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Habitat selection of Rocky Mountain elk in a nonforested environment

Recent expansions by Rocky Mountain elk (Cervus elaphus) into nonforested habitats across the Intermountain West have required managers to reconsider the traditional paradigms of forage and cover as they relate to managing elk and their habitats. We examined seasonal habitat selection patterns of a hunted elk population in a nonforested high-desert region of southwestern Wyoming, USA. We used 35,246 global positioning system locations collected from 33 adult female elk to model probability of use as a function of 6 habitat variables: slope, aspect, elevation, habitat diversity, distance to shrub cover, and distance to road. We developed resource selection probability functions for individual elk, and then we averaged the coefficients to estimate population-level models for summer and winter periods. We used the population-level models to generate predictive maps by assigning pixels across the study area to 1 of 4 use categories (i.e., high, medium-high, medium-low, or low), based on quartiles of the predictions. Model coefficients and predictive maps indicated that elk selected for summer habitats characterized by higher elevations in areas of high vegetative diversity, close to shrub cover, northerly aspects, moderate slopes, and away from roads. Winter habitat selection patterns were similar, except elk shifted to areas with lower elevations and southerly aspects. We validated predictive maps by using 528 locations collected from an independent sample of radiomarked elk (n = 55) and calculating the proportion of locations that occurred in each of the 4 use categories. Together, the high- and medium-high use categories of the summer and winter predictive maps contained 92% and 74% of summer and winter elk locations, respectively. Our population-level models and associated predictive maps were successful in predicting winter and summer habitat use by elk in a nonforested environment. In the absence of forest cover, elk seemed to rely on a combination of shrubs, topography, and low human disturbance to meet their thermal and hiding cover requirements.

Journal of Wildlife Management↗

Bighorn sheep response to road-related disturbances in Rocky Mountain National Park, Colorado

Bighorn sheep (Ovis canadensis) use of Sheep Lakes mineral site, Rocky Mountain National Park, Colorado, USA, has decreased since 1996. Officials were concerned that human disturbance may have been contributing to this decline in use. We evaluated effects of vehicular traffic and other road-related disturbance on bighorn use of Sheep Lakes in the summers of 2002 and 2003. We found that the time and number of attempts required by bighorn to reach Sheep Lakes was positively related to the number of vehicles and people present at Sheep Lakes. Further, the number of bighorn individuals and groups attempting to visit Sheep Lakes were negatively affected by disturbance associated with the site. The number of vehicles recorded the hour before bighorn tried to access Sheep Lakes best predicted an animal's failure to cross Fall River Road and reach Sheep Lakes. We conclude that human and road-related disturbance at Sheep Lakes negatively affected bighorn use of the mineral site. Because Sheep Lakes may be important for bighorn sheep, especially for lamb production and survival, the negative influence of disturbance may compromise health and productivity of the Mummy Range bighorn sheep.

Journal of Wildlife Management↗

Estimation of successful breeding pairs for wolves in the Northern Rocky Mountains, USA

Under the Endangered Species Act, documenting recovery and federally mandated population levels of wolves (Canis lupus) in the Northern Rocky Mountains (NRM) requires monitoring wolf packs that successfully recruit young. United States Fish and Wildlife Service regulations define successful breeding pairs as packs estimated to contain an adult male and female, accompanied by ???2 pups on 31 December of a given year. Monitoring successful breeding pairs will become more difficult following proposed delisting of NRM wolves; alternatives to historically intensive methods, appropriate to the different ecological and regulatory context following delisting, are required. Because pack size is easier to monitor than pack composition, we estimated probability a pack would contain a successful breeding pair based on its size for wolf populations inhabiting 6 areas in the NRM. We also evaluated the extent to which differences in demography of wolves and levels of human-caused mortality among the areas influenced the probability of packs of different sizes would contain successful breeding pairs. Probability curves differed among analysis areas, depending primarily on levels of human-caused mortality, secondarily on annual population growth rate, and little on annual population density. Probabilities that packs contained successful breeding pairs were more uniformly distributed across pack sizes in areas with low levels of human mortality and stable populations. Large packs in areas with high levels of human-caused mortality and high annual growth rates had relatively high probabilities of containing breeding pairs whereas those for small packs were relatively low. Our approach can be used by managers to estimate number of successful breeding pairs in a population where number of packs and their sizes are known. Following delisting of NRM wolves, human-caused mortality is likely to increase, resulting in more small packs with low probabilities of containing breeding pairs. Differing contributions of packs to wolf population growth based on their size suggests monitoring successful breeding pairs will provide more accurate insights into population dynamics of wolves than will monitoring number of packs or individuals only.

Journal of Wildlife Management↗

Den-site characteristics of black bears in Rocky Mountain National Park, Colorado

We compared historic (1985-1992) and contemporary (2003-2006) black bear (Ursus americanus) den locations in Rocky Mountain National Park (RMNP), Colorado, USA, for habitat and physiographic attributes of den sites and used maximum entropy modeling to determine which factors were most influential in predicting den-site locations. We observed variability in the relationship between den locations and distance to trails and elevation over rime. Locations of historic den sites were most associated with slope, elevation, and covertype, whereas contemporary sites were associated with slope, distance to roads, aspect, and canopy height. Although relationships to covariates differed between historic and contemporary periods, preferred den-site characteristics consistently included steep slopes and factors associated with greater snow depth. Distribution of den locations shifted toward areas closer to human developments, indicating little negative influence of this factor on den-site selection by black bears in RMNP.

Journal of Wildlife Management↗

Habitat assessment for giant pandas in the Qinling Mountain region of China

Because habitat loss and fragmentation threaten giant pandas (Ailuropoda melanoleuca), habitat protection and restoration are important conservation measures for this endangered species. However, distribution and value of potential habitat to giant pandas on a regional scale are not fully known. Therefore, we identified and ranked giant panda habitat in Foping Nature Reserve, Guanyinshan Nature Reserve, and adjacent areas in the Qinling Mountains of China. We used Mahalanobis distance and 11 digital habitat layers to develop a multivariate habitat signature associated with 247 surveyed giant panda locations, which we then applied to the study region. We identified approximately 128 km 2 of giant panda habitat in Foping Nature Reserve (43.6% of the reserve) and 49 km 2 in Guanyinshan Nature Reserve (33.6% of the reserve). We defined core habitat areas by incorporating a minimum patch-size criterion (5.5 km 2 ) based on home-range size. Percentage of core habitat area was higher in Foping Nature Reserve (41.8% of the reserve) than Guanyinshan Nature Reserve (26.3% of the reserve). Within the larger analysis region, Foping Nature Reserve contained 32.7% of all core habitat areas we identified, indicating regional importance of the reserve. We observed a negative relationship between distribution of core areas and presence of roads and small villages. Protection of giant panda habitat at lower elevations and improvement of habitat linkages among core habitat areas are important in a regional approach to giant panda conservation.

Qinling Mountains↗

Demographics of an experimentally released population of elk in Great Smoky Mountains National Park

We assessed the potential for reestablishing elk ( Cervus elaphus ) in Great Smoky Mountains National Park (GSMNP), USA, by estimating vital rates of experimentally released animals from 2001 to 2006. Annual survival rates for calves ranged from 0.333 to 1.0 and averaged 0.592. Annual survival for subadult and adult elk (i.e., ≥1 yr of age) ranged from 0.690 to 0.933, depending on age and sex. We used those and other vital rates to model projected population growth and viability using a stochastic individual-based model. The annual growth rate (λ) of the modeled population over a 25-year period averaged 0.996 and declined from 1.059 the first year to 0.990 at year 25. The modeled population failed to attain a positive 25-year mean growth rate in 46.0% of the projections. Poor calf recruitment was an important determinant of low population growth. Predation by black bears ( Ursus americanus ) was the dominant calf mortality factor. Most of the variance of growth projections was due to demographic variation resulting from the small population size ( n = 61). Management actions such as predator control may help increase calf recruitment, but our projections suggest that the GSMNP elk population may be at risk for some time because of high demographic variation.

Great Smoky Mountains National Park↗

Growth and secondary production of aquatic insects along a gradient of Zn contamination in Rocky Mountain streams

Secondary production estimates from several Rocky Mountain streams were used to test hypotheses about the effects of chronic metal contamination on insect populations and ecosystem processes. Quantitative samples of chemistry, habitat, and benthic insects were collected monthly during the ice-free period (May–November) from five 2 nd - to 3 rd -order streams that varied primarily in Zn contamination. Secondary production was estimated for the 19 dominant taxa using increment-summation, size-frequency, and P/B methods. Uncertainty was estimated by bootstrapping estimates of mean abundance, biomass, and cohort production intervals. Secondary production of metal-sensitive Heptageniidae ( Rhithrogena robusta , Cinygmula spp., and Epeorus longimanus ) was lower in lightly to moderately contaminated streams than in reference streams. Experiments were done to determine whether herbivore growth was influenced by food quality in contaminated streams. Growth estimates from field and microcosm experiments revealed that low mayfly production in contaminated streams was caused mostly by reduced population abundances. Production of predatory stoneflies was also lower in contaminated streams than reference streams. Estimates of the trophic basis of production revealed that, although the relative contribution to community production from various food sources was similar among streams, total production attributable to algae and animal prey declined in contaminated streams. Much of the reduction in herbivory in contaminated streams was the result of lower production of heptageniids, especially R. robusta . Assemblage and taxon-specific estimates of secondary production were sensitive to variation in metal contamination and indicated that relatively low metal concentrations may have ecosystem-wide consequences for energy flow.

Journal of the North American Benthological Societ↗

Phytoplankton dynamics in three Rocky Mountain lakes, Colorado, USA

In 1984 and 1985 seasonal changes in phytoplankton were studied in a system of three lakes in Loch Vale, Rocky Mountain National Park, Colorado. Three periods were evident: (1) A spring bloom, during snowmelt, of the planktonic diatom Asterionella formosa , (2) a mid-summer period of minimal algal abundance, and (3) a fall bloom of the blue-green alga Oscillatoria limnetica . Seasonal phytoplankton dynamics in these lakes are controlled partially by the rapid flushing rate during snowmelt and the transport of phytoplankton from the highest lake to the lower lakes by the stream, Icy Brook. During snowmelt, the A. formosa population in the most downstream lake has a net rate of increase of 0.34 d -1 , which is calculated from the flushing rate and from the A. formosa abundance in the inflow from the upstream lake and in the downstream lake. Measurement of photosynthetic rates at different depths during the three periods confirmed the rapid growth of A. formosa during the spring. The decline in A. formosa after snowmelt may be related to grazing by developing zooplankton populations. The possible importance of the seasonal variations in nitrate concentrations were evaluated in situ enrichment experiments. For A. formosa and O. limnetica populations, growth stimulation resulted from 8- or 16-micromolar amendments of calcium nitrate and sulfuric acid, but the reason for this stimulation could not be determined from these experiments.

Colorado↗

Dynamics of Mojave Desert shrub assemblages in the Panamint Mountains, California.

We studied shrub communities in the Panamint Mountains of the Mojave Desert to determine whether vegetational changes after disturbance can be characterized as succession according to modern successional theory. We found, on a variety of disturbed and undisturbed sites, that the rate of change was a function of the type and age of disturbance. Recent debris—flow deposits were colonized by shrub assemblages of different species composition than those on the surrounding, older debris—flow deposits and other geomorphically stable surfaces. Colonization of human—disturbed sites was highly variable, but species compositions were different from the predisturbance species composition. In Grayia—Lycium assemblages, Grayia spinosa reasserted its dominance over colonizers relatively quickly. In Coleogyne assemblages, typically found on older geomorphic surfaces, species composition differences persisted considerably longer, depending on the severity of the initial disturbance. Extremely stable assemblages, dominated by Coleogyne ramosissima, occurred on the oldest, least disturbed surfaces. The variability of species composition among disturbed sites was greater than the variability among undisturbed and geomorphically stable sites, in accord with convergent succession. Models of desert succession should consider several factors: (1) colonization is dependent largely on the severity of disturbances and residual biotic components; (2) the time span for recovery may be longer than past periods of climatic and geomorphic stability; and (3) colonizing species may have considerable range in their life—history strategies.

California↗

Disturbance regime and disturbance interactions in Rocky Mountain subalpine forest

1 The spatial and temporal patterns of fire, snow avalanches and spruce beetle out-breaks were investigated in Marvine Lakes Valley in the Colorado Rocky Mountains in forests of Picea engelmannii , Abies lasiocarpa , Pseudotsuga menziesii and Populus tremuloides . Dates and locations of disturbances were determined by dendrochronological techniques. A geographic information system (GIS) was used to calculate areas affected by the different disturbance agents and to examine the spatial relationships of the different disturbances. 2 In the Marvine Lakes Valley, major disturbance was caused by fire in the 1470s, the 1630s and the 1870s and by spruce beetle outbreak in c. 1716, 1827 and 1949. 3 Since c. 1633, 9% of the Marvine Lakes Valley has been affected by snow avalanches, 38.6% by spruce beetle outbreak and 59.1% by fire. At sites susceptible to avalanches, avalanches occur at a near-annual frequency. The mean return intervals for fire and spruce beetle outbreaks are 202 and 116.5 years, respectively. Turnover times for fire and spruce beetle outbreaks are 521 and 259 years, respectively. 4 Several types of disturbance interaction were identified. For example, large and severe snow avalanches influence the spread of fire. Similarly, following a stand-devastating fire or avalanche, Picea populations will not support a spruce beetle outbreak until individual trees reach a minimum diameter which represents at least 70 years' growth. Thus, recent fires and beetle outbreaks have nonoverlapping distributions.

Journal of Ecology↗

Distribution of corticolous noncrustose lichens on trunks of Rocky Mountain junipers in Boulder County, Colorado.

Nineteen species of noncrustose lichens were found on Juniperus scopulorum bark; 3 species had relatively high cover and frequency values and were characterized as typical lichens of Rocky Mountain junipers: Xanthoria fallax, Phaeophyscia hirsuta and Physcia caesia. Total cover per tree was low (4%) and most species preferred the N and E sides of trunk bases. These distributional trends may reflect gradients of exposure to wind, insolation, and rate of bark exfoliation. -Author Juniperus scopulorum Phaeophyscia hirsuta Physcia caesia Xanthoria fallax.

Colorado↗

The mountain that moved: geologic wonders of the George Washington and Jefferson National Forests

Prehistoric, giant landslides in Montgomery and Craig Counties, Va., in the Blacksburg/Wythe Ranger Districts of the Jefferson National Forest, are the largest known landslides in eastern North America and are among the largest in the world. One of the landslides is more than 3 miles long! The ancient, giant landslides extend for more than 20 miles along the eastern slope of Sinking Creek Mountain. Enormous slabs of rock ranging from about 0.2 to more than 1.5 square miles in size broke loose and slid downslope under the influence of gravity. The movement of some slides may have been slow, but the movement of others was probably sudden and catastrophic.

Virginia↗

Principal facts for gravity stations in the Bearpaw Mountains and vicinity, Montana

Observed gravity values, station locations, terrain corrections, and Bouguer gravity data are provided in tabular form for approximately 500 gravity observations in north-central Montana. These data were used in preparation of -- Peterson, Donald L. and Rambo, William L., 1967, Bouguer gravity anomaly map of the Bearpaw Mountains and vicinity, Montana: U.S. Geol. Survey open-file report.

Montana↗

Remote sensor application studies report, July 1, 1968 to June 30, 1969: Remote sensing reconnaissance, Mill creek area, Arbuckle Mountains, Oklahoma

As part of the U.S. Geological Survey's Remote Sensor Application Studies program, infrared images and several kinds of photographs were obtained on reconnaissance flights over two areas in the Arbuckle Mountains near Mill Creek, Oklahoma. These data were used in a preliminary investigation (1) to determine the diagnostic reflection and emission characteristics of various rock types, and (2) io evaluate the perturbing influence of atmospheric conditions, surface coatings, rock texture, and topography on the observed reflected and emitted energy in the thermal infrared (8-14μ) part of the spectrum

Oklahoma↗

Reconnaissance geology north of the Hoholitna River, Taylor Mountains D-1 1:63,360-scale quadrangle, southwestern Alaska: A section in Geological studies in Alaska by the U.S. Geological Survey, 1999

The lower Paleozoic (Silurian and Ordovician) carbonate stratal succession is divided into six unnamed stratigraphic units in the northern part of the Taylor Mountains D-1 1:63,360-scale quadrangle of southwestern Alaska. Several of these units have previously been recognized in the McGrath and Medfra quadrangles to the northeast in strata of the Nixon Fork subterrane of the Farewell terrane (Decker and others, 1994 ). These rocks occur along the south side of a prominent east-west-trending anticlinoria! axis exposed slightly to the north in the Sleetmute A-2 1:63,360-scale quadrangle. Rocks of the Nixon Fork subterrane are now thought to represent a continental margin sequence rifted from Siberia. The low thermal alteration indices exhibited by the rocks of this area have elicited interest for petroleum exploration. However, low total organic carbon (TOC) values from potential source rocks within this lower Paleozoic succession indicate low petroleum potential.

Alaska↗

Mineral appraisal of the Poker Jim Ridge and Fort Warner areas of the Hart Mountain National Antelope Refuge, Lake County, Oregon

The Poker Jim Ridge and Fort Warner areas, which are candidate areas for inclusion in the National Wilderness Preservation System, are in the Hart Mountain National Antelope Refuge in east-central Lake County, south-central Oregon. The two areas form part of an upraised and tilted fault block--a structural element that is typical of the northern part of the Great Basin province. Rocks in the two areas are principally volcanic in origin. They have been faulted, uplifted, and tilted eastward, the older rocks having been more strongly deformed than the younger ones.

Oregon↗

Manganese deposits in the Drum Mountains, Juab and Millard Counties, Utah

The Drum Mountains are in west-central Utah 30 miles northwest of Delta, between the Sevier Desert on the east and Whirlwind Valley on the west. It is a typically barren desert range comprising a westward-tilted structural unit in which is exposed as much as 9,000 feet of quartzite (Cambrian and Precambrian?) and 3,000 feet of carbonate rocks of Cambrian age. These beds, which strike northward and dip west, are cut by myriad east- to northeast-trending faults with displacements of a few feet to a few thousand feet. Quartz monzonite dikes, pebble dikes, and vein deposits are present locally along the faults. The Cambrian rocks are overlain unconformably by volcanic rocks of probable Tertiary age. Bodies of manganese carbonate ore were formed by replacement of two 20-foot beds of impure dolomite at the base of the sequence of carbonate rocks, along their intersection with certain preore faults. The feeding fissures locally contain veins in which rhodochrosite is associated with base metal sulfides. Downward- moving meteoric water has oxidized the ore bodies to a depth of 100 to 200 feet except where they are sealed off by structural or stratigraphic traps. From 1925 to 1953, 72,462 long tons of manganese ore with an average grade of about 25 percent Mn were shipped.

Utah↗