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Reconnaissance sedimentology of selected tertiary exposures in the upland region bordering the Yukon Flats basin, east-central Alaska

This report summarizes reconnaissance sedimentologic and stratigraphic observations made during six days of helicopter-supported fieldwork in 2002 on Tertiary sedimentary rocks exposed in the upland region around the flanks of the Yukon Flats basin in east-central Alaska (fig. 1). This project was a cooperative effort between the Alaska Division of Geological & Geophysical Surveys (DGGS) and the U.S. Geological Survey (USGS) to investigate the geology of the basin in preparation for an assessment of the undiscovered, technically recoverable hydrocarbon resources (Stanley and others, 2004). Field observations and interpretations summarized in this report are reconnaissance level. At most, no more than a few hours were spent on the ground at any location. Measured sections included in this report are sketch sec- tions and thicknesses shown are approximate. Relatively detailed observations were made by the authors at only three locations, including The Mudbank (Hodzana River), Rampart (east bank of the Yukon River), and Bryant Creek (along the Tintina fault near the Canada border). These three locations are described first in relative detail, then followed by general descriptions of other locations.

Alaska↗

U-Pb Zircon ages from bedrock samples collected in the Tanacross D-1, and parts of the D-2, C-1, and C-2 quadrangles, Alaska

This Alaska Division of Geological & Geophysical Surveys (DGGS) Preliminary Interpretive Report presents U-Pb ages of zircons from 14 sedimentary, metamorphic, and igneous samples collected during 2017 and 2018 field investigations in the northeastern Tanacross Quadrangle, Alaska. The DGGS Northeast Tanacross project is a part of multi-year effort to investigate the geology and mineral-resource potential of the Yukon-Tanana Uplands region in collaboration with the U.S. Geological Survey. The purpose of the U-Pb isotopic study is to better understand the Devonian-to-Mississippian and Mesozoic-to-Early Paleogene episodes of magmatic and tectonic events within the Yukon-Tanana Uplands and the relationship of magmatism to the metallic mineral deposits. This area is characterized by the presence of two Late Devonian to Mississippian metamorphic assemblages-Lake George and Fortymile River (Dusel-Bacon and others, 2006; Foster, 1970). Both assemblages are composed of metasedimentary and metavolcanic rocks that have been intruded by Devonian to Eocene intrusive rocks of varying composition and texture. Paleozoic intrusive rocks are deformed and metamorphosed and include prevalent Late Devonian-Early Mississippian augen orthogneiss, herein called the Divide Mountain suite, that was emplaced into and deformed together with the Lake George assemblage (Aleinikoff and others, 1986). The Fortymile River assemblage is primarily cross-cut by Mississippian to Permian intrusive rocks that are also pervasively deformed and metamorphosed. Following Jurassic to mid-Cretaceous regional metamorphism and deformation, all metamorphic rock packages were intruded by Mid- to Late-Cretaceous volcanic and plutonic rocks (Naibert and others, 2018), some of which have known or suspected potential for gold together with silver, zinc, copper, and lead mineralization. Products included in this data release are: A summary of sample-collection methods, the laboratory report, analytical data tables, and associated metadata. All components of this data release are available on the DGGS website http://doi.org/10.14509/30465 .

Alaska↗

Environmental and human influences on trumpeter swan habitat occupancy in Alaska

Approximately 70-80% of the entire population of the Trumpeter Swan (Cygnus huccinator) depends for reproduction on wetlands in Alaska. This makes the identification of important habitat features and the effects of human interactions important for the species' long-term management. We analyzed the swan's habitat preferences in five areas throughout the state and found that swan broods occupied some wetland types, especially larger closed-basin wetlands such as lakes and ponds, at rates much higher than they occupied other wetland types, such as shrubby or forested wetlands. We also found a negative effect of transportation infrastructure on occupancy by broods in and around the Minto Flats State Game Refuge, Kenai National Wildlife Refuge, and Tetlin National Wildlife Refuge. This finding is of particular interest because much of the Minto Flats refuge has recently been licensed for oil and gas exploration and parts of the Kenai refuge have been developed in the past. We also investigated the potential effects of the shrinkage of closed-basin ponds on habitat occupancy by nesting Trumpeter Swans. We compared nesting swans' use of ponds with changes in the ponds' size and other characteristics from 1982 to 1996 and found no relationships between occupancy and changes in pond size. However, we believe that the recent and rapid growth of Trumpeter Swan populations in Alaska may become limited by available breeding habitat, and anthropogenic and climate-induced changes to the swan's breeding habitats have the potential to limit future production. ?? 2009 by The Cooper Ornithological Society. All rights reserved.

Condor↗

A puzzling migratory detour : Are fueling conditions in Alaska driving the movement of juvenile sharp -tailed sandpipers ?

Making a detour can be advantageous to a migrating bird if fuel-deposition rates at stopover sites along the detour are considerably higher than at stopover sites along a more direct route. One example of an extensive migratory detour is that of the Sharp-tailed Sandpiper ( Calidris acuminata ), of which large numbers of juveniles are found during fall migration in western Alaska. These birds take a detour of 1500–3400 km from the most direct route between their natal range in northeastern Siberia and nonbreeding areas in Australia. We studied the autumnal fueling rates and fuel loads of 357 Sharp-tailed Sandpipers captured in western Alaska. In early September the birds increased in mass at a rate of only 0.5% of lean body mass day -1 . Later in September, the rate of mass increase was about 6% of lean body mass day -1 , among the highest values found among similar-sized shorebirds around the world. Some individuals more than doubled their body mass because of fuel deposition, allowing nonstop flight of between 7100 and 9800 km, presumably including a trans-oceanic flight to the southern hemisphere. Our observations indicated that predator attacks were rare in our study area, adding another potential benefit of the detour. We conclude that the most likely reason for the Alaskan detour is that it allows juvenile Sharp-tailed Sandpipers to put on large fuel stores at exceptionally high rates.

Alaska↗

Age-specific survival of tundra swans on the lower Alaska Peninsula

The population of Tundra Swans (Cygnus columbianus columbianus) breeding on the lower Alaska Peninsula represents the southern extremity of the species' range and is uniquely nonmigratory. We used data on recaptures, resightings, and recoveries of neck-collared Tundra Swans on the lower Alaska Peninsula to estimate collar loss, annual apparent survival, and other demographic parameters for the years 1978–1989. Annual collar loss was greater for adult males fitted with either the thinner collar type (0.34) or the thicker collar type (0.15) than for other age/sex classes (thinner: 0.10, thicker: 0.04). The apparent mean probability of survival of adults (0.61) was higher than that of immatures (0.41) and for both age classes varied considerably by year (adult range: 0.44–0.95, immature range: 0.25–0.90). To assess effects of permanent emigration by age and breeding class, we analyzed post hoc the encounter histories of swans known to breed in our study area. The apparent mean survival of known breeders (0.65) was generally higher than that of the entire marked sample but still varied considerably by year (range 0.26–1.00) and indicated that permanent emigration of breeding swans was likely. We suggest that reductions in apparent survival probability were influenced primarily by high and variable rates of permanent emigration and that immigration by swans from elsewhere may be important in sustaining a breeding population at and near Izembek National Wildlife Refuge.

Alaska↗

An adenovirus linked to mortality and disease in long-tailed ducks (Clangula hyemalis) in Alaska

An adenovirus was isolated from intestinal samples of two long-tailed ducks (Clangula hyemalis) collected during a die-off in the Beaufort Sea off the north coast of Alaska in 2000. The virus was not neutralized by reference antiserum against known group I, II, or III avian adenoviruses and may represent a new serotype. The prevalence of the virus was determined in live-trapped long-tailed ducks at the mortality site and at a reference site 100 km away where no mortality was observed. Prevalence of adenovirus antibodies in serum samples at the mortality site was 86% compared to 10% at the reference site. Furthermore, 50% of cloacal swabs collected at the mortality site and only 7% of swabs from the reference site were positive for adenoviruses. In 2001, no mortality was observed at either of the study areas, and virus prevalence in both serum and cloacal samples was low, providing further evidence that the adenovirus was linked to the mortality event in 2000. The virus was used to infect long-tailed ducks under experimental conditions and resulted in lesions previously described for avian adenovirus infections and similar to those observed in long-tailed duck carcasses from the Beaufort Sea. The status of long-tailed ducks has recently become a concern in Alaska due to precipitous declines in breeding populations there since the mid-1970s. Our findings suggest that the newly isolated adenovirus is a disease agent and source of mortality in long-tailed ducks, and thus could be a contributing factor in population declines.

Alaska↗

Lichens of Izembek National Wildlife Refuge, westernmost Alaska Peninsula

One hundred eighty-two taxa of lichens including two lichen parasites are reported from Izembek National Wildlife Refuge on the tip of the Alaska Peninsula. Metasphaeria tartarina is new to North America; Scoliciosporum umbrinum is new to Alaska. Wide-ranging, arctic-alpine, and boreal species dominate the lichen flora; a coastal element is moderately represented, while amphi-Beringian species form a minor element. Epigeic lichen abundance is described along a lowland to alpine mesotopographic gradient selected to represent major landscape variation in the refuge. Of six major community types identified, three had significant lichen components.

Alaska↗

The prelaying interval of emperor geese on the Yukon-Kuskokwim Delta, Alaska

We marked 136 female Emperor Geese (Chen canagica) in western Alaska with VHF or satellite (PTT) transmitters from 1999 to 2003 to monitor their spring arrival and nest initiation dates on the Yukon Delta, and to estimate prelaying interval lengths once at the nesting area. Ninety-two females with functional transmitters returned to the Yukon Delta in the spring after they were marked, and we located the nests of 35 of these individuals. Prelaying intervals were influenced by when snow melted in the spring and individual arrival dates on the Yukon Delta. The median prelaying interval was 15 days (range = 12-19 days) in a year when snow melted relatively late, and 11 days (range = 4-16 days) in two warmer years when snow melted earlier. In years when snow melted earlier, prelaying intervals of <12 days for 11 of 15 females suggested they initiated rapid follicle development on spring staging areas. The prelaying interval declined by approximately 0.4 days and nest initiation date increased approximately 0.5 days for each day a female delayed her arrival. Thus, females that arrived first on the Yukon Delta had prelaying intervals up to four days longer, yet they nested up to five days earlier, than females that arrived last. The proximity of spring staging areas on the Alaska Peninsula to nesting areas on the Yukon Delta may enable Emperor Geese to alter timing of follicle development depending on annual conditions, and to invest nutrients acquired from both areas in eggs during their formation. Plasticity in timing of follicle development is likely advantageous in a variable environment where melting of snow cover in the spring can vary by 2-3 weeks annually. ?? The Cooper Ornithological Society 2006.

Condor↗

Factors influencing nest survival and productivity of Red-throated Loons ( Gavia stellata ) in Alaska

Red-throated Loon ( Gavia stellata ) numbers in Alaska have fluctuated dramatically over the past 3 decades; however, the demographic processes contributing to these population dynamics are poorly understood. To examine spatial and temporal variation in productivity, we estimated breeding parameters at 5 sites in Alaska: at Cape Espenberg and the Copper River Delta we estimated nest survival, and at 3 sites within the Yukon-Kuskokwim Delta we estimated nest survival and productivity. Nest survival varied broadly among sites and years; annual estimates (lower, upper 95% confidence interval) ranged from 0.09 (0.03, 0.29) at Cape Espenberg in 2001 to 0.93 (0.76, 0.99) at the Copper River Delta in 2002. Annual variation among sites was not concordant, suggesting that site-scale factors had a strong influence on nest survival. Models of nest survival indicated that visits to monitor nests had a negative effect on nest daily survival probability, which if not accounted for biased nest survival strongly downward. The sensitivity of breeding Red-throated Loons to nest monitoring suggests other sources of disturbance that cause incubating birds to flush from their nests may also reduce nest survival. Nest daily survival probability at the Yukon-Kuskokwim Delta was negatively associated with an annual index of fox occurrence. Survival through the incubation and chick-rearing periods on the Yukon-Kuskokwim Delta ranged from 0.09 (0.001, 0.493) to 0.50 (0.04, 0.77). Daily survival probability during the chick-rearing period was lower for chicks that had a sibling in 2 of 3 years, consistent with the hypothesis that food availability was limited. Estimates of annual productivity on the Yukon-Kuskokwim Delta ranged from 0.17 to 1.0 chicks per pair. Productivity was not sufficient to maintain population stability in 2 of 3 years, indicating that nest depredation by foxes and poor foraging conditions during chick rearing can have important effects on productivity.

Alaska↗

Dungeness crab, Cancer magister , do not extrude eggs annually in southeastern Alaska: An in situ study

The reproductive biology of female Dungeness crabs was studied with crab-pot and dive-transect sampling in five bays within or near Glacier Bay National Park and Preserve, southeastern Alaska, in April and September yearly from 1992 to 1998. A large percentage of nonovigerous, mature females was found in April, a time when females were expected to be brooding eggs that hatch in May and June. Our study examined differences between ovigerous and nonovigerous females collected in April and September samples to corroborate our previous laboratory study in which we found nonannual egg extrusion among Dungeness crabs. Seasonal differences in the catches of ovigerous and nonovigerous females, crab sizes, shell condition, and appendage injury were examined. Additionally, all crabs collected from two bays were tagged beginning in the fall of 1995; tagging was conducted twice annually. Our pot and dive data indicate that females, particularly larger ones, do not extrude eggs annually. Larger females have lower molting probabilities, which limits mating potential and increases reliance on stored sperm. The tagging study confirmed that at least some females do not extrude eggs in one year and then extrude eggs at a later time without molting, thus skipping at least one reproductive season. A reproductive cycle of Dungeness crabs in Alaska is introduced which includes earlier egg extrusion by larger females and nonannual egg extrusion.

Journal of Crustacean Biology↗

Fire behavior, weather, and burn severity of the 2007 anaktuvuk river tundra fire, North Slope, Alaska

In 2007, the Anaktuvuk River Fire (ARF) became the largest recorded tundra fire on the North Slope of Alaska. The ARF burned for nearly three months, consuming more than 100,000 ha. At its peak in early September, the ARF burned at a rate of 7000 ha d -1 . The conditions potentially responsible for this large tundra fire include modeled record high summer temperature and record low summer precipitation, a late-season high-pressure system located over the Beaufort Sea, extremely dry soil conditions throughout the summer, and sustained southerly winds during the period of vegetation senescence. Burn severity mapping revealed that more than 80% of the ARF burned at moderate to extreme severity, while the nearby Kuparuk River Fire remained small and burned at predominantly (80%) low severity. While this study provides information that may aid in the prediction of future large tundra fires in northern Alaska, the fact that three other tundra fires that occurred in 2007 combined to burn less than 1000 ha suggests site specific complexities associated with tundra fires on the North Slope, which may hamper the development of tundra fire forecasting models.

Arctic, Antarctic, and Alpine Research↗

Ordovician "sphinctozoan" sponges from Prince of Wales Island, southeastern Alaska

A faunule of silicified hypercalcified "sphinctozoan" sponges has been recovered from a clast of Upper Ordovician limestone out of the Early Devonian Karheen Formation on Prince of Wales Island in southeastern Alaska. Included in the faunule are abundant examples of the new genus Girtyocoeliana , represented by Girtyocoeliana epiporata (Rigby and Potter), and Corymbospongia adnata Rigby and Potter, along with rare Corymbospongia amplia n. sp., and Girtyocoelia (?) sp., plus common Amblysiphonella sp. 1 and rare Amblysiphonella(?) sp. 2. The assemblage is similar to that from Ordovician clasts from the eastern Klamath Mountains of northern California. This indicates that the Alexander terrane of southeastern Alaska is related paleogeographically to the lithologically and paleontologically similar terrane of the eastern Klamath Mountains. This lithology and fossil assemblage of the clast cannot be tied to any currently known local rock units on Prince of Wales Island. Other clasts in the conglomerate appear to have been locally derived, so it is inferred that the limestone clasts were also locally derived, indicating the presence of a previously undocumented Ordovician limestone unit on northern Prince of Wales Island.

Journal of Paleontology↗

Emsian (late Early Devonian) sponges from west-central and south-central Alaska

Relatively common specimens of the hypercalcified agelasiid sponge Hormospongia labyrinthica Rigby and Blodgett, 1983 and specimens of associated species of Hormospongia have been previously reported from Emsian and Eifelian stratigraphic units at several localities in south-central and southeastern Alaska (Rigby and Blodgett, 1983). Those sponges were first described from the type section of the Eifelian Cheeneetnuk Limestone in the McGrath A-5 quadrangle. Since then several additional specimens of Hormospongia labyrinthica have also been collected from a new locality in the Talkeetna C-6 quadrangle in southcentral Alaska (Figs. 1, 2.1), and are documented here.

Alaska↗

Post-fledging movements of juvenile Common Mergansers (mergus merganser) in Alaska as inferred by satellite telemetry

We implanted satellite transmitters into eight juvenile Common Mergansers to investigate post-fledging movements from their natal river in southcentral Alaska. Subsequently, they moved widely throughout portions of western and southcentral Alaska up to 750 km from their natal areas during fall and winter months. Transmitters of two birds (one male and one female) continued to send location data into their second year and allowed us to determine the location and timing of the flightless molt period for each bird. Overall, our data suggest that juvenile Common Mergansers range widely immediately after fledging, that second year males and females may differ in their movement patterns, and that these movements have implications for population genetic structure of this species.

Waterbirds↗

Historic and contemporary mercury exposure and potential risk to yellow-billed loons ( Gavia adamsii ) breeding in Alaska and Canada

The Yellow-billed Loon (Gavia adamsii) is one of the rarest breeding birds in North America. Because of the small population size and patchy distribution, any stressor to its population is of concern. To determine risks posed by environmental mercury (Hg) loads, we captured 115 Yellow-billed Loons between 2002 and 2012 in the North American Arctic and sampled their blood and/or feather tissues and collected nine eggs. Museum samples from Yellow-billed Loons also were analyzed to examine potential changes in Hg exposure over time. An extensive database of published Hg concentrations and associated adverse effects in Common Loons (G. immer) is highly informative and representative for Yellow-billed Loons. Blood Hg concentrations reflect dietary uptake of methylmercury (MeHg) from breeding areas and are generally considered near background levels if less than 1.0 &micro;g/g wet weight (ww). Feather (grown at wintering sites) and egg Hg concentrations can represent a mix of breeding and wintering dietary uptake of MeHg. Based on Common Loon studies, significant risk of reduced reproductive success generally occurs when adult Hg concentrations exceed 2.0 &micro;g/g ww in blood, 20.0 &micro;g/g fresh weight (fw) in flight feathers and 1.0 &micro;g/g ww in eggs. Contemporary mercury concentrations for 176 total samples (across all study sites for 115 Yellow-billed Loons) ranged from 0.08 to 1.45 &micro;g/g ww in blood, 3.0 to 24.9 &micro;g/g fw in feathers and 0.21 to 1.23 &micro;g/g ww in eggs. Mercury concentrations in blood, feather and egg tissues indicate that some individual Yellow-billed Loons in breeding populations across North America are at risk of lowered productivity resulting from Hg exposure. Most Yellow-billed Loons breeding in Alaska overwinter in marine waters of eastern Asia. Although blood Hg concentrations from most breeding loons in Alaska are within background levels, some individuals exhibit elevated feather and egg Hg concentrations, which likely indicate the uptake of MeHg originating from eastern Asia. Feather Hg concentrations tended to be highest in individuals overwintering farthest west (closer to Asia). A retrospective analysis of museum specimens (n = 25) found a two-fold increase in Yellow-billed Loon feather Hg concentrations from the pre-1920s (as early as 1845) to the present. The projected increase in Hg deposition (approximately four-fold by 2050) along with the uncertainty of Hg being released through the thawing of permafrost and Arctic sea ice suggest that Hg body burdens in Yellow-billed Loons may increase. These findings indicate that Hg is a current and potentially increasing environmental stressor for the Yellow-billed Loon and possibly other Nearctic-Palearctic migrant birds.

Alaska↗

eDNA Metabarcoding Analyses of Diet in Yellow-Billed Loons of Northern Alaska

Environmental DNA is a burgeoning tool used to address wide-ranging scientific questions, including determining diets of difficult-to-sample predators. Loons are large piscivorous diving birds that capture and consume prey underwater, making it nearly impossible to visually determine their diet via observation alone. Identifying species' diets is important for understanding basic life history traits, and revealing key prey species can clarify species' roles in complex trophic webs, aid in understanding population and community dynamics, and help identify critical habitat for protection. Current information about loon diet is largely anecdotal, and traditional non-observational methods for quantifying loon diet have limitations. Analysis of eDNA from loon feces may provide biologists with a non-invasive technique for determining diet without negative sampling effects, and with increased resolution as compared to other techniques. We surveyed lakes in two areas of northern Alaska for Yellow-billed Loons ( Gavia adamsii ). Loon fecal samples were collected opportunistically from latrine sites without disturbing any animals and analyzed using novel marker sets to determine loon species and diet. Fish species were detected in all fecal samples, the most common being Alaska blackfish ( Dallia pectoralis ), and ninespine stickleback ( Pungitius pungitius ). This research demonstrates that eDNA metabarcoding analyses of loon fecal samples can determine the specific loon species that deposited the feces and characterize the piscine portion of their diet with limited disturbance to the animals.

Alaska↗

Liverworts from Attu Island, Near Islands, Aleutian Islands, Alaska (USA) with comparison to the Commander Islands (Russia)

The liverwort flora of Attu Island, the westernmost Aleutian Island in the United States, was studied to assess species diversity in the hyperoceanic sector of the northern boreal subzone. The field study was undertaken in sites selected to represent a spectrum of environmental variation, primarily within the eastern part of the island. Data were analyzed using our own collections on Attu Island, supplemented with information from published reports to compare bryophyte distribution patterns at three levels, the Northern Hemisphere, North America, the Commander Islands of Russia, and Alaska. A total of 112 liverworts were identified and a substantial number, 34 species (30%), were new reports from Attu Island and one was new to Alaska. Geographic elements dominating the flora included arctomontane (26%), arctoboreomontane (23%), montane (20%), and boreal (14%) species, while arctic species were almost absent (1%). The liverworts of the Attu Island-Commander Islands region were widespread species with over 70% circumpolar, or nearly circumpolar; nevertheless large gaps were present in some of their distributions with a floristic depression in liverwort distribution between Attu and the Commander Islands.

Alaska↗

Seismic velocity models for the Denali fault zone along the Richardson Highway, Alaska

Crustal-scale seismic-velocity models across the Denali fault zone along the Richardson Highway show a 50-km-thick crust, a near vertical fault trace, and a 5-km-wide damage zone associated with the fault near Trans-Alaska Pipeline Pump Station 10, which provided the closest strong ground motion recordings of the 2002 Denali fault earthquake. We compare models, derived from seismic reflection and refraction surveys acquired in 1986 and 1987, to laboratory measurements of seismic velocities for typical metamorphic rocks exposed along the profiles. Our model for the 1986 seismic reflection profile indicates a 5-km-wide low-velocity zone in the upper 1 km of the Denali fault zone, which we interpret as fault gouge. Deeper refractions from our 1987 line image a 40-km wide, 5-km-deep low-velocity zone along the Denali fault and nearby associated fault strands, which we attribute to a composite damage zone along several strands of the Denali fault zone and to the obliquity of the seismic line to the fault zone. Our velocity model and other geophysical data indicate a nearly vertical Denali fault zone to a depth of 30 km. After-shocks of the 2002 Denali fault earthquake and our velocity model provide evidence for a flower structure along the fault zone consisting of faults dipping toward and truncated by the Denali fault. Wide-angle reflections indicate that the crustal thickness beneath the Denali fault is transitional between the 60-km-thick crust beneath the Alaska Range to the south, and the extended, 30-km-thick crust of the Yukon-Tanana terrane to the north.

Alaska↗