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Using high resolution satellite and telemetry data to track flooded habitats, their use by waterfowl, and evaluate effects of drought on waterfowl and shorebird bioenergetics in California

Wetland managers in the Central Valley of California, a dynamic hydrological landscape, require information regarding the amount and location of existing wetland habitat to make decisions on how to best use water resources to support multiple wildlife objectives, particularly during drought. Scientists from the U.S. Geological Survey Western Ecological Research Center (WERC), Point Blue Conservation Science (Point Blue), and the U.S. Fish and Wildlife Service (USFWS) partnered to learn how wetland and flooded agricultural habitats used by waterfowl and shorebirds change during the non-breeding season (July–April) particularly during drought. During extreme drought conditions, the ability to provide sufficient water for wildlife often depends on the timing of water deliveries to managed wetlands and winter-flooded crop fields and decisions on whether to fallow croplands. Waterfowl and shorebirds could be particularly affected by these decisions because they typically rest and feed in flooded habitats. Poor habitat conditions resulting from spatially or temporally suboptimal water deliveries (that is, mismatch between waterfowl habitat needs and timing and location of flooded habitats) could reduce waterfowl hunting opportunities and body condition. Point Blue scientists developed a system for near real-time tracking of habitats used by waterfowl, shorebirds, and some other wetland-dependent “waterbirds” ( www.pointblue.org/watertracker ) and to assess the impacts of drought on habitat availability and on waterfowl and shorebird bioenergetics. The WERC researchers linked these data with near real-time tracking (telemetry) data of duck locations throughout the Valley. The team used these two datasets to relate duck locations to open-water characteristics and to learn how waterfowl use habitats under spatially and temporally changing conditions during drought and non-drought periods. We found that recent extreme drought (2013–15) significantly changed the timing and magnitude of flooding and consequently reduced the availability of habitats used by waterfowl and shorebirds more than other recent historic droughts 2000–11. Drought reduced irrigations of moist soil seed plants and thus there was lower food energy available for waterfowl. Analyses using bioenergetics models indicated that, overall, extreme drought increased food energy deficits (total number of deficit days) for shorebirds and waterfowl. Our analysis indicated a strong direct relationship between duck locations and classified habitat derived from open-water data during the wintering period (October–March). This result helps confirm the application of dynamic water data to identify flooded areas that provide waterfowl habitat. Presence of open water at a 1-hectare resolution can be used effectively to identify flooded landscape areas available as habitat for ducks. Our discoveries from evaluating use of space by ducks also indicated that nighttime feeding locations of ducks were concentrated nearby primary roosts and that foraging distances could depend on hydrologic dynamics of location (Suisun Marsh versus California excluding Suisun Marsh) and time of season (early, middle, late). Other results indicated that some areas on the California landscape with extremely reliable water supplies could receive consistent use by ducks year after year (in essence, almost drought proof). The Water Tracker is set up to automatically track wetland habitat and food availability each year and is making these data available to water and wetland managers. Results from this research are a significant step toward understanding how waterfowl and shorebird habitats can be optimally managed on the landscape to support desired populations of these migratory birds during extreme drought.

California↗

Physics to fish—Understanding the factors that create and sustain native fish habitat in the San Francisco Estuary

Executive Summary The Bureau of Reclamation (Reclamation) operates the Central Valley Project (CVP), one of the nation’s largest water projects. Reclamation has an ongoing need to improve the scientific basis for adaptive management of the CVP and, by extension, joint operations with California’s State Water Project. The U.S. Geological Survey (USGS) works cooperatively with the Bureau of Reclamation to provide scientific support for the management of Reclamation’s CVP project. Major habitat restoration efforts and a new water-diversion point are planned to benefit delta smelt ( Hypomesus transpacificus ) and other species of concern while ensuring the reliability of water supply. In addition, various flow actions and management activities have been identified as possible methods to increase populations of delta smelt and salmonid ( Oncorhynchus spp.) runs of concern. The overarching goal of this cooperative project was to provide Reclamation with the scientific information needed to evaluate the efficacy of ongoing and future adaptive management actions and to improve the scientific basis for more flexible CVP operations that would achieve water-supply reliability and fish protection. The research and monitoring described in this report comprises the period 2015–19 and focuses on management issues related to native fish species of concern, especially delta smelt. Conserving the delta smelt population while providing a reliable water supply is a primary management and policy issue in California. Our approach for this cooperative project is based on the “physics to fish” concept, the idea that high-quality habitat is generated and sustained by the interaction between physical processes and the landscape. These interactions create a template for chemical and biological processes that can change across a variety of spatial and temporal scales. Following this concept, this project (hereafter referred to as “the physics to fish project”) included monitoring and studies of water flows, sediments, water quality, and invertebrate and fish dynamics across a range of spatial and temporal scales and in regions relevant to resource managers tasked with managing water supplies and ecosystem health in the San Francisco Estuary. The intent of this approach was to document the habitat conditions, important processes, and interactions among them that create high-quality habitat for native fishes so that the likely effects of future management actions (for example, habitat restoration) can be objectively assessed at the local (site-specific), regional (within subregions of the estuary), and landscape (across the entire estuary and beyond) scales. Hydrodynamically, the upper estuary (landward of Carquinez Strait) is characterized by a fixed volume of tidally exchanged water (for example, tidal prism) that interacts with the existing channel network and bathymetry to create regions with differing hydrodynamics. Our results indicate that careful study of construction or reoperation of existing infrastructure to perform management actions can help (1) improve the accuracy of hydrodynamic models; (2) further understanding of ecological effects; and (3) enhance abilities to predict ecological outcomes. At the local scale, we developed a new concept called the Lagrangian to Eulerian (LE) ratio that can be used as a tool for understanding the importance of various hydrodynamic processes in specific channels or channel networks and for forecasting transport dynamics. Channels with LE ratios<1 in a channel network or in a dead-end slough are hydrodynamically able to develop an exchange zone between two parcels of water that may have different chemical and physical properties. In a dead-end channel, there is a landward region with long residence time (no-exchange zone) and a seaward region with short residence time (high-exchange zone) that are well mixed with seaward waters. At the transition (exchange zone) between the high and no-exchange regions, a gradient will form in water-quality constituents that differ in concentration between the landward and seaward waters. Turbidity affects fish habitat and has declined through time in the San Francisco Estuary. Average turbidity across the Sacramento–San Joaquin Delta (hereafter referred to as “the Delta”) is dependent on annual hydrology. In dry years, the region around Cache Slough (known regionally as the “Cache Slough Complex”) in the northern Delta is generally more turbid than Suisun Bay and the lower Sacramento River. When the Yolo By-Pass (known regionally as “Yolo Bypass”), a large flood bypass that runs parallel to the Sacramento River in the northern Delta, is not flooding and river flows are lower, sediment is usually transported into the Cache Slough Complex because flood tides dominate ebb tides, resulting in transport of suspended sediment from seaward areas of the upper estuary into the Cache Slough Complex. These hydrodynamic conditions also favor the formation of turbidity maximums (TMs) in the Cache Slough Complex. The TMs are areas of higher suspended-sediment concentration, providing higher-turbidity habitat favored by some fishes, including delta smelt, and they can also concentrate other constituents, including phytoplankton and organic carbon that can be important in food webs. Pelagic primary production by phytoplankton is the basis for Delta food webs supporting pelagic fishes such as delta smelt; however, phytoplankton abundance in the Delta has declined during recent decades. We examined how nutrients, hydrodynamics, and other factors affect phytoplankton blooms. Based on our results, we developed three new concepts of phytoplankton bloom formation in the Delta, each associated with a distinct set of hydrologic conditions. First, productivity cascades highlighted how local processes can contribute to phytoplankton blooms observed at the regional scale. Second, we observed phytoplankton blooms in the upper San Francisco Estuary that were associated with transport out of Yolo By-Pass (transport blooms). Third, we also documented a series of phytoplankton blooms that were in the confluence area at the landward edge of Suisun Bay. The conditions leading to creation of confluence phytoplankton blooms are not yet understood, but the confluence region connects the Cache Slough Complex with Suisun Marsh. Therefore, blooms in this area have the potential to spread to large areas of the Delta. At the landscape scale, the distribution of the invasive clams ( Potamocorbula amurensis and Corbicula fluminea , hereafter referred to as “ Corbicula ”) is driven by salinity. At smaller spatial scales, the distribution of either species is sensitive to multiple factors affecting survival and reproduction, complicating efforts to predict distribution and abundance without considering local-scale conditions across the area of interest. In the Cache Slough Complex, the area landward of the exchange zone in regions with LE ratio<1 were characterized by low abundances of Corbicula probably because recruits from seaward areas are not transported past the exchange zone and because there are no landward tributaries with adult Corbicula to provide an upstream source of recruits. Corbicula biomass was highest near or downstream from the exchange zone consistent with Corbicula grazing on phytoplankton produced in the exchange zone or transported from the no-exchange zone. The severity of Corbicula grazing could be reduced by manipulating the hydrodynamic characteristics of waterways; however, the beneficial and harmful effects on the organisms meant to benefit from increased phytoplankton production, including zooplankton and fish species of concern, should be thoroughly examined before manipulating hydrodynamic characteristics. The distribution of fishes at the landscape scale is generally driven by the position of the salinity field in the estuary. The physics to fish project compared distributions of fishes at Ryer Island, a tidal wetland in Suisun Bay and a region of variable salinity, with fish distributions at the Cache Slough Complex, a freshwater region. At Ryer Island, there was an absence of freshwater invasive species and an abundance of native species, such as Sacramento splittail ( Pogonichthys macrolepidotus ), tule perch ( Hysterocarpus traskii ), and Sacramento pikeminnow ( Ptychocheilus grandis ). The native species were almost exclusively captured in wetland and nearshore shallow-water habitat regardless of water-quality conditions. In the Cache Slough Complex, our regional scale objective was to elucidate how hydrodynamic-physical habitat interactions drive fish-community structure. Our studies showed that dendritic channel systems were better able to support native species, while intertidal habitats supported those species best able to exploit the transient character of the habitat. Habitats upstream from the exchange zone were especially important in supporting high numbers of native fishes relative to within or downstream from the exchange zone. Many of the native species were associated with tidal marsh in the no-exchange zone. More pelagic-oriented, mobile species, such as Striped Bass ( Morone saxatilis ), threadfin shad ( Dorosoma petenense ), and Sacramento pikeminnow, were more affected by water-quality conditions, such as turbidity. The physics to fish concept developed in this project provides a framework for designing individual projects and for considering the cumulative effects of multiple projects in a region, using the LE ratio as a guiding metric. The physics to fish concept may also provide a suitable framework for coordinating management actions. Tidal wetlands can function in several ways in the hydrodynamic framework. Relatively small tidal wetlands with short channel networks and with LE ratios>1 are not able to maintain a landward no-exchange zone or an exchange zone. This likely means that any contributions to pelagic food webs would be limited to resources derived from wetland vegetation, which can include dissolved and particulate organic matter (detritus) and populations of consumers that can increase in abundance based on those resources. The fate of the contributed production from these channels depends on the characteristics of the receiving waters seaward of the tidal wetland. If these channels join a large system such as Suisun Bay, then any contribution is likely to be rapidly dispersed in the larger volume; however, the channel junction might provide a focal point for consumers, such as fishes, to congregate and feed on material leaving the wetland on ebb tides before it is dispersed in the larger volume. Fishes might also access these resources by entering the wetland. The physics to fish project has established a foundation and several new concepts for understanding how habitat restoration can benefit native fish populations at the local and regional levels. Many of the ideas regarding habitat restoration and channel modifications outlined in this report could help guide management actions that could improve conditions for native fishes at little or no water cost beyond water already dedicated to other management actions. A complete list of products originating from this work is provided in appendix 1.

California↗

Surficial geology and geomorphology of Potter County, Pennsylvania

Potter County is located in the Appalachian Plateaus of north-central Pennsylvania and contains the headwaters of the Genesee River, the Allegheny River, and the Susquehanna River. Drift of Wisconsin age covers the northeastern part of the county. This study includes a detailed survev of the surficial deposits of the Genesee quadrangle in north-central Potter County and a reconnaissance of the remainder of the county; a soil survey and a botanical survey were carried on concurrently. The region is a deeply dissected plateau having extensive areas of steeply sloping land separated by narrow ridges and valleys; there is very little level land. Near the junction of the three watersheds the uplands rise to altitudes of more than 2,500 feet. The maximum relief in the Susquehanna drainage is more than 1,500 feet; in the Genesee and Allegheny drainage it. is about 800 feet. Valley walls are steep (15° to 30°), and the uplands have gentle slopes (0.5° to 10°). The drainage pattern is trellised. The climate is continental. Temperatures range from about -30° F. to more than 100° F. The average annual precipitation ranges approximately from 34 to 42 inches. Floods may occur at any season of the year. The large volumes of water from rain or melting snow carried by small streams come from springs. There is little precise data on frost in the ground, but it is probable that the ground seldom freezes in forested areas. The soils of Potter County have relatively immature profiles with poorly developed horizons that commonly have many characteristics inherited from their parent materials. At the great soil group level, the zonal soils are divided into Podzol soils and Brown Podzolic soils. Many soils have a high silt content in the upper part of the profile, apparently derived (at least partly) from a mantle of eolian silt. Mos~ of Potter County is covered by second-growth forests consisting of 40- to 60-year-old hardwood stands. The present forests growing on slopes and summits are composed approximately of 25 species of trees. The northern hardwood region includes most of the county, with an oak-forest region near the borders, principally along its southern margin. Potter County is underlain by sandstone, siltstone, shale, conglomerate, and minor amounts of coal and calcareous rock that range in age from Late Devonian to Pennsylvanian. These rocks form broad open folds that strike northeast. South of the border of the Wisconsin drift, and possibly at two localities inside the drift border, are scattered remnants of ancient soils (here called paleosol), that were formed in preWisconsin time-probably during the Sangamon interglacial stage. This paleosol ranges in texture from clay loam to silt loam, ranges in color from yellowish red to red, includes a few percent to more than 25 percent of rock fragments, and apparently contains a small percentage of gibbsite and varying amounts of kaolinite. Known thicknesses range from 1 to 33 feet. Paleosol was developed on diverse kinds of parent material, such as till, stratified drift, colluvium, and residuum, at altitudes ranging from a few hundred to 2,400 feet. The climatic conditions under which the paleosol formed are uncertain; however, these ancient soils may record an episode of subtropical climatic conditions during which lateritic soils were formed. Perhaps these soils are analogous to the Red-Yellow Podzolic soils of southeastern United States. Except for one possible remnant, no pre-Wisconsin drift has been recognized in Potter County. The Wisconsin glacial deposits of Potter County belong to either the Iowan or Tazewell substages and are dominantly till with minor amounts of glaciofluvial deposits. Erratics of igneous or metamorphic rock comprise less than 0.1 percent of the total number of rock fragments. The till is slightly weathered to depths ranging from 3 to about 12 feet. The drift border is indefinite and has been drawn at the southern limit of erratics or well-rounded or striated pebbles and is only locally marked by a terminal moraine or by a distinct change in the surficial deposits. The drift border is relatively straight and crosses the Genesee quadrangle in a northwesterly direction with little regard for the major topographic features, thus suggesting that the Wisconsin ice sheet had a relatively straight and steep front. Over most of the unglaciated part of Potter County, the bedrock is concealed beneath rubble that probably was formed during the Iowan or Tazewell substage, almost contemporaneously with the adjacent drift. In general, the rubble is thickest and most extensive within about 10 miles of the drift border, becoming thinner and less continuous farther away. The apparent parallelism between a belt of thick periglacial deposits and the drift border suggests that the deposits result from climatic factors in operation while the Wisconsin ice sheet was nearby. Ancient soil structures or patterned ground occur at, or near, the surface of both the periglacial deposits and the adjacent drift. These ancient soil structures are so similar to modern forms in arctic or alpine environments that they are considered to be the result of vigorous frost action. Many of the structures are believed to be a result of down-slope movement of debris by solifluction, facilitated by a frozen subsoil as much as 10 feet deep. Perennially frozen ground may have been present, but this is not a prerequisite. The periglacial deposits underlie long smooth slopes that extend from ridge crest to valley bottom. Flood plains are absent near the headwaters of many streams, the valley walls forming a V-shaped profile. While frost action was in progress, forests probably were restricted to flood plains, lower slopes, and scattered upland areas. Large parts of the upland were bare or partly covered by tundra vegetation; elsewhere, there were scattered trees but no dense forest. 1 2 SURFICIAL GEOLOGY AND GEOMORPHOLOGY OF POTTER COUNTY, PENNSYLVANIA Recent alluvium and alluvial fans include sand and sandy loams, 1 to 3 feet thick, that overlie gravel. The alluvium contains organic matter and lenses of finer materials. Thickness ranges from a few to more than 100 feet. Along the principal streams the alluvium probably overlies Pleistocene deposits. Most of the alluvial fans are composed of unstratified rubbly, pebbly, cobbly. or bouldery sandy loams to silty clay loams with local lenses of stratified sand and gravel. The alluvial fans mapped in the Genesee quadrangle probably include both Wisconsin stage and Recent deposits. The summits of the A.ppalachian Plateaus in north-central Pennsylvania have long been recognized as the remnants or traces of one or more peneplains. To test this hypothesis, a restored contour map was prepared to show the configuration of a supposed peneplain on the assumption that the plateau tops are remnants of such an old erosion surface. The restored contours delineate a surface that corresponds roughly to rock structure. In general, the uplands slope parallel to the dip of the bedrock. The major streams, such as the West Branch Susquehanna River, cross the ridges and valleys of the restored surface in such a way that it is difficult to suppose that the restored surface was ever graded to these streams. On the contrary, it is probable that the restored surface never existed and that the plateau tops are structurally controlled surfaces held up by sandstone and conglomerate beds in the Pottsville and Pocono formations. The plateau tops may have been lowered by erosion as much as 200 feet during the Pleistocene-in other words, after the major streams were incised. If this portion of the Appalachian Plateaus was ever reduced to a peneplain, such a hypothetical surface must have lain many hundreds of feet above the uplands of the present day. The only alternative that might involve peneplanation is the improbable hypothesis that the plateau tops are remnants of a slightly deformed peneplain and that the peneplain was folded along the axes of the Appalachian orogeny. This remote possibility is not supported by any known evidence. The geomorphic analysis yields no new data on the origin of the cross-axial drainage. Regardless of whether the plateaus are peneplain remnants or are structurally controlled surfaces, the beginning of the major southeastward-flowing streams long antedates the existing landscape. The geomorphic history of Potter County begins with an assumed long interval of erosion during the Mesozoic and early Cenozoic eras, for which no record remains in this area. The southeast master drainage was established by the latter part of the Tertiary period (perhaps at a much earlier date), probably as the result of the northwestward migration of the Atlanticinterior divide. In late Pliocene(?) time, areas adjacent to parts of the West Branch Susquehanna River-and probably elsewhere-had a moderate relief ranging from 300 to 700 feet. Some segments of the West Branch meandered across a broad valley that lay about 900 feet above the present streams. The landscape probably was covered by deep residual soils, perhaps by saprolite. The early Pleistocene history of Potter County is essentially unknown. No deposits of the Kansan stage are known except for a possible trace of pre-Illinoian drift on the uplands in central Potter County (Ayers Hill quadrangle). Some deposits in central and eastern Pennsylvania may be of Kansan age. It is probable that the assumed Aftonian regolith was removed by mass movements and other processes during the Kansan stage, thus resulting in a lowering of the plateau tops by as much as 10 feet. By the close of the Yarmouth(?) interglacial stage the major streams were incised to essentially their present depths. The climates of the Yarmouth interglacial stage probably produced deep residual soils over the landscape, parts of which may still be preserved in the paleosol remnants of the present day. No Illinoian drift is known in Potter County, but drift assigned to this stage occurs in areas to the northwest and to the southeast. Some valleys, such as Kettle Creek valley, were filled with sand and gravel alluvium to depths of as much as 150 feet above their present flood plains. It is assumed that the Yarmouth residual soils were removed by mass movements and other processes induced by a periglacial climate, thus lowering the plateau tops by as much as 10 feet. During the Sangamon interglacial stage, deep (10-to-20 foot) residual soils or paleosol were developed in Potter County and probably throughout much of Pennsylvania, perhaps as a result of lateritic weathering in a subtropical climate. It is possible that the paleosol was largely removed by mass movements and by running water during late Sangamon time. During either the Iowan or Tazewell substages of the Wisconsin (perhaps the Iowan), the ice sheet advanced into the northeastern part of Potter County. The drift is similar to the Olean drift (local usage). The paleosol was almost completely removed by mass movements and other processes induced· by a periglacial climate, prior to drift deposition. This removal probably resulted in a lowering of the plateau tops by as much as 10 feet since Sangamon time. Nearly contemporaneously with drift deposition, the periglacial deposits were formed by frost heaving, solifluction, and fluvial transport in areas outside the drift border. Soil structures or patterned ground were developed on both the drift and the periglacial deposits. It is probable that the forests in the periglacial area were greatly restricted and that large areas on the uplands were essentially treeless. Little is known about the history of Potter County in postOlean time. Presumably, forests completely covered the county by the onset of the next substage, during which the Binghamton drift of MacClintock and Apfel was deposited. This drift also is found in southern New York State. The formation of the alluvium and alluvial fans probably began in the Tazewell substage and continued during the Recent epoch. Since these deposits were formed there has been very little dissection. There is little, if any, difference between soils developed on periglacial deposits and soils developed on drift. The roots of fallen trees have disturbed the soil horizons, and it is unlikely that the existing soil profiles are more than 500 years old. The forested landscape of Potter County has a distinctive microrelief ranging from a few inches to a few feet of mounds and pits produced by the roots of fallen trees. Most mounds and pits range from 10 to 20 feet in length and from 6 to 15 feet in width. On level land, many mounds are oriented with their long axes trending northward, and in some areas the orientation is random. On slopes, the mounds are oriented with their long axes at right angles to the maximum slope as a result of trees falling downslope. The toppling of trees increases the permeability of surficial deposits and mixes and destroys the soil horizons. The microrelief is a factor in forest development. The toppling of trees on slopes is a significant agent of slope erosion. The process loosens, breaks up, or overturns the upper 2 to 3 feet of the forest soil, and it tends to make the surficial layer more stony and to produce features resembling soil structures.

Pennsylvania↗

A multidecade analysis of fluvial geomorphic evolution of the Spirit Lake blockage, Mount St. Helens, Washington

Volcanic eruptions can affect landscapes in many ways and consequently alter erosion and the fluxes of water and sediment. Hydrologic and geomorphic responses to volcanic disturbances are varied in both space and time, and, in some instances, can persist for decades to centuries. Understanding the broad context of how landscapes respond to eruptions can help inform how they may evolve, and therefore provides context for managing and mitigating hazards associated with future volcanic and hydrologic events. Here, we assess the geomorphic evolution of the upper North Fork Toutle River valley, the valley most heavily affected by the Mount St. Helens May 18 and later 1980s eruptions. By doing so, we provide context for the landscape changes caused by the eruptions as they relate to potential hydrological hazards associated with Spirit Lake, an iconic landform at the northern foot of the volcano. The Spirit Lake basin was transformed by the cataclysmic 1980 eruption and had its outlet blocked. The analyses presented provide context for considerations of potential outlets for Spirit Lake, a landform which might be viewed as a “sleeping giant” on this landscape: a giant capable of causing catastrophic downstream consequences if water is released uncontrollably from the lake.

Washington↗

Breeding birds of the upper Mississippi River floodplain forest: One community in a changing forest, 1994 to 1997

Floodplain forest on the upper Mississippi River (UMR), a unique habitat in the Midwest that is important for many bird species, has been reduced and is undergoing continued reduction and changes in structure and species diversity because of river engineering and invasive species. Hydrological changes are causing tree diversity to decline favoring Acer saccharinum (silver maple) and Fraxinus pennsylvanica (green ash). Invasive Phalaris arundinacea (reed canary grass, Phalaris ) threatens tree regeneration, and recent Agrilus planipennis (emerald ash borer) arrival threatens to decimate the important ash component of the forest canopy. During the 1990s, virtually no information was available about breeding songbird species and abundances on the UMR floodplain forest from along many river miles and a broad range of forest situations (for example, mainland, island, edge, interior). From 1994 to 1997, we surveyed breeding birds and sampled vegetation at 391 random points on UMR floodplain forest along a latitudinal gradient from Red Wing, Minnesota, to Clinton, Iowa, to characterize bird assemblages and associations with gradients in forest structure at survey points (local scale) and land cover composition within a 200-meter radius of survey points (landscape scale). Eighty-six bird species were detected during the study, but 28 species comprised 90 percent of all detections. Species that are typically associated with woodland edge or are tolerant of fragmentation were the most common: Setophaga ruticilla (American Redstart), Troglodytes aedon (House Wren), Turdus migratorius (American Robin), Quiscalus quiscula (Common Grackle), and Vireo gilvus (Warbling Vireo). Species typically associated with large forest patches— Setophaga cerulea (Cerulean Warbler), Hylocichla mustelina (Wood Thrush), and Dryocopus pileatus (Pileated Woodpecker)—were rare. Principal components analyses consistently described local habitat gradients related to canopy cover and Phalaris presence and described landscape gradients related to forest area and areas of open land cover types. However, nonmetric multidimensional scaling revealed no pattern in bird assemblages. Canonical correspondence analyses with local habitat variables for each year revealed that bird assemblages were affected by canopy cover, the presence of Phalaris , and the number of tree species. Four bird species were consistently associated with Phalaris presence or negatively with canopy cover, and no species were associated with the number of tree species variable. Although landscape variables were significantly related to the bird assemblage in canonical correspondence analyses, no bird species were consistently related to any landscape variable. These results indicate that there is one assemblage of forest birds on the UMR composed mainly of edge-tolerant species. Species associated with lower canopy cover and Phalaris presence may be favored to increase in abundance as canopy cover opens as trees die and Phalaris becomes more prevalent.

Illinois, Iowa, Minnesota, Wisconsin↗

The Pedestrian Evacuation Analyst: geographic information systems software for modeling hazard evacuation potential

Recent disasters such as the 2011 Tohoku, Japan, earthquake and tsunami; the 2013 Colorado floods; and the 2014 Oso, Washington, mudslide have raised awareness of catastrophic, sudden-onset hazards that arrive within minutes of the events that trigger them, such as local earthquakes or landslides. Due to the limited amount of time between generation and arrival of sudden-onset hazards, evacuations are typically self-initiated, on foot, and across the landscape (Wood and Schmidtlein, 2012). Although evacuation to naturally occurring high ground may be feasible in some vulnerable communities, evacuation modeling has demonstrated that other communities may require vertical-evacuation structures within a hazard zone, such as berms or buildings, if at-risk individuals are to survive some types of sudden-onset hazards (Wood and Schmidtlein, 2013). Researchers use both static least-cost-distance (LCD) and dynamic agent-based models to assess the pedestrian evacuation potential of vulnerable communities. Although both types of models help to understand the evacuation landscape, LCD models provide a more general overview that is independent of population distributions, which may be difficult to quantify given the dynamic spatial and temporal nature of populations (Wood and Schmidtlein, 2012). Recent LCD efforts related to local tsunami threats have focused on an anisotropic (directionally dependent) path distance modeling approach that incorporates travel directionality, multiple travel speed assumptions, and cost surfaces that reflect variations in slope and land cover (Wood and Schmidtlein, 2012, 2013). The Pedestrian Evacuation Analyst software implements this anisotropic path-distance approach for pedestrian evacuation from sudden-onset hazards, with a particular focus at this time on local tsunami threats. The model estimates evacuation potential based on elevation, direction of movement, land cover, and travel speed and creates a map showing travel times to safety (a time map) throughout a hazard zone. Model results provide a general, static view of the evacuation landscape at different pedestrian travel speeds and can be used to identify areas outside the reach of naturally occurring high ground. In addition, data on the size and location of different populations within the hazard zone can be integrated with travel-time maps to create tables and graphs of at-risk population counts as a function of travel time to safety. As a decision-support tool, the Pedestrian Evacuation Analyst provides the capability to evaluate the effectiveness of various vertical-evacuation structures within a study area, both through time maps of the modeled travel-time landscape with a potential structure in place and through comparisons of population counts within reach of safety. The Pedestrian Evacuation Analyst is designed for use by researchers examining the pedestrian-evacuation potential of an at-risk community. In communities where modeled evacuation times exceed the event (for example, tsunami wave) arrival time, researchers can use the software with emergency managers to assess the area and population served by potential vertical-evacuation options. By automating and managing the modeling process, the software allows researchers to concentrate efforts on providing crucial and timely information on community vulnerability to sudden-onset hazards.

Techniques and Methods↗

Widespread occupancy of the endangered northern myotis on northeastern Atlantic Coastal Plain islands

Northern myotis Myotis septentrionalis are one of the bat species most affected by white-nose syndrome (WNS), and disease-induced declines may cause compounding effects when combined with other threats such as habitat loss and fragmentation. Recent evidence suggests that peripheral populations are persisting in post-WNS years; however, the environmental factors that influence the occurrence of this species along the Atlantic Coastal Plain are virtually unknown. We conducted a large-scale acoustic survey on 3 islands: Long Island, New York, and Martha’s Vineyard and Nantucket, Massachusetts, USA, and used a multi-scale occupancy modeling approach to determine the landscape and abiotic factors affecting the distribution of northern myotis. Our estimates of occupancy and detection probability suggest widespread presence across the islands. At the local (200 m) scale, we identified strong negative effects of development on Long Island and Nantucket and a strong positive effect of forest habitat on Martha’s Vineyard. None of the variables we measured sufficiently explained the landscape (1 km 2 ) occupancy of this species, which was very high (ψ = 0.81–0.97), representing an outlier for this species in the post-WNS landscape. The lack of association at the landscape scale suggests that general differences in land cover are not a driving factor of higher occupancy of peripheral northern myotis populations, while local site- specific conditions may be supporting critical foraging or roosting habitat. Because islands are particularly vulnerable to human-driven habitat alteration due to the constraint of limited space, and development pressure is expected to increase, this study provides a baseline to enable managers to assess the effects of future environmental disturbances and monitor population trends to support long-term survival of northern myotis.

Massachusetts, New York↗

Predicted changes in climatic niche and climate refugia of conservation priority salamander species in the northeastern United States

Global climate change represents one of the most extensive and pervasive threats to wildlife populations. Amphibians, specifically salamanders, are particularly susceptible to the effects of changing climates due to their restrictive physiological requirements and low vagility; however, little is known about which landscapes and species are vulnerable to climate change. Our study objectives included, (1) evaluating species-specific predictions (based on 2050 climate projections) and vulnerabilities to climate change and (2) using collective species responses to identify areas of climate refugia for conservation priority salamanders in the northeastern United States. All evaluated salamander species were projected to lose a portion of their climatic niche. Averaged projected losses ranged from 3%–100% for individual species, with the Cow Knob Salamander ( Plethodon punctatus ), Cheat Mountain Salamander ( Plethodon nettingi ), Shenandoah Mountain Salamander ( Plethodon virginia ), Mabee’s Salamander ( Ambystoma mabeei ), and Streamside Salamander ( Ambystoma barbouri ) predicted to lose at least 97% of their landscape-scale climatic niche. The Western Allegheny Plateau was predicted to lose the greatest salamander climate refugia richness ( i.e. , number of species with a climatically-suitable niche in a landscape patch), whereas the Central Appalachians provided refugia for the greatest number of species during current and projected climate scenarios. Our results can be used to identify species and landscapes that are likely to be further affected by climate change and potentially resilient habitats that will provide consistent climatic conditions in the face of environmental change.

Forests↗

Predicting habitat and distribution of an interior highlands regional endemic winter stonefly (Allocapnia mohri) in Arkansas using random forest models

Stoneflies are a globally threatened aquatic insect order. In Arkansas, a diverse group of winter stonefly (Capniidae: Allocapnia ) have not been surveyed since the 1980s, likely because species-level identification requires the rarely-collected adult form. Allocapnia mohri , a regional endemic, was previously commonly found in mountainous, intermittent streams from the Ouachita Mountains ecoregion north to the Ozark Highlands, but no species distributional models including land use or climate variables exist to our knowledge. We collected adults from 71 stream reaches from the historic Arkansas range from November to April 2020 and 2021. We modeled distributions using random forest (RF) models populated with landscape, climate, and both data to determine which were most predictive of species presence. Correlations between landscape or climate variables and presence were examined using multiple logistic regression. The landscape RF models performed better than the climate or landscape + climate RF models. A. mohri presence sites tended to have a greater elevation, a lower mean July temperature, and a greater percentage of very slow infiltration soils in the watershed, compared to absence sites. A. mohri was absent at the Ouachita Mountains sites and may be experiencing a range contraction or migration northward.

Arkansas↗

Fire and the distribution and uncertainty of carbon sequestered as above-ground tree biomass in Yosemite and Sequoia & Kings Canyon National Parks

Fire is one of the principal agents changing forest carbon stocks and landscape level distributions of carbon, but few studies have addressed how accurate carbon accounting of fire-killed trees is or can be. We used a large number of forested plots (1646), detailed selection of species-specific and location-specific allometric equations, vegetation type maps with high levels of accuracy, and Monte Carlo simulation to model the amount and uncertainty of aboveground tree carbon present in tree species (hereafter, carbon) within Yosemite and Sequoia & Kings Canyon National Parks. We estimated aboveground carbon in trees within Yosemite National Park to be 25 Tg of carbon (C) (confidence interval (CI): 23–27 Tg C), and in Sequoia & Kings Canyon National Park to be 20 Tg C (CI: 18–21 Tg C). Low-severity and moderate-severity fire had little or no effect on the amount of carbon sequestered in trees at the landscape scale, and high-severity fire did not immediately consume much carbon. Although many of our data inputs were more accurate than those used in similar studies in other locations, the total uncertainty of carbon estimates was still greater than ±10%, mostly due to potential uncertainties in landscape-scale vegetation type mismatches and trees larger than the ranges of existing allometric equations. If carbon inventories are to be meaningfully used in policy, there is an urgent need for more accurate landscape classification methods, improvement in allometric equations for tree species, and better understanding of the uncertainties inherent in existing carbon accounting methods.

California↗

Exploiting differential vegetation phenology for satellite-based mapping of semiarid grass vegetation in the southwestern United States and northern Mexico

We developed and evaluated a methodology for subpixel discrimination and large-area mapping of the perennial warm-season (C 4 ) grass component of vegetation cover in mixed-composition landscapes of the southwestern United States and northern Mexico. We describe the methodology within a general, conceptual framework that we identify as the differential vegetation phenology (DVP) paradigm. We introduce a DVP index, the Normalized Difference Phenometric Index (NDPI) that provides vegetation type-specific information at the subpixel scale by exploiting differential patterns of vegetation phenology detectable in time-series spectral vegetation index (VI) data from multispectral land imagers. We used modified soil-adjusted vegetation index (MSAVI 2 ) data from Landsat to develop the NDPI, and MSAVI 2 data from MODIS to compare its performance relative to one alternate DVP metric (difference of spring average MSAVI 2 and summer maximum MSAVI 2 ), and two simple, conventional VI metrics (summer average MSAVI 2 , summer maximum MSAVI 2 ). The NDPI in a scaled form (NDPI s ) performed best in predicting variation in perennial C 4 grass cover as estimated from landscape photographs at 92 sites (R 2 = 0.76, p < 0.001), indicating improvement over the alternate DVP metric (R 2 = 0.73, p < 0.001) and substantial improvement over the two conventional VI metrics (R 2 = 0.62 and 0.56, p < 0.001). The results suggest DVP-based methods, and the NDPI in particular, can be effective for subpixel discrimination and mapping of exposed perennial C 4 grass cover within mixed-composition landscapes of the Southwest, and potentially for monitoring of its response to drought, climate change, grazing and other factors, including land management. With appropriate adjustments, the method could potentially be used for subpixel discrimination and mapping of grass or other vegetation types in other regions where the vegetation components of the landscape exhibit contrasting seasonal patterns of phenology.

Arizona↗

Impacts of climate changes and amplified natural disturbance on global ecosystems

Natural disturbances maintain biological diversity and landscape heterogeneity and initiate ecosystem renewal and reorganization. However, the severity, frequency, and extent of many disturbances have increased substantially in recent decades as the result of anthropogenic climate change. Disturbances can be discrete, short-duration events, such as wildfires or hurricanes, or can exert persistent, cumulative stresses on an ecosystem (for example, ongoing warming of ocean or land surface temperatures). Landscape and ecosystem impacts can occur from a single disturbance, from several disturbances acting independently, or from the interactions of multiple, linked disturbances. Key, climate-related disturbances affecting global biomes and ecosystems include shifting temperature and hydrologic regimes (for example, warming surface temperatures and increasing aridity), increased frequency and magnitude of extreme events such as heatwaves, severe droughts, storms, and hurricanes, warming-induced permafrost thaw, and heightened wildfire activity and insect-caused tree mortality. For ecosystems and landscapes, the consequences of climate-amplified disturbance include forced poleward and upward movement of plant and animal species, widespread tree mortality and reduced forest productivity, changes in plant community structure and species distributions, reduced biodiversity, increased erosion, debris flows, wetland dynamism, declining sea ice extent, more frequent storm-driven tides and saltwater intrusion, and increased landscape flammability.

Book chapter↗

Steady-state forms of channel profiles shaped by debris flow and fluvial processes

Debris flows regularly traverse bedrock channels that dissect steep landscapes, but our understanding of bedrock erosion by debris flows and their impact on steepland morphology is still rudimentary. Quantitative models of steep bedrock channel networks are based on geomorphic transport laws designed to represent erosion by water-dominated flows. To quantify the impact of debris flow erosion on steep channel network form, it is first necessary to develop methods to estimate spatial variations in bulk debris flow properties (e.g., flow depth, velocity) throughout the channel network that can be integrated into landscape evolution models. Here, we propose and evaluate two methods to estimate spatial variations in bulk debris flow properties along the length of a channel profile. We incorporate both methods into a model designed to simulate the evolution of longitudinal channel profiles that evolve in response to debris flow and fluvial processes. To explore this model framework, we propose a general family of debris flow erosion laws where erosion rate is a function of debris flow depth and channel slope. Model results indicate that erosion by debris flows can explain the occurrence of a scaling break in the slope–area curve at low-drainage areas and that upper-network channel morphology may be useful for inferring catchment-averaged erosion rates in quasi-steady landscapes. Validating specific forms of a debris flow incision law, however, would require more detailed model–data comparisons in specific landscapes where input parameters and channel morphometry can be better constrained. Results improve our ability to interpret topographic signals within steep channel networks and identify observational targets critical for constraining a debris flow incision law.

Earth Surface Dynamics↗

Assessing range-wide habitat suitability for the Lesser Prairie-Chicken

Population declines of many wildlife species have been linked to habitat loss incurred through land-use change. Incorporation of conservation planning into development planning may mitigate these impacts. The threatened Lesser Prairie-Chicken ( Tympanuchus pallidicinctus ) is experiencing loss of native habitat and high levels of energy development across its multijurisdictional range. Our goal was to explore relationships of the species occurrence with landscape characteristics and anthropogenic effects influencing its distribution through evaluation of habitat suitability associated with one particular habitat usage, lekking. Lekking has been relatively well-surveyed, though not consistently, in all jurisdictions. All five states in which Lesser Prairie-Chickens occur cooperated in development of a Maxent habitat suitability model. We created two models, one with state as a factor and one without state. When state was included it was the most important predictor, followed by percent of land cover consisting of known or suspected used vegetation classes within a 5000 m area around a lek. Without state, land cover was the most important predictor of relative habitat suitability for leks. Among the anthropogenic predictors, landscape condition, a measure of human impact integrated across several factors, was most important, ranking third in importance without state. These results quantify the relative suitability of the landscape within the current occupied range of Lesser Prairie-Chickens. These models, combined with other landscape information, form the basis of a habitat assessment tool that can be used to guide siting of development projects and targeting of areas for conservation.

Colorado, Kansas, New Mexico, Oklahoma, Texas↗

Ecological influence and pathways of land use in sagebrush

Land use in sagebrush (Artemisia spp.) landscapes influences all sage-grouse (Centrocer-cus spp.) populations in western North America. Croplands and the network of irrigation canals cover 230,000 km2 and indirectly influence up to 77% of the Sage-Grouse Conservation Area and 73% of sagebrush land cover by subsidizing synanthropic predators on sage-grouse. Urbanization and the demands of human population growth have created an extensive network of con-necting infrastructure that is expanding its influence on sagebrush landscapes. Over 2,500 km 2 are now covered by interstate highways and paved roads; when secondary roads are included, 15% of the Sage-Grouse Conservation Area and 5% of existing sagebrush habitats are 2.5 km from roads. Density of secondary roads often exceeds 5 km/km 2 , resulting in widespread motorized access for recreation, creating extensive travel corridors for management actions and resource development, subsidizing predators adapted to human presence, and facilitating spread of exotic or invasive plants. Sagebrush lands also are being used for their wilderness and recreation values, including off highway vehicle use. Approximately 12,000,000 animal use months (AUM amount of forage to support one livestock unit per month) are permitted for grazing livestock on public lands in the western states. Direct effects of grazing on sage-grouse populations or sagebrush landscapes are not possible to assess from current data. However, management of lands grazed by livestock has influenced sagebrush ecosystems by vegetation treatments to increase forage and reduce sagebrush and other plant species unpalatable to livestock. Fences (2 km/km 2 in some regions), roads, and water developments to manage livestock movements further modify the landscape. Oil and gas development influences 8% of the sagebrush habitats with the highest intensities occurring in the eastern range of sage-grouse; 20% of the sagebrush distribution is indirectly influenced in the Great Plains, Wyoming Basin, and Colorado Plateau SMZs. Energy development physically removes habitat to construct well pads, roads, power lines, and pipelines; indirect effects include habitat fragmentation, soil disturbance, and facilitation of exotic plant and animal spread. More recent development of alternative energy, such as wind and geothermal, creates infrastructure in new regions of the sage-grouse distribution. Land use will continue to be a dominant stressor on sage-brush systems; its individual and cumulative effects will challenge long-term conservation of sage-grouse populations.

Book chapter↗

Modeling mountain pine beetle disturbance in Glacier National Park using multiple lines of evidence

Temperate forest ecosystems are subject to various disturbances which contribute to ecological legacies that can have profound effects on the structure of the ecosystem. Impacts of disturbance can vary widely in extent, duration and severity over space and time. Given that global climate change is expected to increase rates of forest disturbance, an understanding of these events are critical in the interpretation of contemporary forest patterns and those of the near future. We seek to understand the impact of the 1970s mountain pine beetle outbreak on the landscape of Glacier National Park and investigate any connection between this event and subsequent decades of extensive wildfire. The lack of spatially explicit data on the mountain pine beetle disturbance represents a major data gap and inhibits our ability to test for correlations between outbreak severity and fire severity. To overcome this challenge, we utilized multiple lines of evidence to model forest canopy mortality as a proxy for outbreak severity. We used historical aerial and landscape photos, reports, aerial survey data, a six year collection of Landsat imagery and abiotic data in combination with regression analysis. The use of remotely sensed data is critical in large areas where subsequent disturbance (fire) has erased some of the evidence from the landscape. Results indicate that this method is successful in capturing the spatial heterogeneity of the outbreak in a topographically complex landscape. Furthermore, this study provides an example on the use of existing data to reduce levels of uncertainty associated with an historic disturbance.

Conference Paper↗

The quantification of instream flow rights to water

Energy development of all types continues to grow in the Rocky Mountain Region of the western United States. Federal resource managers increasingly need to balance energy demands, their effects on the natural and human landscape, and public perceptions towards these issues. The Western Energy Citation Clearinghouse (WECC v.1.0), part of a suite of data and information management tools developed and managed by the Wyoming Landscape Conservation Initiative (WLCI), provides resource managers with a searchable online database of citations that covers a broad spectrum of energy and landscape related topics relevant to resource managers, such as energy sources, natural and human landscape effects, and new research, methods and models. Based on the 2011 USGS Open-file Report "Abbreviated bibliography on energy development" (Montag, et al. 2011), WECC is an extensive collection of energy-related citations, as well as categorized lists of additional online resources related to oil and gas development, best practices, energy companies and Federal agencies. WECC incorporates the powerful web services of Sciencebase 2.0, the enterprise data and information platform for USGS scientists and partners, to provide secure, role-based data management features. For example, public/unauthenticated WECC users have full search and read access to the entire energy citation collection, while authenticated WLCI data stewards can manage WECC's citation collection using Sciencebase data management forms.

Book↗

Proceedings of the fourth conference on research in California's National Parks

The papers in this proceedings were selected from the 63 presentations given at the Fourth Biennial Conference on Research in National Parks in California. The overall theme for this meeting was a recurring one: “The Integration of Research into National Park Service Resource Management Decisions.” The conference was held at the University of California, Davis, on 10-12 September 1991 and was sponsored by the National Park Service Cooperative Park Studies Unit and the Institute of Ecology at the University of California, Davis. This proceedings highlights a variety of research and resource management efforts to improve the stewardship of our most treasured landscapes. In the future, it will become increasingly more important for federal and state agencies, university scientists and students, and the public to cooperate fully to improve the quantity and quality of science and resource management programs in units of the National Park System. As many of the papers in this proceedings attest, we must look beyond the political boundaries of protected landscapes to incorporate entire ecosystems. Competing resource uses inside and outside parklands must be reexamined to weave a common thread of biological conservation. As scientists, our studies must bridge the gap from plots to landscapes and from landscapes to regions. Our studies must built on information from species and populations to ecosystems and the processes that influence them. The papers in this proceedings are modest but important contribution to those ideals. Each paper represents original research and has been peer-reviewed. Many agencies, institutions, and individuals contributed in the development of this product. In the planning stages, National Park Service Western Region scientists provided advice and assistance in structuring the conference format. University of California, Davis, graduate students, directed by Sharon Lynch, assisted with logistics at the meeting, and provided general assistance with the paper sessions. We thank the students, technicians, and support staff who labored unselfishly behind the scenes to collect data and transfer information important to the completion of each paper. Financial support for this publication was provided with funds from the Western Region Resource Management and Resource Program and the Washington Office Servicewide Publications Program of the National Park Service. We thank Denny Fenn, Stan Albright, Bruce Kilgore, Dave Cherry, Gene Hester, and Donna O'Leary of the National Park Service and Charles Goldman of the University of California, Davis, Institute of Ecology for their support throughout the development of the proceedings. Finally, we are entering into a new age of research in the U.S. Department of the Interior. Recently, plans were announced for the creation of the National Biological Survey, dedicated to improving the quantity and quality of research necessary for the wise stewardship of our natural resources and wildlands. We welcome our new partners and pledge our full support to both the new and old agencies. The papers presented here--and past research efforts in national parks--provide an important stepping stone to the future. While many of us will be transferred administratively to the National Biological Survey, our emotional ties to the National Park Service mission of preserving resources for future generations will remain with us forever.

California↗