycles over the last ~150 years, yet there have been few efforts to synthesize this information to facilitate comparisons among taxa. Most existing host-parasite databases focus on particular host taxa, do not distinguish final from intermediate hosts, and lack parasite life-history information. We summarized the known life cycles of trophically transmitted parasitic acanthocephalans, cestodes, and nematodes. For 973 parasite species, we gathered information from the literature on the hosts infected at each stage of the parasite life cycle (8510 host-parasite species associations), what parasite stage is in each host, and whether parasites need to infect certain hosts to complete the life cycle. We also collected life-history data for these parasites at each life cycle stage, including 2313 development time measurements and 7660 body size measurements. The result is the most comprehensive data summary available for these parasite taxa. In addition to identifying gaps in our knowledge of parasite life cycles, these data can be used to test hypotheses about life cycle evolution, host specificity, parasite life-history strategies, and the roles of parasites in food webs.

Ecology↗

Variation in the vital rates of an Antarctic marine predator: the role of individual heterogeneity

Variation in life‐history traits such as lifespan and lifetime reproductive output is thought to arise, in part, due to among‐individual differences in the underlying probabilities of survival and reproduction. However, the stochastic nature of demographic processes can also generate considerable variation in fitness‐related traits among otherwise‐identical individuals. An improved understanding of life‐history evolution and population dynamics therefore depends on evaluating the relative role of each of these processes. Here, we used a 33‐yr data set with reproductive histories for 1,274 female Weddell seals from Erebus Bay, Antarctica, to assess the strength of evidence for among‐individual heterogeneity in the probabilities of survival and reproduction, while accounting for multiple other sources of variation in vital rates. Our analysis used recent advances in Bayesian model selection techniques and diagnostics to directly compare model fit and predictive power between models that included individual effects on survival and reproduction to those that did not. We found strong evidence for costs of reproduction to both survival and future reproduction, with breeders having rates of survival and subsequent reproduction that were 3% and 6% lower than rates for non‐breeders. We detected age‐related changes in the rates of survival and reproduction, but the patterns differed for the two rates. Survival rates steadily declined from 0.92 at age 7 to 0.56 at the maximal age of 31 yr. In contrast, reproductive rates increased from 0.68 at age 7 to 0.79 at age 16 and then steadily declined to 0.37 for the oldest females. Models that included individual effects explained more variation in observed life histories and had better estimated predictive power than those that did not, indicating their importance in understanding sources of variation among individuals in life‐history traits. We found that among‐individual heterogeneity in survival was small relative to that for reproduction. Our study, which found patterns of variation in vital rates that are consistent with a series of predictions from life‐history theory, is the first to provide a thorough assessment of variation in important vital rates for a long‐lived, high‐latitude marine mammal while taking full advantage of recent developments in model evaluation.

Ecology↗

Southern California and range‐wide raccoon gastrointestinal helminth database

Local and global measurements of parasite prevalence and abundance are critical for understanding the dynamics that underlie the diversity, distribution, and evolution of infectious diseases. Here, we present a data set of gut helminths found in (1) raccoons throughout their range, based on primary literature from 1925–2017 and (2) raccoons in Santa Barbara County, California, USA surveyed from 2012 to 2015. The range‐wide data set has 1,256 parasite entries from 217 literature sources across three continents and 32 states in the USA. This data set includes a list of all recorded raccoon gut helminths (n = 100) and their presence and prevalence in surveyed raccoon populations. The Santa Barbara data set includes gut helminth data from 182 raccoons from one Southern California County. In addition to the presence and abundance data for 13 parasite species, this data set includes measurements of 7,465 individual raccoon roundworms (Baylisascaris procyonis). For both range‐wide and Santa Barbara data sets, we include information on parasite site of infection in host, sampling method, and sample size. We also provide geographic coordinates for infected raccoon populations (range‐wide database) and individuals (Santa Barbara). In the associated metadata, we include sampling methods and summary figures for both the range‐wide and Santa Barbara raccoon gut helminth records. There are no copyright or proprietary restrictions for research and/or teaching purposes. S. B. Weinstein and J. C. Van Wert contributed equally to this manuscript and are shared first authors.

California↗

Spatiotemporal causal inference with mechanistic ecological models: Evaluating targeted culling on chronic wasting disease dynamics in cervids

Spatiotemporal causal inference methods are needed to detect the effect of interventions on indirectly measured epidemiological outcomes that go beyond studying spatiotemporal correlations. Chronic wasting disease (CWD) causes neurological degeneration and eventual death to white-tailed deer ( Odocoileus virginianus ) in Wisconsin. Targeted culling involves removing deer after traditional hunting seasons in areas with high CWD prevalence. The evaluation of the causal effects of targeted culling in the spread and growth of CWD is an important unresolved research and CWD management question that can guide surveillance efforts. Reaction–diffusion partial differential equations (PDEs) can be used to mechanistically model the underlying spatiotemporal dynamics of wildlife diseases, like CWD, allowing researchers to make inference about unobserved epidemiological quantities. These models indirectly regress spatiotemporal covariates on diffusion and growth rates parameterizing such PDEs, obtaining associational conclusions. In this work we develop an innovative method to obtain causal estimators for the effect of targeted culling interventions on CWD epidemiological processes using an inverse-probability-of-treatment-weighted technique by means of marginal structural models embedded in the PDE fitting process. Additionally we establish a novel scheme for sensitivity analysis under unmeasured confounder for testing the hypothesis of a significant causal effect in the indirectly measured epidemiological outcomes. Our methods can be broadly used to study the impact of spatiotemporal interventions and treatment exposures in the epidemiological evolution of infectious diseases that can help to inform future efforts to mitigate public health implications and wildlife disease burden.

Wisconsin↗

Estimation of regional material yield from coastal landslides based on historical digital terrain modelling

High-resolution historical (1942) and recent (1994) digital terrain models were derived from aerial photographs along the Big Sur coastline in central California to measure the long-term volume of material that enters the nearshore environment. During the 52-year measurement time period, an average of 21 000 ?? 3100 m3 km-1 a-1 of material was eroded from nine study sections distributed along the coast, with a low yield of 1000 ?? 240 m3 km-1 a-1 and a high of 46 700 ?? 7300 m3 km-1 a-1. The results compare well with known volumes from several deep-seated landslides in the area and suggest that the processes by which material is delivered to the coast are episodic in nature. In addition, a number of parameters are investigated to determine what influences the substantial variation in yield along the coast. It is found that the magnitude of regional coastal landslide sediment yield is primarily related to the physical strength of the slope-forming material. Coastal Highway 1 runs along the lower portion of the slope along this stretch of coastline, and winter storms frequently damage the highway. The California Department of Transportation is responsible for maintaining this scenic highway while minimizing the impacts to the coastal ecosystems that are part of the Monterey Bay National Marine Sanctuary. This study provides environmental managers with critical background data on the volumes of material that historically enter the nearshore from landslides, as well as demonstrating the application of deriving historical digital terrain data to model landscape evolution. Published in 2005 by John Wiley & Sons, Ltd.

Earth Surface Processes and Landforms↗

The interactions between vegetation and erosion: new directions for research at the interface of ecology and geomorphology

Vegetation and processes of erosion and deposition are interactive. An objective of this paper is to review selected studies that emphasize the interdependencies. The reviews suggest new directions for research uniting ecology and geomorphology – the sub‐discipline of biogeomorphology. The research, which recently has become vigorous, includes the sources, movement, and fates of fluvial loads of sediment, organic carbon, nutrients, contaminants, and woody debris to low‐energy storage sites; the function of biota in causing soil evolution, stability, and sequestration of carbon; the development of new methods to characterize watersheds based on edaphic conditions; and the refinement of current empirical and conceptual models and dendrochronological techniques to measure landscape change. These well acknowledged topics and others less well anticipated ensure that biogeomorphology will remain vibrant.

Earth Surface Processes and Landforms↗

Multitemporal ALSM change detection, sediment delivery, and process mapping at an active earthflow

Remote mapping and measurement of surface processes at high spatial resolution is among the frontiers in Earth surface process research. Remote measurements that allow meter-scale mapping of landforms and quantification of landscape change can revolutionize the study of landscape evolution on human timescales. At Mill Gulch in northern California, USA, an active earthflow was surveyed in 2003 and 2007 by airborne laser swath mapping (ALSM), enabling meter-scale quantification of landscape change. We calculate four-year volumetric flux from the earthflow and compare it to long-term catchment average erosion rates from cosmogenic radionuclide inventories from adjacent watersheds. We also present detailed maps of changing features on the earthflow, from which we can derive velocity estimates and infer dominant process. These measurements rely on proper digital elevation model (DEM) generation and a simple surface-matching technique to align the multitemporal data in a manner that eliminates systematic error in either dataset. The mean surface elevation of the earthflow and an opposite slope that was directly influenced by the earthflow decreased 14 ± 1 mm/yr from 2003 to 2007. By making the conservative assumption that these features were the dominant contributor of sediment flux from the entire Mill Gulch drainage basin during this time interval, we calculate a minimum catchment-averaged erosion rate of 0·30 ± 0·02 mm/yr. Analysis of beryllium-10 ( 10 Be) concentrations in fluvial sand from nearby Russian Gulch and the South Fork Gualala River provide catchment averaged erosion rates of 0·21 ± 0·04 and 0·23 ± 0·03 mm/yr respectively. From translated landscape features, we can infer surface velocities ranging from 0·5 m/yr in the wide upper ‘source’ portion of the flow to 5 m/yr in the narrow middle ‘transport’ portion of the flow. This study re-affirms the importance of mass wasting processes in the sediment budgets of uplifting weak lithologies.

California↗

Initial hydrologic and geomorphic response following a wildfire in the Colorado front range

A wildfire in May 1996 burned 4690 hectares in two watersheds forested by ponderosa pine and Douglas fir in a steep, mountainous landscape with a summer, convective thunderstorm precipitation regime. The wildfire lowered the erosion threshold in the watersheds, and consequently amplified the subsequent erosional response to shorter time interval episodic rainfall and created both erosional and depositional features in a complex pattern throughout the watersheds. The initial response during the first four years was an increase in runoff and erosion rates followed by decreases toward pre-fire rates. The maximum unit-area peak discharge was 24 m3 s-1 km-2 for a rainstorm in 1996 with a rain intensity of 90 mm h-1. Recovery to pre-fire conditions seems to have occurred by 2000 because for a maximum 30-min rainfall intensity of 50 mm h-1, the unit-area peak discharge in 1997 was 6.6 m3 s-1 km-2, while in 2000 a similar intensity produced only 0.11 m3 s-1 km-2. Rill erosion accounted for 6 per cent, interrill erosion for 14 per cent, and drainage erosion for 80 per cent of the initial erosion in 1996. This represents about a 200-fold increase in erosion rates on hillslopes which had a recovery or relaxation time of about three years. About 67 per cent of the initially eroded sediment is still stored in the watersheds after four years with an estimated residence time greater than 300 years. This residence time is much greater than the fire recurrence interval so erosional and depositional features may become legacies from the wildfire and may affect landscape evolution by acting as a new set of initial conditions for subsequent wildfire and flood sequences.

Colorado↗

Relations between rainfall–runoff-induced erosion and aeolian deposition at archaeological sites in a semi-arid dam-controlled river corridor

Process dynamics in fluvial-based dryland environments are highly complex with fluvial, aeolian, and alluvial processes all contributing to landscape change. When anthropogenic activities such as dam-building affect fluvial processes, the complexity in local response can be further increased by flood- and sediment-limiting flows. Understanding these complexities is key to predicting landscape behavior in drylands and has important scientific and management implications, including for studies related to paleoclimatology, landscape ecology evolution, and archaeological site context and preservation. Here we use multi-temporal LiDAR surveys, local weather data, and geomorphological observations to identify trends in site change throughout the 446-km-long semi-arid Colorado River corridor in Grand Canyon, Arizona, USA, where archaeological site degradation related to the effects of upstream dam operation is a concern. Using several site case studies, we show the range of landscape responses that might be expected from concomitant occurrence of dam-controlled fluvial sand bar deposition, aeolian sand transport, and rainfall-induced erosion. Empirical rainfall-erosion threshold analyses coupled with a numerical rainfall–runoff–soil erosion model indicate that infiltration-excess overland flow and gullying govern large-scale (centimeter- to decimeter-scale) landscape changes, but that aeolian deposition can in some cases mitigate gully erosion. Whereas threshold analyses identify the normalized rainfall intensity (defined as the ratio of rainfall intensity to hydraulic conductivity) as the primary factor governing hydrologic-driven erosion, assessment of false positives and false negatives in the dataset highlight topographic slope as the next most important parameter governing site response. Analysis of 4+ years of high resolution (four-minute) weather data and 75+ years of low resolution (daily) climate records indicates that dryland erosion is dependent on short-term, storm-driven rainfall intensity rather than cumulative rainfall, and that erosion can occur outside of wet seasons and even wet years. These results can apply to other similar semi-arid landscapes where process complexity may not be fully understood.

Arizona↗

Three-dimensional flow structure and patterns of bed shear stress in an evolving compound meander bend

Compound meander bends with multiple lobes of maximum curvature are common in actively evolving lowland rivers. Interaction among spatial patterns of mean flow, turbulence, bed morphology, bank failures and channel migration in compound bends is poorly understood. In this paper, acoustic Doppler current profiler (ADCP) measurements of the three-dimensional (3D) flow velocities in a compound bend are examined to evaluate the influence of channel curvature and hydrologic variability on the structure of flow within the bend. Flow structure at various flow stages is related to changes in bed morphology over the study timeframe. Increases in local curvature within the upstream lobe of the bend reduce outer bank velocities at morphologically significant flows, creating a region that protects the bank from high momentum flow and high bed shear stresses. The dimensionless radius of curvature in the upstream lobe is one-third less than that of the downstream lobe, with average bank erosion rates less than half of the erosion rates for the downstream lobe. Higher bank erosion rates within the downstream lobe correspond to the shift in a core of high velocity and bed shear stresses toward the outer bank as flow moves through the two lobes. These erosion patterns provide a mechanism for continued migration of the downstream lobe in the near future. Bed material size distributions within the bend correspond to spatial patterns of bed shear stress magnitudes, indicating that bed material sorting within the bend is governed by bed shear stress. Results suggest that patterns of flow, sediment entrainment, and planform evolution in compound meander bends are more complex than in simple meander bends. Moreover, interactions among local influences on the flow, such as woody debris, local topographic steering, and locally high curvature, tend to cause compound bends to evolve toward increasing planform complexity over time rather than stable configurations.

Earth Surface Processes and Landforms↗

Parsing anomalous versus normal diffusive behavior of bedload sediment particles

Bedload sediment transport is the basic physical ingredient of river evolution. Formulae exist for estimating transport rates, but the diffusive contribution to the sediment flux, and the associated spreading rate of tracer particles, are not clearly understood. The start-and-stop motions of sediment particles transported as bedload on a streambed mimic aspects of the Einstein–Smoluchowski description of the random-walk motions of Brownian particles. Using this touchstone description, recent work suggests the presence of anomalous diffusion, where the particle spreading rate differs from the linear dependence with time of Brownian behavior. We demonstrate that conventional measures of particle spreading reveal different attributes of bedload particle behavior depending on details of the calculation. When we view particle motions over start-and-stop timescales obtained from high-speed (250 Hz) imaging of coarse-sand particles, high-resolution measurements reveal ballistic-like behavior at the shortest (10 −2 s) timescale, followed by apparent anomalous behavior due to correlated random walks in transition to normal diffusion (>10 −1 s) – similar to Brownian particle behavior but involving distinctly different physics. However, when treated as a ‘virtual plume’ over this timescale range, particles exhibit inhomogeneous diffusive behavior because both the mean and the variance of particle travel distances increase nonlinearly with increasing travel times, a behavior that is unrelated to anomalous diffusion or to Brownian-like behavior. Our results indicate that care is needed in suggesting anomalous behavior when appealing to conventional measures of diffusion formulated for ideal particle systems.

Earth Surface Processes and Landforms↗

Strong variation in weathering of layered rock maintains hillslope‐scale strength under high precipitation

The evolution of volcanic landscapes and their landslide potential are both dependent upon the weathering of layered volcanic rock sequences. We characterize critical zone structure using shallow seismic V p and V s profiles and vertical exposures of rock across a basaltic climosequence on Kohala peninsula, Hawai’i, and exploit the dramatic gradient in mean annual precipitation (MAP) across the peninsula as a proxy for weathering intensity. Seismic velocity increases rapidly with depth and the velocity–depth gradient is uniform across three sites with 500–600 mm/yr MAP, where the transition to unaltered bedrock occurs at a depth of 4 to 10 m. In contrast, velocity increases with depth less rapidly at wetter sites, but this gradient remains constant across increasing MAP from 1000 to 3000 mm/yr and the transition to unaltered bedrock is near the maximum depth of investigation (15–25 m). In detail, the profiles of seismic velocity and of weathering at wet sites are nowhere monotonic functions of depth. The uniform average velocity gradient and the greater depths of low velocities may be explained by the averaging of velocities over intercalated highly weathered sites with less weathered layers at sites where MAP > 1000 mm/yr. Hence, the main effect of climate is not the progressive deepening of a near‐surface altered layer, but rather the rapid weathering of high permeability zones within rock subjected to precipitation greater than ~1000 mm/yr. Although weathering suggests mechanical weakening, the nearly horizontal orientation of alternating weathered and unweathered horizons with respect to topography also plays a role in the slope stability of these heterogeneous rock masses. We speculate that where steep, rapidly evolving hillslopes exist, the sub‐horizontal orientation of weak/strong horizons allows such sites to remain nearly as strong as their less weathered counterparts at drier sites, as is exemplified by the 50°–60° slopes maintained in the amphitheater canyons on the northwest flank of the island.

Hawai'i↗

Factors controlling landslide frequency-area distributions

A power‐law relation for the frequency–area distribution (FAD) of medium and large landslides (e.g. tens to millions of square meters) has been observed by numerous authors. But the FAD of small landslides diverges from the power‐law distribution, with a rollover point below which frequencies decrease for smaller landslides. Some studies conclude that this divergence is an artifact of unmapped small landslides due to lack of spatial or temporal resolution; others posit that it is caused by the change in the underlying failure process. An explanation for this dilemma is essential both to evaluate the factors controlling FADs of landslides and power‐law scaling, which is a crucial factor regarding both landscape evolution and landslide hazard assessment. This study examines the FADs of 45 earthquake‐induced landslide inventories from around the world in the context of the proposed explanations. We show that each inventory probably involves some combination of the proposed explanations, though not all explanations contribute to each case. We propose an alternative explanation to understand the reason for the divergence from a power‐law. We suggest that the geometry of a landslide at the time of mapping reflects not just one single movement but many, including the propagation of numerous smaller landslides before and after the main failure. Because only the resulting combination of these landslides can be observed due to a lack of temporal resolution, many smaller landslides are not taken into account in the inventory. This reveals that the divergence from the power‐law is not necessarily attributed to the incompleteness of an inventory. This conceptual model will need to be validated by ongoing observation and analysis. Also, we show that because of the subjectivity of mapping procedures, the total number of landslides and total landslide areas in inventories differ significantly, and therefore the shapes of FADs also differ considerably.

Earth Surface Processes and Landforms↗

Six years of fluvial response to a large dam removal on the Carmel River, California, USA

Measuring river response to dam removal affords a rare, important opportunity to study fluvial response to sediment pulses on a large field scale. We present a before–after/control–impact study of the Carmel River, California, measuring fluvial geomorphic and grain-size evolution over 8 years, six of which postdated removal of a 32 m-high dam (one of the largest dams removed worldwide) and included 11 flow events exceeding the 2-year flood magnitude. We find that the reservoir-sediment pulse following dam removal was relatively small (97 000 ± 24 000 t over 4 years), owing to deliberate reservoir-sediment stabilization. Scaled to the size of the Carmel River watershed and compared against long-term bedrock denudation rates, the post-dam-removal sediment release was slightly less than the annualized long-term sediment export from this basin. New sediment transited >30 km to the river mouth in less than 2 years, assisted by floods 2 and 4 years after dam removal. The sediment pulse fined the downstream riverbed while causing mostly low-magnitude bed-elevation changes: commonly 0.5 to 1 m or smaller, occurring as discontinuous sediment patches or interstitial deposits, aside from the filling and subsequent partial scour of deep pools. There was no major geomorphic reset downstream from the dam site. Geomorphic changes were driven almost entirely by flow rather than by the modest increase in sediment supply, in contrast to recent examples from other large dam removals. The relatively minor disturbance caused by dam removal on the Carmel River is likely analogous to many future dam removals: a relatively small sediment pulse after deliberate limitation of reservoir-sediment erosion, and with an upstream dam remaining in place. Thus, a large dam removal need not lead to major downstream impacts.

California↗

Fluviomorphic trajectories for dryland ephemeral stream channels following extreme flash floods

Ephemeral alluvial streams pose globally significant flood hazards to human habitation in drylands, but sparse data for these regions limit understanding of the character and impacts of extreme flooding. In this study, we document decadal changes in dryland ephemeral channel patterns at two sites in the lower Colorado River Basin (southwestern United States) that were ravaged by extraordinary flash floods in the 1970s: Bronco Creek, Arizona (1971), and Eldorado Canyon, Nevada (1974). We refer to these two floods as ‘fluviomorphic erasure events’, because they produced blank slates for the channels that were gradually moulded by more frequent but much smaller flood events. We studied georectified aerial photos that span ~60 years at each site to show that both study sites recovered to their pre-flood condition after ~25 years. We employ channel network metrics: stream-link area (SLA), geometric braiding index and junction-node density. Each metric decreased during the short-duration extreme flood erasure events. Subsequently, a fluviomorphic trajectory at a decadal tempo returned the channels to pre-flood values. The SLA decreased at rates of 3.6%–4.1% per year in the decade following the floods. The extreme flood events decreased the pre-flood geometric braiding index at the two sites by 56%–68%, and it took 15–24 years for this index to recover to pre-flood values. In contrast, it took 30–35 years for the channels to recover to a uniform pre-flood channel form, as indicated by the spatial distribution of bars and junction nodes. Our results document baseline examples of ephemeral stream channel evolution trajectories, as future climatic change will likely accelerate increases in the magnitudes and frequencies of extreme floods and geomorphic erasure events.

Earth Surface Processes and Landforms↗

The contribution of a surge event to infilling in a barrier-enclosed estuary: Insights from field observations

Many estuaries worldwide face increasing sediment loading caused by catchment land use change and intensification, creating subsequent adverse effects on estuarine ecosystems. Extreme weather events can disproportionately alter sediment pathways and loading. Although storm-driven sediment exchange has been widely examined at open coasts and inlets, key transport mechanisms within constricted, sheltered estuaries remain understudied. This study presents an observational dataset capturing the impact of a 99th percentile water-level event (based on 20 years of records) on sediment transport pathways in a sheltered, barrier-enclosed estuary. This event, driven by a 3-day storm surge (>0.5 m) combined with a spring tide, was recorded during a 3-week field campaign. Sediment transport pathways and riverine contributions were analysed, and observations revealed substantial changes in suspended sediment concentrations increasing from 18 mg/l to 70 mg/l during the event. The elevated water levels and resulting pressure gradient at the constricted study site entrance caused by the storm surge increased local flood dominance. Combined with higher flow velocities and resuspension, the storm led to a sixfold increase in sediment import at the estuary entrance and a 600-fold increase in sediment flux to the upper estuary. The decoupling of peak suspended sediment concentrations from streamflow indicates that the resuspension of estuarine legacy sediment, rather than catchment inputs, dominated the system's response. These findings challenge assumptions about estuarine sediment budgets and emphasise that incorporating high water-level surge events into models can enhance the prediction of long-term estuarine evolution. Given projected increases in storm frequency under climate change, understanding these episodic but highly consequential sediment pulses can support the assessment of wetland resilience and inform estuarine management strategies.

Whangateau Estuary↗

Wildfires as an ecosystem service

Wildfires are often viewed as destructive disturbances. We propose that when including both evolutionary and socioecological scales, most ecosystem fires can be understood as natural processes that provide a variety of benefits to humankind. Wildfires provide open habitats that enable the evolution of a diversity of shade-intolerant plants and animals that have long been used by humans. There are many provisioning, regulating, and cultural services that we obtain from wildfires; prescribed fires and wildfire management are tools for mimicking the ancestral role of wildfires in a highly populated world.

Frontiers in Ecology and the Environment↗

Geoarchaeology of stratified paleoindian deposits at the Big Eddy site, Southwest Missouri, U.S.A

The Big Eddy site (23CE426) in the Sac River valley of southwest Missouri is a rare recorded example of distinctly stratified Early through Late Paleoindian cultural deposits. Early point types recovered from the site include Gainey, Sedgwick, Dalton (fluted and unfluted), San Patrice, Wilson, and Packard. The Paleoindian record at Big Eddy represents only a fraction of the site's prehistoric cultural record; stratified cultural deposits in alluvium above the Paleoindian components span the entire known prehistoric sequence, and terminal Pleistocene alluvium may contain pre-Early Paleoindian cultural deposits. This study focused on the paleogeomorphic setting, stratigraphy, depositional environments, pedology, geochronology, and history of landscape evolution of the late Pleistocene and early Holocene alluvium at the site. The Paleoindian sequence is associated with a complex buried soil 2.85 m below the modern surface (T1a) of the first terrace of the Sac River valley in the site vicinity. This soil formed at the top of the early submember of the Rodgers Shelter Member (underlying the T1c paleogeomorphic surface) and contains at least 70 cm of stratified Paleoindian cultural deposits, all in floodplain and upper point-bar facies. A suite of 36 radiocarbon ages indicates that the alluvium hosting the Paleoindian sequence aggraded between ca. 13,250 and 11,870 cal yr B.P. (11,380 and 10,180 14C yr B.P.). Underlying deposits accumulated between ca. 15,300 and 13,250 cal yr B.P. (12,950 and 11,380 14C yr B.P.). By ca. 11,250 cal yr B.P. (9,840 14C yr B.P.) the T1c paleogeomorphic surface was buried by the earliest increment of a thick sequence of overbank sheetflood facies, ultimately resulting in deep burial and preservation of the Paleoindian record. The landform-sediment assemblage that hosts the Paleoindian and possibly earlier cultural deposits at Big Eddy is both widespread and well preserved in the lower Sac River valley. Moreover, the terminal Pleistocene and early Holocene depositional environments were favorable for the preservation of the archaeological record. ?? 2007 Wiley Periodicals, Inc.

Geoarchaeology↗