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R.J. Cooper

Publications and source records attributed to R.J. Cooper.

32 records · Page 2Linked to original sources

Population and habitat assessment: Monitoring bird populations over large areas: Introduction

Monitoring provides essential information about status and change in bird populations. For Neotropical Migrant Birds (NTMBs), the North American Breeding Bird Survey (BBS) has been particularly influential in documenting regional population change and often is cited as justification for management actions. However, as with most bird surveys, the design of the BBS, and the geographic scale of the information, often limits its use either in evaluating the response of bird populations to management, or in identifying causes of population change.

Book chapter↗

Does nonrandom nest placement imply nonrandom nest predation?: A reply

In response to the critique by Schmidt and Whelan (Condor 101(4):916-920, 1999), we find that the relationship between nest success and tree selectivity is dependent upon inclusion or exclusion of particular tree species, whether or not years are pooled, and the selectivity index used. We question their use of point estimates of nest success with extremely high variances, defend our index, question the application of the Chesson (1983) index to our data, and explain the need to analyze years separately. Bottomland hardwood forest systems are extremely variable; hydroperiods alter the suitability of nesting substrates, availability of alternative food, and behavior of predators and their prey. Given these features, actively searching for Acadian Flycatcher (Empidonax virescens) nests is seldom an efficient predator foraging strategy. Therefore, these predation events are best described as random; nests are principally encountered opportunistically by generalist predators while searching for other prey.

Condor↗

Acadian flycatcher nest placement: Does placement influence reproductive success?

We located 511 Acadian Flycatcher (Empidonax virescens) nests in bottomland hardwood forest of eastern Arkansas. Microhabitat characteristics were measured and their relationship with nest success evaluated. Fifty-two percent of all nesting attempts resulted in predation. Attributes of nest placement were similar between successful and unsuccessful nests, although successful nests were placed higher. Similarly, nonparasitized nests were typically higher than parasitized nests. Nests initiated late in the breeding season were placed in larger trees with higher canopy bases resulting in increased vegetation around the nest. Fifteen different tree species were used for nesting. Acadian Flycatchers chose nest trees in a nonrandom fashion, selecting Nuttall oak (Quercus nuttallii) and possumhaw (Ilex decidua) in greater proportions than their availability. However, there was no relationship between tree species used for nesting and nest success. Nest height was positively correlated with concealment at the nest site, supporting the predator-avoidance theory. No other attribute of nest placement differentiated successful nest sites, suggesting that nest predation is likely a function of random events in space and time.

Condor↗

The seventy-second Christmas bird count. 302. Southern Dorchester County, Md

We located 511 Acadian Flycatcher (Empidonax virescens) nests in bottomland hardwood forest of eastern Arkansas. Microhabitat characteristics were measured and their relationship with nest success evaluated. Fifty-two percent of all nesting attempts resulted in predation. Attributes of nest placement were similar between successful and unsuccessful nests, although successful nests were placed higher. Similarly, nonparasitized nests were typically higher than parasitized nests. Nests initiated late in the breeding season were placed in larger trees with higher canopy bases resulting in increased vegetation around the nest. Fifteen different tree species were used for nesting. Acadian Flycatchers chose nest trees in a nonrandom fashion, selecting Nuttall oak (Quercus nuttallii) and possumhaw (Ilex decidua) in greater proportions than their availability. However, there was no relationship between tree species used for nesting and nest success. Nest height was positively correlated with concealment at the nest site, supporting the predator-avoidance theory. No other attribute of nest placement differentiated successful nest sites, suggesting that nest predation is likely a function of random events in space and time.

Condor↗

Breeding biology of Acadian flycatchers in a bottomland hardwood forest

From 1993-1995, we located and monitored 601 Acadian Flycatcher (Empidonax virescens) nests in a large contiguous tract of bottomland hardwood forest on the White River National Wildlife Refuge, Arkansas. Annual reproductive success was significantly different among years; ranging from 10-25% (Mayfield estimate) over the three years of the study. There was no significant difference in nest success among study plots, with nesting success showing a trend of increasing late in the breeding season. Clutch size for non-parasitized nests averaged 2.9 ? 0.02 (SE) eggs with a mode of 3. Rates of Brown-headed Cowbird (Molothrus ater) parasitism were low (21%), accounting for 7% of all nest failures. However, parasitism by cowbirds resulted in a reduction of clutch size for nests initiated early (i.e., first nests and replacements) in the breeding season. Predation was the leading cause of nest failures, accounting for 75% of all failures. Snakes and avian predators were thought to be the leading cause of nest failures. Although additional factors must be investigated, preliminary results indicate that nest predation is a major influence on this population, despite the size of the forest tract.

The Wilson Bulletin↗

Increasing point-count duration increases standard error

We examined data from point counts of varying duration in bottomland forests of west Tennessee and the Mississippi Alluvial Valley to determine if counting interval influenced sampling efficiency. Estimates of standard error increased as point count duration increased both for cumulative number of individuals and species in both locations. Although point counts appear to yield data with standard errors proportional to means, a square root transformation of the data may stabilize the variance. Using long (>10 min) point counts may reduce sample size and increase sampling error, both of which diminish statistical power and thereby the ability to detect meaningful changes in avian populations.

Journal of Field Ornithology↗

A land manager's guide to point counts of birds in the Southeast

Current widespread concern for the status of neotropical migratory birds has sparked interest in techniques for inventorying and monitoring populations of these and other birds in southeastern forest habitats. The present guide gives detailed instructions for conducting point counts of birds. It further presents a detailed methodology for the design and conduct of inventorial and monitoring surveys based on point counts, including discussion of sample size determination, distribution of counts among habitats, cooperation among neighboring land managers, vegetation sampling, standard data format, and other topics. Appendices provide additional information, making this guide a stand-alone text for managers interested in developing inventories of bird populations on their lands.

General Technical Report↗

Sample size and allocation of effort in point count sampling of birds in bottomland hardwood forests

To examine sample size requirements and optimum allocation of effort in point count sampling of bottomland hardwood forests, we computed minimum sample sizes from variation recorded during 82 point counts (May 7-May 16, 1992) from three localities containing three habitat types across three regions of the Mississippi Alluvial Valley (MAV). Also, we estimated the effect of increasing the number of points or visits by comparing results of 150 four-minute point counts obtained from each of four stands on Delta Experimental Forest (DEF) during May 8-May 21, 1991 and May 30-June 12, 1992. For each stand, we obtained bootstrap estimates of mean cumulative number of species each year from all possible combinations of six points and six visits. ANOVA was used to model cumulative species as a function of number of points visited, number of visits to each point, and interaction of points and visits. There was significant variation in numbers of birds and species between regions and localities (nested within region); neither habitat, nor the interaction between region and habitat, was significant. For a = 0.05 and a = 0.10, minimum sample size estimates (per factor level) varied by orders of magnitude depending upon the observed or specified range of desired detectable difference. For observed regional variation, 20 and 40 point counts were required to accommodate variability in total individuals (MSE = 9.28) and species (MSE = 3.79), respectively, whereas ? 25 percent of the mean could be achieved with five counts per factor level. Sample size sufficient to detect actual differences of Wood Thrush (Hylocichla mustelina) was >200, whereas the Prothonotary Warbler (Protonotaria citrea) required <10 counts. Differences in mean cumulative species were detected among number of points visited and among number of visits to a point. In the lower MAV, mean cumulative species increased with each added point through five points and with each additional visit through four visits. Although no interaction was detected between number of points and number of visits, when paired reciprocals were compared, more points invariably yielded a significantly greater cumulative number of species than more visits to a point. Still, 36 point counts per stand during each of two breeding seasons detected only 52 percent of the known available species pool in DEF.

Book chapter↗

Sampling bird communities in bottomland hardwood forests of the Mississippi Alluvial Valley: Number of points visited versus number of visits to a point

Within each of 4 forest stands on Delta Experimental Forest (DEF), 25 points were visited 5 to 7 times from 8 May to 21 May 1991, and 6 times from 30 May to 12 June 1992. During each visit to a point, all birds detected, visuallyor aurally, at any distance were recorded during a 4-minute interval. Using these data, our objectives were to recommend the number of point counts and the number of visits to a point which provide the greatest efficiency for estimating the cumulative number of species in bottomland hardwood forest stands within the Mississippi Alluvial Valley, and to ascertain if increasing the number of visits to points is equivalent to adding more points. Because the total number of species detected in DEF were different between years, 39 species in 1991 and 55 species in 1992, we considered each year independently. Within each stand, we obtained bootstrap estimates of the mean cumulative number of species obtained from all possible combinations of six points and six visits (i.e., 36 means/stand). These bootstrap estimates were subjected to ANOVA; we modelled cumulative number of species as a function of the number of points visited, the number of visits to each point, and their interaction. As part of the same ANOVA we made an a priori, simultaneous comparison of the 15 possible reciprocal treatments (i.e., 1 point-2 visits vs. 2 points-1 visit, etc.). Results of analyses for each year were similar. Although no interaction was detected between the number of points and the number of visits, when reciprocals were compared, more points visited yielded significantly greater cumulative number of species than more visits to each point. Significant differences were detected among both the number of points visited and among the number of visits to a point. Scheffe's test of differences among means indicated that the cumulative number of species increased significantly with each added point, through five points, but six points did not differ from five points in 1991. Similarly, the cumulative number of species increased significantlywith each revisit, up to four visits, but four visits did not differ significantly from five visits. Starting with one point, which yielded about 33 percent of the total species pool when averaged among one through six points, each subsequent point resulted in an increase of about 9 percent, 5 percent, 3 percent, and 3 percent, respectively. Each sequential increase in the number of visits, however, only resulted in increases of 7 percent, 4 percent, 2 percent, and 2 percent of the total species pool.

Book chapter↗

Point counts of landbirds in bottomland hardwood forests of the Mississippi Alluvial Valley: How long and how many?

To quantify efficacy of point count sampling in bottomland hardwood forests, we examined the influence of point count duration on corresponding estimates of number of individuals and species recorded. To accomplish this we conducted a totalof 82 point counts 7 May-16 May 1992distributed among three habitats (Wet, Mesic, Dry) in each of three regions within the lower Mississippi Alluvial Valley (MAV). Each point count consisted of recording the number of individual birds (all species) seen or heard during the initial three minutes and per each minute thereafter for a period totaling ten minutes. In addition, we included 384 point counts recorded during an 8-week period in each of 3 years (1985-1987) among 56 randomly-selected forest patches within the bottomlands of western Tennessee. Each point count consisted of recording the number of individuals (excluding migrating species) during each of four, 5 minute intervals for a period totaling 20 minutes. To estimate minimum sample size, we determined sampling variation at each level (region, habitat, and locality) with the 82 point counts from the lower (MAV) and applied the procedures of Neter and Wasserman (1974:493; Applied linear statistical models). Neither the cumulative number of individuals nor number of species per sampling interval attained an asymptote after 10 or 20 minutes of sampling. For western Tennessee bottomlands, total individual and species counts relative to point count duration were similar among years and comparable to the pattern observed throughout the lower MAV. Across the MAV, we recorded a total of 1,62 1 birds distributed among 52 species with the majority (8721/1621) representing 8 species. More birds were recorded within 25-50 m than in either of the other distance categories. There was significant variation in numbers of individuals and species among point counts. For both, significant differences between region and patch (nested within region) occurred; neither habitat nor interaction between habitat and region was significant. For = 0.05 and L3 = 0.10, minimum sample size estimates (per factor level) varied by orders of magnitude depending upon the observed or specified range of desired detectable difference. For observed regional variation, 20 and 40 point counts were required to accommodate variability in total birds (MSE = 9.28) and species (MSE = 3.79), respectively; 25 percent of the mean could be achieved with 5 counts per factor level. Corresponding sample sizes required to detect differences of rarer species (e.g., Wood Thrush) were 500; for common species (e.g., Northern Cardinal) this same level of precision could be achieved with 100 counts.

Book chapter↗

Point counts of birds in bottomland hardwood forests of the Mississippi Alluvial Valley: Duration, minimal sample size, and points versus visits

The purpose of the research was to evaluate the efficacy of point count surveys in bottomland hardwood forests. The specific objectives were to determine: (1) whether the recommended distance measures for point count circular plots are useful in bottomland hardwood forests, (2) the optimum duration for each point count, (3) the optimum number of points to sample at each locality, (4) the optimum number of counts at each point during a season, and (5) the minimum sample size to accommodate the variation in bird species distribution and relative abundance throughout the lower MAV.

Research Paper↗