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At least 1,225 records · Page 68Linked to original sources

Stability of steep slopes in cemented sands

The analysis of steep slope and cliff stability in variably cemented sands poses a significant practical challenge as routine analyses tend to underestimate the actually observed stability of existing slopes. The presented research evaluates how the degree of cementation controls the evolution of steep sand slopes and shows that the detailed slope geometry is important in determining the characteristics of the failure mode, which in turn, guide the selection of an appropriate stability analysis method. Detailed slope-profile cross sections derived from terrestrial lidar surveying of otherwise inaccessible cemented sand cliffs are used to investigate failure modes in weakly cemented [unconfined compressive strength ( UCS ) &lt; 30 &#x2002; kPa "> ( UCS ) < 30 kPa (UCS)<30 kPa ] and moderately cemented ( 30 &lt; UCS &lt; 400 &#x2002; kPa ) "> ( 30 < UCS < 400 kPa ) (30<UCS<400 kPa) sands and their role in the evolution of the geometry of the slopes. The results show that high-resolution slope topography, such as can be obtained with terrestrial lidar, is essential for identifying altogether different failure modes in weakly cemented (shear-mode) and moderately cemented (tensile-mode) sand slopes. Analyses show that the standard Culmann method for steep slopes is inappropriate for modeling the stability of cemented sand slopes since it tends to overpredict expected crest retreat and underestimate failure plane angle. Instead, a simplified analysis using infinite slope assumptions, but applied to a slope with finite dimensions subject to changing geometric conditions, such as toe erosion and slope steepening, is suggested for analysis of weakly cemented sand slopes. For moderately cemented sand slopes, a limit equilibrium analysis directly comparing the cliff tensile stress and cemented sand tensile strength is shown to reasonably predict failure conditions and timing as a result of either slope steepening or tensile strength loss, presumably from wetting in most cases.

Journal of Geotechnical and Geoenvironmental Engin↗

The response of spit shapes to wave-angle climates

We investigate spit formation and evolution in light of the high-wave-angle instability in shoreline shape arising from a maximizing angle for wave-driven alongshore sediment transport. Single spits emerge in a simple one-contour line numerical model that evolves the coast using morphodynamic feedbacks and a 'climate' of waves approaching the shore from variable directions. Analysis of sediment transport and shoreline stability metrics illustrate how spits can be generated, demonstrating how waves from all angles can play a role in spit formation and evolution. Simulations suggest that regardless of whether high- or low-angle waves dominate relative to the general shoreline trend, as spits extend offshore, they tend to orient themselves such that most of their coast barely experiences low-angle waves and alongshore sediment transport to the spit end is maximized. This 'graded' spit shape minimizes gradients in sediment flux, while the recurve at the spit end experiences larger gradients and a region of high-angle instability. Examining hindcast wave data, similar trends are seen along the natural example of Long Point, Lake Erie, Canada. ?? 2007 ASCE.

Conference Paper↗

Hindcasting potential hurricane impacts on rapidly changing barrier islands

Hindcasts of the coastal impact of Hurricane Ivan on Santa Rosa Island, Florida, using a storm-impact scaling model that compares hurricane-induced water levels to local dune morphology, were found to have an accuracy of 68% in predicting the occurrence of one of four impact regimes: swash, collision, overwash, and inundation. Errors were overwhelming under-predictions of the regime where the observed response was more extreme than had been expected. This is related to the evolution of the profile during the storm. Mean pre-storm dune elevations decreased by 1.9 m over the 75-km long island as most of the dunes were completely eroded during the storm. Dramatic morphologic change during a hurricane makes barrier islands more vulnerable to overwash and inundation than will be predicted based on pre-storm dune parameters. Incorporation of the timing of rising water levels relative to storm-induced profile evolution is required to improve model accuracy.

Conference Paper↗

An introduced and a native vertebrate hybridize to form a genetic bridge to a second native species

The genetic impacts of hybridization between native and introduced species are of considerable conservation concern, while the possibility of reticulate evolution affects our basic understanding of how species arise and shapes how we use genetic data to understand evolutionary diversification. By using mitochondrial NADH dehydrogenase subunit 2 (ND2) sequences and 467 amplified fragment-length polymorphism nuclear DNA markers, we show that the introduced white sucker (Catostomus commersoni) has hybridized with two species native to the Colorado River Basin - the flannelmouth sucker (Catostomus latipinnis) and the bluehead sucker (Catostomus discobolus). Hybrids between the flannelmouth sucker and white sucker have facilitated introgression between the two native species, previously isolated by reproductive barriers, such that individuals exist with contributions from all three genomes. Most hybrids had the mitochondrial haplotype of the introduced white sucker, emphasizing its pivotal role in this three-way hybridization. Our findings highlight how introduced species can threaten the genetic integrity of not only one species but also multiple previously reproductively isolated species. Furthermore, this complex three-way reticulate (as opposed to strictly bifurcating) evolution suggests that seeking examples in other vertebrate systems might be productive. Although the present study involved an introduced species, similar patterns of hybridization could result from natural processes, including stream capture or geological formations (e.g., the Bering land bridge). ?? 2008 by The National Academy of Sciences of the USA.

Proceedings of the National Academy of Sciences of↗

Polar and brown bear genomes reveal ancient admixture and demographic footprints of past climate change

Polar bears (PBs) are superbly adapted to the extreme Arctic environment and have become emblematic of the threat to biodiversity from global climate change. Their divergence from the lower-latitude brown bear provides a textbook example of rapid evolution of distinct phenotypes. However, limited mitochondrial and nuclear DNA evidence conflicts in the timing of PB origin as well as placement of the species within versus sister to the brown bear lineage. We gathered extensive genomic sequence data from contemporary polar, brown, and American black bear samples, in addition to a 130,000- to 110,000-y old PB, to examine this problem from a genome-wide perspective. Nuclear DNA markers reflect a species tree consistent with expectation, showing polar and brown bears to be sister species. However, for the enigmatic brown bears native to Alaska's Alexander Archipelago, we estimate that not only their mitochondrial genome, but also 5–10% of their nuclear genome, is most closely related to PBs, indicating ancient admixture between the two species. Explicit admixture analyses are consistent with ancient splits among PBs, brown bears and black bears that were later followed by occasional admixture. We also provide paleodemographic estimates that suggest bear evolution has tracked key climate events, and that PB in particular experienced a prolonged and dramatic decline in its effective population size during the last ca. 500,000 years. We demonstrate that brown bears and PBs have had sufficiently independent evolutionary histories over the last 4–5 million years to leave imprints in the PB nuclear genome that likely are associated with ecological adaptation to the Arctic environment.

Proceedings of the National Academy of Sciences↗

Landscape changes elevate the risk of avian influenza virus diversification and emergence in the East Asian–Australasian Flyway

Highly pathogenic avian influenza viruses (HPAIV) persistently threaten wild waterfowl, domestic poultry, and public health. The East Asian–Australasian Flyway plays a crucial role in HPAIV dynamics due to its large populations of migratory waterfowl and poultry. Over recent decades, this flyway has undergone substantial landscape changes, including both losses and gains of waterfowl habitats. These changes can affect waterfowl distributions, increase contact with poultry, and consequently alter ecological conditions that favor avian influenza virus (AIV) evolution. However, limited research has assessed these likely impacts. Here, we integrated empirical data and an individual-based model to simulate AIV transmission in migratory waterfowl and domestic poultry, including wild-to-poultry spillover and reassortment dynamics in poultry, across landscapes representing the years 2000 and 2015. We used the reassortment incidence as a proxy for ecological and transmission conditions that support viral diversification and the emergence of novel subtypes. Our simulations show that landscape change reshaped the waterfowl distribution, facilitated bird aggregation at improved habitats, increased coinfection, and raised reassortment rate by 1,593%, indicating a substantially higher potential for viral diversification and emergence. Model-generated risk maps show expanded and increased reassortment risk in southeastern China, the Yellow River Basin, and northeastern China. These findings suggest the importance of landscape change as a driver of potential AIV diversification and subtype emergence. This underscores the need for interdisciplinary approaches that integrate landscape dynamics, host movement, and viral evolution to better assess and mitigate future risk.

East Asian–Australasian Flyway↗

Does clutch size evolve in response to parasites and immunocompetence?

Parasites have been argued to influence clutch size evolution, but past work and theory has largely focused on within-species optimization solutions rather than clearly addressing among-species variation. The effects of parasites on clutch size variation among species can be complex, however, because different parasites can induce age-specific differences in mortality that can cause clutch size to evolve in different directions. We provide a conceptual argument that differences in immunocompetence among species should integrate differences in overall levels of parasite-induced mortality to which a species is exposed. We test this assumption and show that mortality caused by parasites is positively correlated with immunocompetence measured by cell-mediated measures. Under life history theory, clutch size should increase with increased adult mortality and decrease with increased juvenile mortality. Using immunocompetence as a general assay of parasite-induced mortality, we tested these predictions by using data for 25 species. We found that clutch size increased strongly with adult immunocompetence. In contrast, clutch size decreased weakly with increased juvenile immunocompetence. But, immunocompetence of juveniles may be constrained by selection on adults, and, when we controlled for adult immunocompetence, clutch size decreased with juvenile immunocompetence. Thus, immunocompetence seems to reflect evolutionary differences in parasite virulence experienced by species, and differences in age-specific parasite virulence appears to exert opposite selection on clutch size evolution.

Proceedings of the National Academy of Sciences of↗

Tertiary volcanic rocks from the southern Pannonian Basin, Croatia

The Tertiary volcanic rocks of the southern part of the Pannonian Basin are related to a sequence of tectonic events that occurred along the northern margin of the Dinarides when subduction ceased and after uplift of the region in middle Eocene time. The oldest rocks are andesites and dacites erupted during an Egerian to Eggenburgian marine transpression phase. In early Miocene time (about 17 Ma), rifting started, probably caused by the rise of the mantle, resulting in E-W extension and strike-slip faulting leading to formation of the Pannonian Basin. The initial rifting was accompanied by trachyandesitic volcanism of Karpathian age, followed by Badenian-age (15 to 13 Ma) volcanic activity producing basalts, andesites, and dacites. During these volcanic phases the Pannonian Basin, or the Paratethys, probably was periodically connected with the Mediterranean Sea. The final period of volcanism (about 9 to 7 Ma) produced alkalic basalts and basalts and occurred in fresh-water environments. These volcanic rocks have had a complex petrological evolution, related to an active continental margin. Egerian-Eggenburgian andesites and dacites can be explained by upper crustal assimilation with fractional crystallization (ACF) of primary basalt magma. The three younger associations originated by partial melting of rocks from both the upper mantle and the lower crust. Their evolution was accompanied by crustal contamination.

Carpathian Basin, Pannonian Basin↗

Occam's shadow: levels of analysis in evolutionary ecology - where to next?

Evolutionary ecology is the study of evolutionary processes, and the ecological conditions that influence them. A fundamental paradigm underlying the study of evolution is natural selection. Although there are a variety of operational definitions for natural selection in the literature, perhaps the most general one is that which characterizes selection as the process whereby heritable variation in fitness associated with variation in one or more phenotypic traits leads to intergenerational change in the frequency distribution of those traits. The past 20 years have witnessed a marked increase in the precision and reliability of our ability to estimate one or more components of fitness and characterize natural selection in wild populations, owing particularly to significant advances in methods for analysis of data from marked individuals. In this paper, we focus on several issues that we believe are important considerations for the application and development of these methods in the context of addressing questions in evolutionary ecology. First, our traditional approach to estimation often rests upon analysis of aggregates of individuals, which in the wild may reflect increasingly non-random (selected) samples with respect to the trait(s) of interest. In some cases, analysis at the aggregate level, rather than the individual level, may obscure important patterns. While there are a growing number of analytical tools available to estimate parameters at the individual level, and which can cope (to varying degrees) with progressive selection of the sample, the advent of new methods does not reduce the need to consider carefully the appropriate level of analysis in the first place. Estimation should be motivated a priori by strong theoretical analysis. Doing so provides clear guidance, in terms of both (i) assisting in the identification of realistic and meaningful models to include in the candidate model set, and (ii) providing the appropriate context under which the results are interpreted. Second, while it is true that selection (as defined) operates at the level of the individual, the selection gradient is often (if not generally) conditional on the abundance of the population. As such, it may be important to consider estimating transition rates conditional on both the parameter values of the other individuals in the population (or at least their distribution), and population abundance. This will undoubtedly pose a considerable challenge, for both single- and multi-strata applications. It will also require renewed consideration of the estimation of abundance, especially for open populations. Thirdly, selection typically operates on dynamic, individually varying traits. Such estimation may require characterizing fitness in terms of individual plasticity in one or more state variables, constituting analysis of the norms of reaction of individuals to variable environments. This can be quite complex, especially for traits that are under facultative control. Recent work has indicated that the pattern of selection on such traits is conditional on the relative rates of movement among and frequency of spatially heterogeneous habitats, suggesting analyses of evolution of life histories in open populations can be misleading in some cases.

Journal of Applied Statistics↗

Reduced swimming performance repeatedly evolves upon loss of migration in landlocked populations of Alewife

Whole-organism performance tasks are accomplished by the integration of morphological traits and physiological functions. Understanding how evolutionary change in morphology and physiology influences whole-organism performance will yield insight into the factors that shape its own evolution. We demonstrate that nonmigratory populations of alewife ( Alosa pseudoharengus ) have evolved reduced swimming performance in parallel, compared with their migratory ancestor. In contrast to theoretically and empirically based predictions, poor swimming among nonmigratory populations is unrelated to the evolution of osmoregulation and occurs despite the fact that nonmigratory alewives have a more fusiform (torpedo-like) body shape than their ancestor. Our results suggest that elimination of long-distance migration from the life cycle has shaped performance more than changes in body shape and physiological regulatory capacity.

Physiological and Biochemical Zoology↗

Determinants and consequences of dispersal in vertebrates with complex life cycles: a review of pond-breeding amphibians

Dispersal is a central process in ecology and evolution. It strongly influences the dynamics of spatially structured populations, by affecting population growth rate and local colonization-extinction processes. Dispersal can also influence evolutionary processes because it determines rates and patterns of gene flow in spatially structured populations and is closely linked to local adaptation. For these reasons, dispersal has received considerable attention from ecologists and evolutionary biologists. However, although it has been studied extensively in taxa such as birds and mammals, much less is known about dispersal in vertebrates with complex life cycles such as pond-breeding amphibians. Over the past two decades, researchers have taken an interest in amphibian dispersal and initiated both fundamental and applied studies, using a broad range of experimental and observational approaches. This body of research reveals complex dispersal patterns, causations and syndromes, with dramatic consequences for the demography and genetics of amphibian populations. In this review, our goals are to (1) redefine and clarify the concept of amphibian dispersal, (2) review current knowledge about the effects of individual (i.e., condition-dependent dispersal) and environmental (i.e., context-dependent dispersal) factors during the three stages of dispersal (i.e., emigration, immigration, transience), (3) identify the demographic and genetic consequences of dispersal in spatially structured amphibian populations, and (4) propose new research avenues to extend our understanding of amphibian dispersal. In particular, we emphasize the need to (1) quantify dispersal rate and distance rigorously using suitable model systems, (2) investigate the genetic basis and dispersal evolution patterns, and (3) examine dispersal-related eco-evolutionary dynamics. These proposed research avenues tap from the recent advances in quantitative and molecular methods and have the potential to improve our understanding of dispersal in organisms with complex life cycles.

The Quarterly Review of Biology↗

Daily nest predation rates decrease with body size in passerine birds

Body size evolution is generally framed by the benefits of being large, while costs are largely overlooked. An important putative cost of being large is the need to extend development periods, which should increase exposure to predation and potentially select against larger size. In birds, this selection pressure can be important because predation is the main source of offspring mortality and predators should more readily detect the larger nests associated with larger body sizes. Here, we show for diverse passerine birds across the world that counter to expectations, larger species suffer lower daily nest predation rates than smaller species. This pattern is consistent despite latitudinal variation in predation and does not seem to reflect a tendency of larger species to use more protected nests or less exposed nest locations. Evidence instead suggests that larger species attack a wider array of predator sizes, which could reduce predation rates in nests of large-bodied species. Regardless of the mechanism, the lower daily nest predation rates of larger species yield slightly lower predation rates over the entire development period compared with smaller species. These results highlight the importance of behavior as a mechanism to alter selection pressures and have implications for body size evolution.

The American Naturalist↗

Female sea lamprey shift orientation toward a conspecific chemical cue to escape a sensory trap

The sensory trap model of signal evolution hypothesizes that signalers adapt to exploit a cue used by the receiver in another context. Although exploitation of receiver biases can result in conflict between the sexes, deceptive signaling systems that are mutually beneficial drive the evolution of stable communication systems. However, female responses in the nonsexual and sexual contexts may become uncoupled if costs are associated with exhibiting a similar response to a trait in both contexts. Male sea lamprey ( Petromyzon marinus ) signal with a mating pheromone, 3-keto petromyzonol sulfate (3kPZS), which may be a match to a juvenile cue used by females during migration. Upstream movement of migratory lampreys is partially guided by 3kPZS, but females only move toward 3kPZS with proximal accuracy during spawning. Here, we use in-stream behavioral assays paired with gonad histology to document the transition of female preference for juvenile- and male-released 3kPZS that coincides with the functional shift of 3kPZS as a migratory cue to a mating pheromone. Females became increasingly biased toward the source of synthesized 3kPZS as their maturation progressed into the reproductive phase, at which point, a preference for juvenile odor (also containing 3kPZS naturally) ceased to exist. Uncoupling of female responses during migration and spawning makes the 3kPZS communication system a reliable means of synchronizing mate search. The present study offers a rare example of a transition in female responses to a chemical cue between nonsexual and sexual contexts, provides insights into the origins of stable communication signaling systems.

Behavioral Ecology↗

Evolutionary fire ecology: An historical account and future directions

The idea that fire acts as an evolutionary force contributing to shaping species traits started a century ago, but had not been widely recognized until very recently. Among the first to realize this force were E dward B . Poulton, R. D ale Guthrie, and E dwin V . Komarek in animals and W illis L . Jepson, W alter W . Hough, T om M . Harris, P hilip V . Wells, and R obert W . Mutch in plants. They were all ahead of their time in their evolutionary thinking. Since then, evolutionary fire ecology has percolated very slowly into the mainstream ecology and evolutionary biology; in fact, this topic is still seldom mentioned in textbooks of ecology or evolution. Currently, there is plenty of evidence suggesting that we cannot understand the biodiversity of our planet without considering the key evolutionary role of fire. But there is still research to be done in order to fully understand fire's contribution to species evolution and to predicting species responses to rapid global changes.

BioScience↗

A reference genome assembly for the continentally distributed ring-necked snake, Diadophis punctatus

Snakes in the family Colubridae include more than 2,000 currently recognized species, and comprise roughly 75% of the global snake species diversity on Earth. For such a spectacular radiation, colubrid snakes remain poorly understood ecologically and genetically. Two subfamilies, Colubrinae (788 species) and Dipsadinae (833 species), comprise the bulk of colubrid species richness. Dipsadines are a speciose and diverse group of snakes that largely inhabit Central and South America, with a handful of small-body-size genera that have invaded North America. Among them, the ring-necked snake, Diadophis punctatus , has an incredibly broad distribution with 14 subspecies. Given its continental distribution and high degree of variation in coloration, diet, feeding ecology, and behavior, the ring-necked snake is an excellent species for the study of genetic diversity and trait evolution. Within California, six subspecies form a continuously distributed “ring species” around the Central Valley, while a seventh, the regal ring-necked snake, Diadophis punctatus regalis is a disjunct outlier and Species of Special Concern in the state. Here, we report a new reference genome assembly for the San Diego ring-necked snake, D. p. similis , as part of the California Conservation Genomics Project. This assembly comprises a total of 444 scaffolds spanning 1,783 Mb and has a contig N50 of 8.0 Mb, scaffold N50 of 83 Mb, and BUSCO completeness score of 94.5%. This reference genome will be a valuable resource for studies of the taxonomy, conservation, and evolution of the ring-necked snake across its broad, continental distribution.

Journal of Heredity↗

The population history of endogenous retroviruses in mule deer (Odocoileus heminous)

Mobile elements are powerful agents of genomic evolution and can be exceptionally informative markers for investigating species and population-level evolutionary history. While several studies have utilized retrotransposon-based insertional polymorphisms to resolve phylogenies, few population studies exist outside of humans. Endogenous retroviruses are LTR-retrotransposons derived from retroviruses that have become stably integrated in the host genome during past infections and transmitted vertically to subsequent generations. They offer valuable insight into host-virus co-evolution and a unique perspective on host evolutionary history because they integrate into the genome at a discrete point in time. We examined the evolutionary history of a cervid endogenous gammaretrovirus (CrERVγ) in mule deer ( Odocoileus hemionus ). We sequenced 14 CrERV proviruses (CrERV-in1 to -in14), and examined the prevalence and distribution of 13 proviruses in 262 deer among 15 populations from Montana, Wyoming, and Utah. CrERV absence in white-tailed deer ( O. virginianus ), identical 5′ and 3′ long terminal repeat (LTR) sequences, insertional polymorphism, and CrERV divergence time estimates indicated that most endogenization events occurred within the last 200000 years. Population structure inferred from CrERVs (F ST = 0.008) and microsatellites (θ = 0.01) was low, but significant, with Utah, northwestern Montana, and a Helena herd being particularly differentiated. Clustering analyses indicated regional structuring, and non-contiguous clustering could often be explained by known translocations. Cluster ensemble results indicated spatial localization of viruses, specifically in deer from northeastern and western Montana. This study demonstrates the utility of endogenous retroviruses to elucidate and provide novel insight into both ERV evolutionary history and the history of contemporary host populations.

Montana;Utah;Wyoming↗

Replicated landscape genomics identifies evidence of local adaptation to urbanization in wood frogs

Native species that persist in urban environments may benefit from local adaptation to novel selection factors. We used double-digest restriction-side associated DNA (RAD) sequencing to evaluate shifts in genome-wide genetic diversity and investigate the presence of parallel evolution associated with urban-specific selection factors in wood frogs ( Lithobates sylvaticus ). Our replicated paired study design involved 12 individuals from each of 4 rural and urban populations to improve our confidence that detected signals of selection are indeed associated with urbanization. Genetic diversity measures were less for urban populations; however, the effect size was small, suggesting little biological consequence. Using an F ST outlier approach, we identified 37 of 8344 genotyped single nucleotide polymorphisms with consistent evidence of directional selection across replicates. A genome-wide association study analysis detected modest support for an association between environment type and 12 of the 37 F ST outlier loci. Discriminant analysis of principal components using the 37 F ST outlier loci produced correct reassignment for 87.5% of rural samples and 93.8% of urban samples. Eighteen of the 37 F ST outlier loci mapped to the American bullfrog ( Rana [Lithobates] catesbeiana ) genome, although none were in coding regions. This evidence of parallel evolution to urban environments provides a powerful example of the ability of urban landscapes to direct evolutionary processes.

Maine↗

The generation of oceanic rhyolites by crystal fractionation: the basalt-rhyolite association at Volcán Alcedo, Galápagos archipelago

Alcedo volcano is one of six shield volcanoes on Isabela Island in the western Galápagos Islands. Although Alcedo is dominantiy basaltic, it is unusual in that it also has erupted ∼1 km 3 of rhyolite. The rhyolitic phase marked a 10-fold decrease in the mass-eruption rate of the volcano, and the volcano has returned to erupting basalt. The basalts are tholeiitic and range from strongly to sparsely porphyritic. Olivine and plagiodase are the liquidus phases in the most primitive basalts. The MgO and Ni concentrations in the most primitive basalts indicate that they have undergone substantial differentiation since extraction from the mantle. The rhyolites contain the assemblage oligoclase-augite-titanomagnetite-fayalite-apatite and sparse xenoliths of quenched basalt and cumulate gabbros. Intermediate rocks are very rare, but some are apparently basaltrhyolite hybrids, and others resulted from differentiation of tholeiitic magma. Several modeling approaches and Sr-, Nd-, and O-isotopic data indicate that the rhyolites resulted from ∼ 90% fractionation (by weight) of plagiodase, augite, titanomagnetite, olivine, and apatite from the most primitive olivine tholeiite. The data are inconsistent with the rhyolites originating by crustal anatexis. The extreme Daly gap may be caused by the large increase in viscosity as the basaltic magma differentiates to intermediate and siliceous compositions; highly evolved magmas are eruptible only after they become saturated with volatiles by second boiling. The close association of the hybrid intermediate magmas and magmatic inclusions with the climactic plinian eruption indicates mixing between mafic and silicic magmas immediately before eruption. Rhyolite production was favored by the decrease in supply of basaltic magma as Alcedo was carried away from the focus of the Galápagos hotspot. A three-stage model for the magmatic evolution of a Galápagos volcano is proposed. In the first stage, the supply of basaltic magma is large. Basaltic magma continually intrudes the subcaldera magma chamber, buffering the magmas' compositional and thermal evolution. As the volcano is carried away from the basaltic source, the magma chamber is allowed to cool and differentiate, as exemplified by Alcedo's rhyolitic phase. Finally, the volcano receives even smaller influx of basalt, so a large magma chamber cannot be sustained, and the volcano shifts to isolated basaltic eruptions.

Galapagos Islands↗