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Visualizing social-ecological intensities for management of recreation visitors in a multiuse system

Accounting for the variation of visitor conflicts and ecological disturbance of outdoor recreation activities across space and time can cause difficulty for managers seeking to make decisions in social-ecological systems (SESs). We develop a method to quantify and visualize social and ecological intensities resulting from outdoor recreation. We demonstrate the utility of our method at Valentine National Wildlife Refuge, where we conducted onsite surveys for an entire year of recreationists participating in consumptive (i.e., hunting), intermediate-consumptive (i.e., fishing) and nonconsumptive (e.g., hiking) activities. We use survey results and combine them with expert consensus by engaging refuge managers and scientists (i.e., Delphi method) to chart patterns in social (e.g., visitor conflicts) and ecological (e.g., damages to natural resources) intensities across multiple spatial and temporal scales. We highlight unexpected patterns that are revealed by collectively considering multi-activity groups through space and time and combining different survey methods (onsite, Delphi method). Based on the consensus reached using the Delphi method , the consumptive group had the greatest potential for social conflicts and ecological disturbances. Social and ecological intensities (i.e., hotspots) of recreation varied across lake types and seasons, highlighting high-intensity areas and periods on the refuge. Accounting for diverse outdoor recreation activities and coinciding social and ecological intensities will allow managers of SESs the ability to concomitantly preserve ecological resources, prioritize conservation efforts, and minimize visitor conflicts. We demonstrate the utility and ease of use of this technique, which can be implemented by managers and scientists within their respective SES of interest.

Nebraska↗

Does age matter? The influence of age on response rates in a mixed-mode survey

The appeal of cost savings and faster results has fish and wildlife management agencies considering the use of Internet surveys instead of traditional mail surveys to collect information from their constituents. Internet surveys, however, may suffer from differential age-related response rates, potentially producing biased results if certain age groups respond to Internet surveys differently than they do to mail surveys. We examined this concern using data from a mixed-mode angler survey conducted in South Dakota following the 2011 fishing season. Results indicated that young anglers (16–18) had the lowest return rates and senior anglers (65+) had the highest, regardless of survey mode. Despite this consistency in response rates, we note two concerns: (a) lower Internet response rates and (b) different age groups represented by the Internet and mail survey samples differed dramatically. Findings indicate that constituent groups may be represented differently with the use of various survey modes.

Human Dimensions of Wildlife: An International Jou↗

Integrated rangeland fire management strategy actionable science plan completion assessment— Climate and weather topic, 2015–20

Loss and degradation of sagebrush ( Artemisia spp.) rangelands due to an accelerated invasive annual grass-wildfire cycle and other stressors are significant management, conservation, and economic issues in the western U.S. These sagebrush rangelands comprise a unique biome spanning 11 states, support over 350 wildlife species, and provide important ecosystem services that include stabilizing the economies of western communities. Impacts to sagebrush ecosystem processes over large areas due to the annual grass-wildfire cycle necessitated the development of a coordinated, science-based strategy for improving efforts to achieve long-term protection, conservation, and restoration of sagebrush rangelands, which was framed in 2015 under the Integrated Rangeland Fire Management Strategy (IRFMS). Central to this effort was the development of an Actionable Science Plan (Plan) that identified 37 priority science needs (hereinafter, “Needs”) for informing the actions proposed under the five topics (Fire, Invasives, Restoration, Sagebrush and Sage-Grouse ( Centrocercus urophasianus ), Climate and Weather) that were part of the collective focus of the IRFMS. Notable keys to this effort were identification of the Needs co-produced by managers and researchers, and a focus on resulting science being “actionable.” Substantial investments aimed at fulfilling the Needs identified in the Plan have been made since its release in 2016. While the state of the science has advanced considerably, the extent to which knowledge gaps remain relative to identified Needs is relatively unknown. Moreover, new Needs have likely emerged since the original strategy as results from actionable science reveal new questions and possible (yet untested) solutions. A quantifiable assessment of the progress made on the original science Needs can identify unresolved gaps and new information that can help inform prioritization of future research efforts. This report details a systematic literature review that evaluated how well peer-reviewed journal articles and formal technical reports published between January 1, 2015, and December 31, 2020, addressed four Needs identified under the Climate and Weather topic in the Plan. The topic outlined research Needs broadly focused on understanding the potential effects of climate change on vegetative resilience to inform restoration of sagebrush rangelands. We established the level of progress towards addressing each Need following a standardized set of criteria, and developed summaries detailing how research objectives nested within Needs identified in the Plan (hereinafter, “Next Steps”) were either addressed well, partially addressed or remain outstanding (that is, addressed poorly) in the literature through 2020. Our searches resulted in the inclusion of 92 science products that at least partially addressed a Need identified in the Climate and Weather topic. The Needs that were well and partially addressed included: studies of the complex set of climatic relationships that influence sagebrush rangeland restoration and seeding success; the identification of seed collection areas across the range of environmental variability inhabited by target restoration species; and develop predictive models to assess targeted restoration species’ responses to mid-century climatic conditions. The Need addressed poorly was the identification of native plant species, genotypes and ecotypes, and seed mixes that may be resilient to a changing climate. The information provided in this assessment will assist updating the Plan, and can inform updates of other relevant science planning documents as needed.

Open-File Report↗

Using distance sampling with camera traps to estimate densities of ungulates on tropical oceanic islands

Reliable population estimates are one of the most elementary needs for the management of wildlife, particularly for introduced ungulates on oceanic islands. We aimed to produce accurate and precise density estimates of Philippine deer ( Rusa marianna ) and wild pigs ( Sus scrofa ) on Guam using motion-triggered cameras combined with distance sampling to estimate densities from observations of unmarked animals while accounting for imperfect detection. We used an automated digital data processing pipeline for species recognition and to estimate the distance to detected species. Our density estimates were slightly lower than published estimates, consistent with management to reduce populations. We estimated the number of camera traps needed to obtain a 0.1 coefficient of variation was substantial, requiring > ten-fold increase in camera traps, while estimates with precision of 0.2 or 0.3 were more achievable, requiring doubling to quadrupling the number of camera traps. We provide best practices for establishing and conducting distance sampling with camera trap surveys for density estimation based on lessons learned during this study. Future studies should consider distance sampling with camera traps to efficiently survey and monitor unmarked animals, particularly medium-sized ungulates, in tropical, oceanic island ecosystems.

Journal of Tropical Ecology↗

Modeling habitat connectivity to inform reintroductions: a case study with the Chiricahua Leopard Frog

Managing species with intensive tools such as reintroduction may focus on single sites or entire landscapes. For vagile species, long-term persistence will require colonization and establishment in neighboring habitats. Therefore, both suitable colonization sites and suitable dispersal corridors between sites are required. Assessment of landscapes for both requirements can contribute to ranking and selection of reintroduction areas, thereby improving management success. Following eradication of invasive American Bullfrogs ( Lithobates catesbeianus ) from most of Buenos Aires National Wildlife Refuge (BANWR; Arizona, United States), larval Chiricahua Leopard Frogs ( Lithobates chiricahuensis ) from a private pond were reintroduced into three stock ponds. Populations became established at all three reintroduction sites followed by colonization of neighboring ponds in subsequent years. Our aim was to better understand colonization patterns by the federally threatened L. chiricahuensis which could help inform other reintroduction efforts. We assessed the influence of four landscape features on colonization. Using surveys from 2007 and information about the landscape, we developed a habitat connectivity model, based on electrical circuit theory, that identified potential dispersal corridors after explicitly accounting for imperfect detection of frogs. Landscape features provided little insight into why some sites were colonized and others were not, results that are likely because of the uniformity of the BANWR landscape. While corridor modeling may be effective in more-complex landscapes, our results suggest focusing on local habitat will be more useful at BANWR. We also illustrate that existing data, even when limited in spatial or temporal resolution, can provide information useful in formulating management actions.

Journal of Herpetology↗

Importance of riparian forest corridors for the ocelot in agricultural landscapes

Worldwide, private lands have attracted increased attention from conservationists, not only because most of the globe is privately owned, but also because private lands can be an asset to the protected area conservation strategy. In Brazil, the riverine Areas of Permanent Protection (APPs) is a key instrument of the Forest Code to protect native vegetation on private lands. Although APPs were conceived to function as potential wildlife corridors, this putative role has been rarely assessed. Further, recent debatable changes in the Forest Code have decreased the extent of APPs. Given this lack of information and the declining area being protected along with the growing demand for agricultural and biofuel production, which stimulate additional deforestation, it is timely to assess the role of APPs in wildlife conservation. We did this analyzing how ocelot (Leopardus pardalis) occurrence is influenced by covariates in 3 cerrado landscapes dominated by sugarcane and managed forests of eucalyptus and pine trees. We collected detection/non-detection data by camera trapping during 2 dry seasons (2013 and 2014) at 208 stations (6606 camera-days). We estimated ocelot detection and mean relative abundance using a single-species/single-season occupancy model that accounts for heterogeneous detection probability induced by variation in abundance. Modeling results showed that percentage of native forest was the most important covariate to explain ocelot mean relative abundance. This parameter was also affected positively by APPs, with ocelots being more abundant inside than outside APPs and, unexpectedly, by sugarcane. Given study design, however, the positive effect of this crop likely reflects the contact zone between sugarcane and native forest. Our findings show that landscape composition affects ocelot abundance and highlight the importance of APPs and Legal Reserves in agricultural landscapes. We conclude that, in such landscapes, compliance to the Forest Code by private land owners is key to supporting ocelot occurrence.

Journal of Mammalogy↗

Managing plague on prairie dog colonies: Insecticides as ectoparasiticides

Human health practitioners and wildlife biologists use insecticides to manage plague by suppressing fleas (Siphonaptera), but insecticides can also kill other ectoparasites. We investigated effects of deltamethrin and fipronil on ectoparasites from black-tailed prairie dogs ( Cynomys ludovicianus , BTPDs). In late July, 2018, we treated three sites with 0.05% deltamethrin dust and 5 sites with host-fed 0.005% fipronil grain. Three non-treated sites functioned as experimental baselines. We collected ectoparasites before treatments (June-July, 2018) and after treatments (August-October, 2018, June-July, 2019). Both deltamethrin and fipronil suppressed fleas for at least 12 months. Deltamethrin had no detectable effect on mites (Arachnida). Fipronil suppressed mites for at least 12 months. Lice (Phthiraptera) were scarce on non-treated sites throughout the study, complicating interpretation. Concentrating on eight sites where all three ectoparasites where found in June-July, 2018 (before treatments), flea intensity was greatest on BTPDs carrying many lice and mites. These three ectoparasites co-occurred at high numbers, which might facilitate plague transmission in some cases. Lethal effects of insecticides on ectoparasite communities are potentially advantageous in the context of plague management.

South Dakota↗

Bioeconomic analysis of selected conservation practices on soil erosion and freshwater fisheries

Farmers can generate environmental benefits (improved water quality and fisheries and wildlife habitat), but they may not be able to quantify them. Furthermore, farmers may reduce their incomes from managing lands to produce these positive externalities but receive little monetary compensation in return. This study simulated the relationship between agricultural practices, water quality, fish responses to suspended sediment and farm income within two small watersheds, one of a cool water stream and one of a warm water stream. Using the Agricultural Drainage and Pesticide Transport (ADAPT) model, this study related best management practices (BMPs) to calculated instream suspended sediment concentrations by estimating sediment delivery, runoff, base flow, and streambank erosion to quantify the effects of suspended sediment exposure on fish communities. By implementing selected BMPs in each watershed, annual net farm income declined $18,000 to $28,000 (1 to 3 percent) from previous levels. “Lethal” fish events from suspended sediments in the cool water watershed decreased by 60 percent as conservation tillage and riparian buffers increased. Despite reducing suspended sediments by 25 percent, BMPs in the warm water watershed did not reduce the negative response of the fisheries. Differences in responses (physical and biological) between watersheds highlight potential gains in economic efficiency by targeting BMPs or by offering performance based “green payments.”

Journal of the American Water Resources Associatio↗

Compensating for diminishing natural water: Predicting the impacts of water development on summer habitat of desert bighorn sheep

Artificial water sources have been used for decades to enhance and restore wildlife habitat but the benefits of their use have been subject to debate. During the past century, the number of natural springs in Joshua Tree National Park, California, USA, has declined. In response to concerns about the viability of the bighorn sheep (Ovis canadensis nelsoni) population, a number of water developments were constructed throughout the park. We modeled potential historical and present-day summer habitat of female bighorn sheep to evaluate the effectiveness of the artificial and remaining natural water sources in maintaining habitat and to determine how loss of artificial sources might affect future habitat availability. Prior to 1950, 583.5 km2 of summer habitat was potentially available. Presently, only 170.6 km2 of habitat is available around natural water sources and 153.5 km2 is available around guzzlers. When all perennial water sources are included in the habitat model (minus overlap), 302.3 km2 of summer habitat is potentially available. This represents only 51.7% of summer habitat available prior to 1950. Without artificial water developments, 47.7% of present-day summer habitat would be lost, which raises important management questions regarding the debate about what is natural or artificial within otherwise protected areas.

Journal of Arid Environments↗

Why let the dogs out? Exploring variables associated with dog confinement and general characteristics of the free-ranging owned-dog population in a peri-urban area

Free-ranging dogs (FRDs), are a problem in several countries, with impacts on humans, domestic animals, and wildlife, although increasing evidence suggests that most FRDs are owned. Therefore, understanding dog ownership on a fine scale is critical. The main objectives of this study were to explore dog management in rural localities from central Chile focusing on modeling owner-related variables associated with dog confinement and characterize confined and FRDs populations. Interviews (170) were carried out in Paine municipality, reporting a human:dog ratio of 1.5:1, and dogs in most households (85.9%, 146/170). Thirty-seven percent (54/146) of those households did not confine their dog(s) to some degree, and 41% (196/472) of surveyed dogs were FRD. Based on multivariable logistic regression models, non-confinement was decreased by (i) negative opinion of owners toward roaming behavior of their dogs, (ii) negative opinion toward FRDs, among others. Dog confinement increased along with owners' concerns about the impacts of their dogs on others. Owned-FRDs tended to have poorer general care than confined dogs. Our findings represent a contribution to the understanding of the human dimensions behind FRDs and provide critical quantitative elements to consider when planning effective control strategies.

Journal of Applied Animal Welfare Science↗

Trends and natural variability of North American spring onset as evaluated by a new gridded dataset of spring indices

Climate change is expected to modify the timing of seasonal transitions this century, impacting wildlife migrations, ecosystem function, and agricultural activity. Tracking seasonal transitions in a consistent manner across space and through time requires indices that can be used for monitoring and managing biophysical and ecological systems during the coming decades. Here a new gridded dataset of spring indices is described and used to understand interannual, decadal, and secular trends across the coterminous United States. This dataset is derived from daily interpolated meteorological data, and the results are compared with historical station data to ensure the trends and variations are robust. Regional trends in the first leaf index range from 20.8 to 21.6 days decade21, while first bloom index trends are between20.4 and 21.2 for most regions. However, these trends are modulated by interannual to multidecadal variations, which are substantial throughout the regions considered here. These findings emphasize the important role large-scale climate modes of variability play in modulating spring onset on interannual to multidecadal time scales. Finally, there is some potential for successful subseasonal forecasts of spring onset, as indices from most regions are significantly correlated with antecedent large-scale modes of variability.

Journal of Climate↗

Trail resource impacts and an examination of alternative assessment techniques

Trails are a primary recreation resource facility on which recreation activities are performed. They provide safe access to non-roaded areas, support recreational opportunities such as hiking, biking, and wildlife observation, and protect natural resources by concentrating visitor traffic on resistant treads. However, increasing recreational use, coupled with poorly designed and/or maintained trails, has led to a variety of resource impacts. Trail managers require objective information on trails and their conditions to monitor trends, direct trail maintenance efforts, and evaluate the need for visitor management and resource protection actions. This paper reviews trail impacts and different types of trail assessments, including inventory, maintenance, and condition assessment approaches. Two assessment methods, point sampling and problem assessment, are compared empirically from separate assessments of a 15-mile segment of the Appalachian Trail in Great Smoky Mountains National Park. Results indicate that point sampling and problem assessment methods yield distinctly different types of quantitative information. The point sampling method provides more accurate and precise measures of trail characteristics that are continuous or frequent (e.g., tread width or exposed soil). The problem assessment method is a preferred approach for monitoring trail characteristics that can be easily predefined or are infrequent (e.g., excessive width or secondary treads), particularly when information on the location of specific trail impact problems is needed. The advantages and limitations of these two assessment methods are examined in relation to various management and research information needs. The choice and utility of these assessment methods are also discussed.

Journal of Park and Recreation Administration↗

A system to evaluate the scientific quality of biological and restoration objectives using National Wildlife Refuge Comprehensive Conservation Plans as a case study

It is widely accepted that plans for restoration projects should contain specific, measurable, and science-based objectives to guide restoration efforts. The United States Fish and Wildlife Service (USFWS) is in the process of developing Comprehensive Conservation Plans (CCPs) for more than 500 units in the National Wildlife Refuge System (NWRS). These plans contain objectives for biological and ecosystem restoration efforts on the refuges. Based on USFWS policy, a system was developed to evaluate the scientific quality of such objectives based on three critical factors: (1) Is the objective specific, measurable, achievable, results-oriented, and time-fixed? (2) What is the extent of the rationale that explains the assumptions, logic, and reasoning for the objective? (3) How well was available science used in the development of the objective? The evaluation system scores each factor on a scale of 1 (poor) to 4 (excellent) according to detailed criteria. The biological and restoration objectives from CCPs published as of September 2004 (60 total) were evaluated. The overall average score for all biological and restoration objectives was 1.73. Average scores for each factor were: Factor 1-1.97; Factor 2-1.86; Factor 3-1.38. The overall scores increased from 1997 to 2004. Future restoration efforts may benefit by using this evaluation system during the process of plan development, to ensure that biological and restoration objectives are of the highest scientific quality possible prior to the implementation of restoration plans, and to allow for improved monitoring and adaptive management.

Journal for Nature Conservation↗

Panmixia in a sea ice-associated marine mammal: evaluating genetic structure of the Pacific walrus (Odobenus rosmarus divergens) at multiple spatial scales

The kin structure of a species at relatively fine spatial scales impacts broad-scale patterns in genetic structure at the population level. However, kin structure rarely has been elucidated for migratory marine mammals. The Pacific walrus ( Odobenus rosmarus divergens ) exhibits migratory behavior linked to seasonal patterns in sea ice dynamics. Consequently, information on the spatial genetic structure of the subspecies, including kin structure, could aid wildlife managers in designing future studies to evaluate the impacts of sea ice loss on the subspecies. We sampled 8,303 individual walruses over a 5-year period and used 114 single-nucleotide polymorphisms to examine both broad-scale patterns in genetic structure and fine-scale patterns in relatedness. We did not detect any evidence of genetic structure at broad spatial scales, with low F ST values (≤ 0.001) across all pairs of putative aggregations. To evaluate kin structure at fine spatial scales, we defined a walrus group as a cluster of resting individuals that were less than one walrus body length apart. We found weak evidence of kin structure at fine spatial scales, with 3.72% of groups exhibiting mean relatedness values greater than expected by chance, and a significantly higher overall observed mean value of relatedness within groups than expected by chance. Thus, the high spatiotemporal variation in the distribution of resources in the Pacific Arctic environment likely has favored a gregarious social system in Pacific walruses, with unrelated animals forming temporary associations.

Bering Sea↗

An assessment of Idaho's wildlife management areas for the protection of wildlife

Since 1940, Idaho Department of Fish and Game has developed a network of 31 Wildlife Management Areas (WMAs) across the state. This program has been focused mostly on conservation of game species and their habitats. We assessed the contribution of Idaho's WMAs to conservation of all Idaho's wildlife and other aspects of ecological diversity. Predicted occurrences of species' breeding habitats and other data were used to evaluate the representation of wildlife habitat and other ecological conditions. We found 33 of 39 natural land cover types were mapped as occurring in WMAs. WMAs occurred in 10 of 15 of Bailey's ecoregion sections, absent only from two sections that occupy greater than 1% of Idaho. Percent area of WMAs by elevation followed a pattern similar to percent area of Idaho; however, mean elevation for WMAs was lower than for the state and other protected areas in Idaho. We predicted breeding habitat for 98.4% of Idaho's wildlife and all federal and state listed threatened, endangered, or candidate terrestrial vertebrates to occur in at least one WMA. We predicted habitat for 39 species to occur on five or fewer WMAs, and predicted no habitat on WMAs for five species. We found that a system of WMAs established mainly to protect game species potentially conserves many other aspects of Idaho's ecological diversity, may provide habitat for more than 98% of Idaho's wildlife, and complements other protected areas in the state.

Idaho↗

Gray wolf (Canis lupus) predation patterns following recent recolonization in a multi-predator, multi-prey system

Predator–prey interactions are among the most fundamental of ecological relationships. Recolonizing gray wolf ( Canis lupus Linnaeus, 1758) populations present new challenges for wildlife management in multi-prey, multi-carnivore systems. We documented diet composition and kill rates for wolves in a recently recolonized area over winter and summer seasons (2014–2015). Elk ( Cervus canadensis Erxleben, 1777) were the primary ungulate prey (63%) located at wolf kill sites. Deer (mule deer ( Odocoileus hemionus (Rafinesque, 1817)) and white-tailed deer ( Odocoileus virginianus (Zimmermann, 1780))) were less prevalent than elk in wolf diets, but the amount of deer in diets (40%–50%) varied by pack and season. Juvenile elk were the most prevalent class of prey in wolf diets during summer (63.3%) and winter (36.3%), with adult elk (32.5%) observed nearly as often as juveniles in winter. Kill rates varied by season, with rates 2.3 times higher in summer (mean = 3.5 ungulates/week per pack) than winter (mean = 1.5 ungulates/week per pack), consistent with increased availability and use of neonate prey. Prey biomass acquisition did not vary by pack or season (summer = 243 kg/week per pack; winter = 182 kg/week per pack). Our study quantified predation patterns for a recolonizing wolf population, and patterns that we documented were similar to other multi-prey systems in North America.

Oregon↗

Habitat use by tiger prey in Thailand’s Western Forest Complex: What will it take to fill a half-full tiger landscape?

Tiger populations are declining globally, and depletion of major ungulate prey is an important contributing factor. To better understand factors affecting prey distribution in Thailand’s Western Forest Complex (WEFCOM), we conducted sign surveys for gaur (Bos gaurus ), banteng ( Bos javanicus ), and sambar ( Rusa unicolor ) along 3517 1-km transects and used occupancy models to identify important covariates associated with habitat use by each species. Habitat use by both gaur and sambar was lowest in areas closest to human settlements, although sambar preferred lower slopes near streams whereas gaur preferred steeper slopes at higher elevations. Banteng were found in only one of 17 protected areas (Huai Kha Khaeng [HKK] Wildlife Sanctuary), where they used low elevations and low slopes. We used these modeled relationships to predict occurrence of gaur, sambar, and banteng across each square km of the 19,000 km 2 WEFCOM landscape, using > 60 % occupancy probability to define suitable habitat use for each species. Based on this criterion, gaur and sambar occupied 28 and 50 % of suitable habitat in WEFCOM, and banteng occupied 57 % of suitable habitat in HKK. We used our models to assess the effectiveness of two hypothetical conservation initiatives. First, we modeled the impact of decreasing human activities around nine villages in the core of WEFCOM, which increased predicted suitable habitat in WEFCOM to 68 and 75 % for guar and sambar. We also modeled the extent of potential banteng habitat that still remains in the other 16 protected areas. This could result in a 4-fold increase in banteng suitable habitat in WEFCOM. This is the first study to use occupancy surveys to determine where large prey species can be restored to support management to increase the distribution of tigers, and potentially fill a half-full tiger landscape.

Journal for Nature Conservation↗

Daily survival rates for nests of Black Skimmers from a core breeding area of the Southeastern USA

Little is known about the reproductive success of Black Skimmers ( Rynchops niger ) throughout the southeastern USA where availability of undisturbed beaches for nesting is limited. Daily survival rates (DSR) of nests were examined at three nesting sites in Cape Romain National Wildlife Refuge (CRNWR), South Carolina, USA, 2009&ndash;2010. The percent of successful nests ( n = 346 nests) ranged from 42&ndash;69% among colony sites when data were pooled across both years. The DSR of nests was primarily related to colony site, predation risk, height of high tide, and clutch size. Predation and overwash were the principal causes of identifiable nest loss, each accounting for &sim;33% of nest failures during the two study years. Because of the challenges of resighting skimmer chicks, we were not able to measure chick survival effectively and therefore accurate measures of productivity remain elusive. High variability in nest success among sites within close proximity to each other (<20 km) suggests factors at local scales such as disturbance, predation, and overwash events strongly influenced nest success of Black Skimmers during these 2 years as opposed to more region-wide stressors such as tropical storms or food availability. Although time-intensive techniques to control predators do exist, management options to limit flooding and overwash are far more limited. Conservation of Black Skimmers in the southeastern USA would benefit from coordinated, multi-state efforts to measure nest and chick survival.

South Carolina↗