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Lightning fires in North Dakota grasslands and in pine-savanna lands of South Dakota and Montana

Lightning strike fires which occurred between 1940 and 1981 were studied in mixed-grass prairie grasslands and in pine-savanna lands in the Northern Great Plains region. A majority (73%) of ignitions occurred during July and August, while a lesser number was recorded in April, May, June, and September. The April-September period is also the average time of the freeze-free period and approximates the average distribution period for thunderstorm activity in this region. The area burned by each of 293 lightning fires (most of which were suppressed) ranged from 0.004-1158.3 ha ( X̄ = 10.8 ha). The frequency of lightning fires in mixed-grass prairie grasslands averaged 6.0/yr per 10,000 km 2 in eastern North Dakota, 22.4/yr per 10,000 km 2 in southcentral North Dakota, 24.7/yr per 10,000 km 2 in western North Dakota, and 91.7/yr per 10,000 km 2 in pine-savanna lands in northwestern South Dakota and southeastern Montana. The ecological role of lightning-set fires is discussed relative to the development of resource research and management plans and to the interpretation of historical records of natural fire occurrence in the Northern Great Plains region.

Journal of Range Management

Nesting habitat selection by sage grouse in south-central Washington

To characterize western sage grouse ( Centrocercus urophasianus phaios Bonaparte) nesting habitat in sagebrush-steppe habitat in Washington, we initiated a study on the Yakima Training Center to determine nesting habitat characteristics and whether these characteristics differed between successful and depredated nests. Most nests (71%) were in big sagebrush ( Artemesia tridentata Nutt.)/bunchgrass communities. Nest habitat was characterized by greater shrub cover, shrub height, vertical cover height, residual cover, and litter than at random locations. Successful 1-m 2 nest sites within big sagebrush/bunchgrass in 1992 had less shrub cover (51%) and shrub height (64 cm) than depredated nest sites (70% and 90 cm, respectively). Successful 77-m 2 nest areas in big sagebrush/bunchgrass in 1993 had more tall grass ( ≥ 18 cm) than depredated nest areas. Management that protects the big sagebrush/bunchgrass community is essential for maintaining nesting habitat for sage grouse.

Journal of Range Management

Response of mountain meadows to grazing by recreational pack stock

Effects of recreational pack stock grazing on mountain meadows in Yosemite National Park were assessed in a 5-year study. Yosemite is a designated wilderness, to be managed such that its natural conditions are preserved. Studies were conducted in 3 characteristic meadow types: shorthair sedge ( Carex filifolia Nutt.), Brewer's reed grass ( Calamagrostis breweri Thurber), and tufted hairgrass [ Deschampsia cespitosa (L.) Beauv.]. Horses and mules grazed experimental plots at intensities of 15 to 69% utilization for 4 seasons. In all 3 meadows, grazing caused decreases in productivity. The mean reduction after 4 years of grazing was 18% in the shorthair sedge meadow, 17% in the Brewer's reed grass meadow, and 22% in the tufted hairgrass meadow. Grazing also caused shifts in basal groundcover (usually a reduction in vegetation cover and increase in bare soil cover), and changes in species composition. Productivity and vegetation cover decreased as percent utilization increased, while bare soil cover increased as utilization increased. Changes in species composition were less predictably related to differences in grazing intensity. Passive management of grazing is insufficient in wilderness areas that are regularly used by groups with recreational stock. Wilderness managers need to monitor meadow conditions and the grazing intensities that occur. Our study suggests that biomass and ground cover are more sensitive indicators of grazing impact than species composition. Managers must make decisions about maximum acceptable levels of grazing impact and then develop guidelines for maximum use levels, based on data such as ours that relates grazing intensity to meadow response.

Journal of Range Management

Grassland birds associated with agricultural riparian practices in southwestern Wisconsin

Rotational grazing has been proposed as a Best Management Practice (BMP) for minimizing runoff in Wisconsin agricultural riparian areas. The influence of this land management practice on grassland birds has not been evaluated in relation to more traditional agricultural land management systems in Midwestern riparian areas. This study compared the grassland bird community in riparian areas in Wisconsin that were rotationally grazed to 2 common land use practices along streams in Wisconsin: continuously grazed pastures and rowcrop fields with 10-m-wide ungrazed buffer strips located along the stream. We calculated total number of birds, the Berger-Parker Index of Dominance, and number of birds ha-1 for each site. Vegetation variables used were height-density, litter depth, and percent bare ground. Bird species richness, species dominance, and density did not differ among land use types. In contrast, grassland bird species of management concern [Savannah Sparrow ( Passerculus sandwichensis Gmelin), Eastern Meadowlark ( Sturnella magna L.), and Bobolink ( Dolichonyx oryzivorus L.)] were found on continuous and rotational pastures but very rarely or never occurred on buffer strips. Contrary to previous research, however, rotationally grazed pastures did not support more of these species than continuously grazed pastures. Bird density was related to vegetation structure, with higher densities found on sites with deeper litter. Within the pasture land use types, there were no consistent differences between species richness and density near the stream (<10 m) and away (>10 m).

Wisconsin

Rangeland health attributes and indicators for qualitative assessment

Panels of experts from the Society for Range Management and the National Research Council proposed that status of rangeland ecosystems could be ascertained by evaluating an ecological site's potential to conserve soil resources and by a series of indicators for ecosystem processes and site stability. Using these recommendations as a starting point, we developed a rapid, qualitative method for assessing a moment-in-time status of rangelands. Evaluators rate 17 indicators to assess 3 ecosystem attributes (soil and site stability, hydrologic function, and biotic integrity) for a given location. Indicators include rills, water flow patterns, pedestals and terracettes, bare ground, gullies, wind scour and depositional areas, litter movement, soil resistance to erosion, soil surface loss or degradation, plant composition relative to infiltration, soil compaction, plant functional/structural groups, plant mortality, litter amount, annual production, invasive plants, and reproductive capability. In this paper, we detail the development and evolution of the technique and introduce a modified ecological reference worksheet that documents the expected presence and amount of each indicator on the ecological site. In addition, we review the intended applications for this technique and clarify the differences between assessment and monitoring that lead us to recommend this technique be used for moment-in-time assessments and not be used for temporal monitoring of rangeland status. Lastly, we propose a mechanism for adapting and modifying this technique to reflect improvements in understanding of ecosystem processes. We support the need for quantitative measures for monitoring rangeland health and propose some measures that we believe may address some of the 17 indicators.

Journal of Range Management

Changes in plant functional groups, litter quality, and soil carbon and nitrogen mineralization with sheep grazing in an Inner Mongolian Grassland

This study reports on changes in plant functional group composition, litter quality, and soil C and N mineralization dynamics from a 9-year sheep grazing study in Inner Mongolia. Addressed are these questions: 1) How does increasing grazing intensity affect plant community composition? 2) How does increasing grazing intensity alter soil C and N mineralization dynamics? 3) Do changes in soil C and N mineralization dynamics relate to changes in plant community composition via inputs of the quality or quantity of litter? Grazing plots were set up near the Inner Mongolia Grassland Ecosystem Research Station (IMGERS) with 5 grazing intensities: 1.3, 2.7, 4.0, 5.3, and 6.7 sheep ha −1 ·yr −1 . Plant cover was lower with increasing grazing intensity, which was primarily due to a dramatic decline in grasses, Carex duriuscula , and Artemisia frigida . Changes in litter mass and percentage organic C resulted in lower total C in the litter layer at 4.0 and 5.3 sheep ha −1 ·yr −1 compared with 2.7 sheep ha −1 ·yr −1 . Total litter N was lower at 5.3 sheep ha −1 ·yr −1 compared with 2.7 sheep ha −1 ·yr −1 . Litter C:N ratios, an index of litter quality, were significantly lower at 4.0 sheep ha −1 ·yr −1 relative to 1.3 and 5.3 sheep ha −1 ·yr −1 . Cumulative C mineralized after 16 days decreased with increasing grazing intensity. In contrast, net N mineralization ( NH 4 NO − 3 ) after a 12-day incubation increased with increasing grazing intensity. Changes in C and N mineralization resulted in a narrowing of CO 2 -C:net N min ratios with increasing grazing intensity. Grazing explained 31% of the variability in the ratio of CO 2 -C:net N min . The ratio of CO 2 -C:net N min was positively correlated with litter mass. Furthermore, there was a positive correlation between litter mass and A. frigida cover. Results suggest that as grazing intensity increases, microbes become more C limited resulting in decreased microbial growth and demand for N.

Journal of Range Management

Changes in vegetation and grazing capacity following honey mesquite control

Honey mesquite kill and suppression, vegetation response, and changes in grazing use and capacity were evaluated following brush control in north-central Texas. Tree grubbing was most effective for eliminating honey mesquite, but because of soil and plant damage the treatment did not increase grazing capacity or improve range condition compared to nontreated rangeland. Aerial application of 2,4,5-T + picloram was more effective in klllmg and defoliating honey mesquite than 2,4,5-T alone, but both treatments significantly increased forage production. The 2,4,5-T + picloram and 2,4,5-T sprays provided a 7 to 16% increase in grazing capacity over a 4-year period on light and heavy honey mesquite infested pastures, respectively.

Texas

Frequency sampling in microhistological studies: An alternative model

Frequency sampling in microhistological studies is discussed in terms of sampling procedures, statistical properties, and biological inferences. Two sampling approaches are described and contrasted, and some standard methods for improving the stability of density estimators are discussed. Possible sources of difficulty are highlighted in terms of sampling design and statistical analysis. An alternative model is proposed that accounts for 2-stage sampling, and yields reasonable, we!!-behaved estimates of relative densities.

Journal of Range Management

Copper deficiency in Tule Elk at Point Reyes, California

Tule elk ( Cervus elaphus nannodes ) reintroduced to Point Reyes, Calif., in 1978 exhibited gross signs of copper deficiency by June 1979. Copper levels in liver (x=5.9 ppm) and serum (0.42 ppm) of elk in Point Reyes were below levels in adult tule elk from other locations in California (liver, x=80 ppm; serum, x=1.4 ppm). These levels were consistent with documented copper deficiencies in wild and domestic ruminants. Copper serum levels increased in response to copper enriched dietary supplements and declined after the elk stopped eating the supplements. Analysis of plant and soil samples showed both are deficient in copper and normal in molybdenum and sulfur-sulfates. Deficiency in plants and soils at Point Reyes are probably due to low copper levels in the underlying granitic parent material.

California

Prescribed grazing as a secondary impact in a western riparian floodplain

The effect of late-autumn cattle grazing on plant biomass was examined in a western Great Plains cottonwood riparian zone prone to catastrophic flooding every 5-8 years. Following 1 year of pre-treatment data collection in 1982, five 16-ha pastures were grazed from 1982 to 1984 and compared to 5 control pastures within the South Platte River floodplain in northeastern Colorado. At a prescribed grazing level of 0.46 ha/AUM, riparian vegetation proved to be resilient to the impacts of grazing. We detected only a few significant treatment effects for above-ground biomass after succeeding growing seasons. Willows ( Salix spp.) responded negatively to grazing whereas biomass of prairie cordgrass ( Spartina pectinata Link) was greater on grazed plots. Yearly changes in above-ground biomass, especially dramatic following a severe flood in 1983, suggest that periodic, catastrophic flooding is a major perturbation to the ecosystem, and in conjunction with our results on grazing impacts, indicate that dormant-season grazing within Soil Conservation Service (SCS) guidelines is a comparatively minor impact within the floodplain. In addition, grazing impacts were probably further mitigated by a major forage supplement of cottonwood leaves which was available at the time of cattle introductions. This local forage supplement ultimately created a lighter grazing treatment than that originally prescribed.

Journal of Range Management

Comparison of rangeland vegetation sampling techniques in the Central Grasslands

Maintaining native plant diversity, detecting exotic species, and monitoring rare species are becoming important objectives in rangeland conservation. Four rangeland vegetation sampling techniques were compared to see how well they captured local pant diversity. The methods tested included the commonly used Parker transects, Daubenmire transects as modified by the USDA Forest Service, a new transect and 'large quadrat' design proposed by the USDA Agricultural Research Service, and the Modified-Whittaker multi-scale vegetation plot. The 4 methods were superimposed in shortgrass steppe, mixed grass prairie, northern mixed prairie, and tallgrass prairie in the Central Grasslands of the United States with 4 replicates in each prairie type. Analysis of variance tests showed significant method effects and prairie type effects, but no significant method X type interactions for total species richness, the number of native species, the number of species with less than 1 % cover, and the time required for sampling. The methods behaved similarly in each prairie type under a wide variety of grazing regimens. The Parker, large quadrat, and Daubenmire transects significantly underestimated the total species richness and the number of native species in each prairie type, and the number of species with less than 1 % cover in all but the tallgrass prairie type. The transect techniques also consistently missed half the exotic species, including noxious weeds, in each prairie type. The Modified-Whittaker method, which included an exhaustive search for plant species in a 20 x 50 m plot, served as the baseline for species richeness comparisons. For all prairie types, the Modified-Whittaker plot captured an average of 42. ( ± 2.4; 1 S.E.) plant species per site compared to 15.9 ( ± 1.3), 18.9 ( ± 1.2), and 22.8 ( ± 1.6) plant species per site using the Parker, large quadrat, and Daubenmire transect methods, respectively. The 4 methods captured most of the dominant species at each site and thus produced similar results for total foliar cover and soil cover. The detection and measurement of exotic plant species were greatly enhanced by using ten 1 m 2 subplots in a multi-scale sampling design and searching a larger area (1,000 m 2 ) at each site. Even with 4 replicate sites, the transect methods usually captured, and thus would monitor, 36 to 66 % of the plant species at each site. To evaluate the status and trends of common, rare, and exotic plant species at local, regional, and national scales, innovative, multi-scale methods must replace the commonly used transect methods to the past.

Journal of Range Management

Breeding bird response to juniper woodland expansion

In recent times, pinyon ( Pinus spp. )-juniper ( Juniperus spp. ) woodlands have expanded into large portions of the Southwest historically occupied by grassland vegetation. From 1997-1998, we studied responses of breeding birds to one-seed juniper ( J. monosperma ) woodland expansion at 2 grassland study areas in northern Arizona. We sampled breeding birds in 3 successional stages along a grassland-woodland gradient: un-invaded grassland, grassland undergoing early stages of juniper establishment, and developing woodland. Species composition varied greatly among successional stages and was most different between endpoints of the gradient. Ground-nesting grassland species predominated in uninvaded grassland but declined dramatically as tree density increased. Tree- and cavity-nesting species increased with tree density and were most abundant in developing woodland. Restoration of juniper-invaded grasslands will benefit grassland-obligate birds and other wildlife.

Journal of Range Management

Lone star tick abundance, fire, and bison grazing in tall-grass prairie

Lone star ticks ( Amblyomma americanum L.) were collected by drag samples of 1 km transects on 12 watersheds at Konza Prairie Research Natural Area near Manhattan, Kans., during summer 1995-1996. Watersheds were treated to 2 experimental treatments: 3 burn intervals (1-year, 4-year, and 20-year) and 2 grazing treatments (grazed by bison ( Bos bison L.) or ungrazed). The objectives were to determine whether fire interval, time since most recent burn, and the presence of large ungulate grazers would cause changes in lone star tick abundance in tallgrass prairie in central Kansas. Watersheds burned at 1-year intervals had fewer larvae and adults than watersheds burned at 4-year or 20-year intervals. Watersheds burned during the year of sampling had fewer ticks than watersheds burned one or more years in the past. For watersheds burned 1 or more years in the past there was no effect from time since burn. The presence of bison did not affect tick abundance. Spring burning is an effective method to reduce tick populations in tallgrass prairie during the year of the burn.

Journal of Range Management