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Unique challenges facing Southwestern tribes: Chapter 17

Executive Summary When considering climate change, risks to Native American lands, people, and cultures are noteworthy. Impacts on Native lands and communities are anticipated to be both early and severe due to their location in marginal environments. Because Native American societies are socially, culturally, and politically unique, conventional climate change adaptation planning and related policies could result in unintended consequences or conflicts with Native American governments, or could prove to be inadequate if tribal consultation is not considered. Therefore, it is important to understand the distinct historical, legal, and economic contexts of the vulnerability and adaptive capacity of Southwestern Native American communities. The key messages presented in this chapter are: • Vulnerability of Southwestern tribes is higher than that for most groups because it is closely linked to endangered cultural practices, history, water rights, and socio-economic and political marginalization, characteristics that most Indigenous people share. (high confidence) • Very little data are available that quantify the changes that are occurring or that establish baseline conditions for many tribal communities. Additional data are crucial for understanding impacts on tribal lands for resource monitoring and scientific studies. (high confidence) • The scant data available indicate that at least some tribes may already be experiencing climate change impacts. (medium confidence) • Tribes are taking action to address climate change by instituting climate-change mitigation initiatives, including utility-scale, alternative-energy projects, and energy-conservation projects. Tribes are also evaluating their existing capacity to engage in effective adaptation planning, even though financial and social capital is limited.

Book chapter↗

Present investigations of radioactive raw materials by the Geological Survey and a recommended program for future work

The Geological Survey's program of investigation of radioactive raw materials is presented herewith under present investigations, plans for future investigations, plan of operation, and cost of operation. This report was prepared at the request of the Atomic Energy Commission. Present investigations are summarized to show the scope of the present Trace Elements program, grouping individual projects into related types of investigations. Plans for future investigations on an expanded scale are outlined. These should provide sufficient data and knowledge of the occurrence and availability of uranium, thorium, and related elements, to permit a more complete evaluation of domestic resources. Reconnaissance projects are designed to discover possible new sources of uranium and thorium and to select areas and materials warranting further investigation. Typical projects leading to the estimation of reserves are the investigation of the carnotite ores of the Colorado Plateau by geologic mapping, exploratory drilling, and related research, and investigation of asphaltic sandstone in Emery County, Utah. Extensive research will be undertaken to establish the principles governing the geological and geochemical relations of uranium, thorium, and associated elements as an essential guide in appraising domestic resources. Particular emphasis will be placed on phosphatic rocks and black shales which offer ultimate resources of uranium far greater than carnotite ores. All the foregoing investigations will be accompanied by chemical, gephysical, and mineralogical research and analytical work. Under plan of operation is discussed the organization of the Trace Elements Unit, space requirements for laboratory and office, the scheduling of investigations, and other related problems. The proposed scheduling of work calls for approximately 109, 173, and 203 man years in fiscal years 1948, 1949, and 1950 respectively. Definite plans have been formulated only for the next three fiscal years, by which time it is assumed the program will reach stable proportions or can be altered as experience dictates. Under cost of operation is set forth the funds available in fiscal year 1947, the status of funds transferred from Atomic Services (14-217/80920), and funds necessary in succeeding fiscal years. The estimate for fiscal year 1948 inclues a non-recurring item of $1,025,000 for establishing adequate laboratories for chemical, physical, spectrographic and mineralogic research and analytical work. The total funds required in fiscal years 1948, 1949, and 1950 to support the proposed program will be $2,440,000, $2,161,000 and $2,198,000 respectively. The Geological survey anticipates contributing from its appropriation in fiscal years 1948, 1949 and 1950 approximately $243,000, $350,000, and $400,000 respectively; the balance of the necessary funds to be contributed by the Atomic Energy Commission in fiscal years 1948, 1949, and 1950 will be approximately $2,196,900, $1,811,000, and $1,798,000 respectively.

Trace Elements Investigations↗

Guide for benthic invertebrate studies in support of Natural Resource Damage Assessment and Restoration

This guide is intended to assist with characterizing injury to freshwater benthic macroinvertebrates (BMIs) in Natural Resource Damage Assessment and Restoration (NRDAR) cases. The contents are narrowly focused on insects, crustaceans, snails, and other invertebrate fauna that are typically considered part of BMI communities and are not intended to address studies of injury to larger benthic taxa such as freshwater mussels, crayfish, or benthic fishes or amphibians. Although some percentage of the community functions as predators, BMIs are predominantly primary consumers (for example, scrapers, shredders, and filterer/gatherer feeding groups) that play an essential role in converting carbon and nitrogen from plant tissues into animal biomass for higher-order consumers, especially in flowing waters. Aquatic contaminants can disrupt the quantity and quality of energy transferred (ecosystem function) by reducing invertebrate biomass and diversity. Additionally, the accumulation of toxic residues in invertebrate tissues may be a source of exposure leading to adverse effects in higher trophic levels. The goal of NRDAR BMI assessments is to establish direct linkages of contaminant exposure to injuries reflected by changes in community structure (for example, reduced density and taxa richness) or by effects at the individual population level (for example, survival, growth, and reproduction). BMIs are infrequently the U.S. Department of Interior (DOI)-managed resource in a NRDAR case, with managed resources more frequently including migratory birds, fish, or other insectivorous vertebrates. Therefore, it is critical to have clearly defined objectives for evaluating BMIs and an understanding of how invertebrate data relate to the quantification of injuries to the DOI-managed resource. This guide is intended to assist decisions on whether or not to proceed with BMI studies, use of existing information and data for screening purposes, and what types of studies can support a BMI-injury determination. This document is intended to provide general considerations and best practices for assessing BMIs. Relevant guidance and references are listed throughout the report as sources for specific methods and analysis.

Open-File Report↗

Resource allocation in offspring provisioning: An evaluation of the conditions favoring the evolution of matrotrophy

We used analytic and simulation models to determine the ecological conditions favoring evolution of a matrotrophic fish from a lecithotrophic ancestor given a complex set of trade‐offs. Matrotrophy is the nourishment of viviparous embryos by resources provided between fertilization and parturition, while lecithotrophy describes embryo nourishment provided before fertilization. In fishes and reptiles, embryo nourishment encompasses a continuum from solely lecithotrophic to primarily matrotrophic. Matrotrophy has evolved independently from lecithotrophic ancestors many times in many groups. We assumed matrotrophy increased the number of offspring a viviparous female could gestate and evaluated conditions of food availability favoring lecithotrophy or matrotrophy. The matrotrophic strategy was superior when food resources exceeded demand during gestation but at a risk of overproduction and reproductive failure if food intake was limited. Matrotrophic females were leaner during gestation than lecithotrophic females, yielding shorter life spans. Our models suggest that matrotrophic embryo nourishment evolved in environments with high food availability, consistently exceeding energy requirements for maintaining relatively large broods. Embryo abortion with some resorption of invested energy is a necessary preadaptation to the evolution of matrotrophy. Future work should explore trade‐offs of age‐specific mortality and reproductive output for females maintaining different levels of fat storage during gestation.

American Naturalist↗

Aquatic pollution increases use of terrestrial prey subsidies by stream fish

Stream food webs are connected with their riparian zones through cross-ecosystem movements of energy and nutrients. The use and impact of terrestrial subsidies on aquatic consumers is determined in part by in situ biomass of aquatic prey. Thus, stressors such as aquatic pollutants that greatly reduce aquatic secondary production could increase the need for and reliance of stream consumers on terrestrial resource subsidies. To test this hypothesis, we surveyed stream fish, their diets and resource availability in 16 subalpine streams over a regional gradient of trace metals known to strongly impact aquatic insect communities (i.e. fish prey) in the Colorado Rocky Mountains, USA. Fish increased their reliance on terrestrial insect prey as stream metals increased. Relative biomass of terrestrial insects in stomach contents of Brook and Brown Trout increased with respect to aquatic insect biomass and total stomach contents. Drifting insect biomass showed a declining trend for aquatic, but not terrestrial insects, over the metal gradient. Trout densities were unrelated to metal concentrations in streams where we found fish. Synthesis and applications . Our results indicate that diets of aquatic consumers can become more terrestrial as aquatic stressors that limit in situ food production increase and that these subsidies may compensate for loss of aquatic resources. This work implies an important connection between preserving aquatic–terrestrial linkages and management of fish populations in stressed watersheds. Specifically, intact riparian zones and aquatic–terrestrial linkages are likely to be important for maintaining trout production in streams with moderate metal contamination.

Colorado↗

Reaeration-coefficient measurements of 10 small streams in Wisconsin using radioactive tracers : with a section on the energy-dissipation model

Reaeration-rate coefficients were measured for 10 small streams in Wisconsin using the radioactive-tracer method. The coefficients ranged from 2.06 to 55.2 per day (base e at 25 degrees Celsius). Stream discharges ranged from 0.3 to 37.0 cubic feet per second, most discharges being less than 10 cubic feet per second. Data also were collected for evaluation of the energy-dissipation model. The escape coefficient of 0.090 per foot at 25 degrees Celsius may be used for small streams similar to those studied. (Woodward-USGS).

Water-Resources Investigations Report↗

Hydrogeology and sources of water to select springs in Black Canyon, south of Hoover Dam, Lake Mead National Recreation Area, Nevada and Arizona

Springs in Black Canyon of the Colorado River, directly south of Hoover Dam in the Lake Mead National Recreation Area, Nevada and Arizona, are important hydrologic features that support a unique riparian ecosystem including habitat for endangered species. Rapid population growth in areas near and surrounding Black Canyon has caused concern among resource managers that such growth could affect the discharge from these springs. The U.S. Geological Survey studied the springs in Black Canyon between January 2008, and May 2014. The purposes of this study were to provide a baseline of discharge and hydrochemical data from selected springs in Black Canyon and to better understand the sources of water to the springs. Various hydrologic, hydrochemical, geochemical, and geologic data were collected and analyzed during this study. More than 100 hydrologic sites consisting of springs, seeps, pools, rivers, reservoirs, and wells were investigated, and measurements were taken at 75 of these sites. Water levels were measured or compiled for 42 wells and samples of water were collected from 36 unique sites and submitted for laboratory analyses of hydrochemical constituents. Measurements of discrete discharge were made from nine unique spring areas and four sites in Black Canyon were selected for continuous monitoring of discharge. Additionally, samples of rock near Hoover Dam were collected and analyzed to determine the age of spring deposits. Results of hydrochemical analyses indicate that discharge from springs in Black Canyon is from two sources: (1) Lake Mead, and (2) a local and (or) regional source. Discharge from springs closest to Hoover Dam contains a substantial percentage (>50 percent) of water from Lake Mead. This includes springs that are between Hoover Dam and Palm Tree Spring. Discharge from springs south of Palm Tree Spring contains a substantial percentage (>50 percent) of the water that is believed to come from a combination of other local and regional sources, although the exact location and nature of these sources is not clear. The unique hydrochemistry of some springs, such as Bighorn Sheep Spring and Latos Pool, suggests that little if any water discharging from these springs comes from Lake Mead. Geochronological results of spring deposits at several sites near Hoover Dam indicate that most deposits are young and likely formed after the construction of Hoover Dam. Several major faults, including the Salt Cedar Fault and the Palm Tree Fault, play an important role in the movement of groundwater. Groundwater may move along these faults and discharge where faults intersect volcanic breccias or fractured rock. Vertical movement of groundwater along faults is suggested as a mechanism for the introduction of heat energy present in groundwater from many of the springs. Groundwater altitudes in the study area indicate a potential for flow from Eldorado Valley to Black Canyon although current interpretations of the geology of this area do not favor such flow. If groundwater from Eldorado Valley discharges at springs in Black Canyon then the development of groundwater resources in Eldorado Valley could result in a decrease in discharge from the springs. Geology and structure indicate that it is not likely that groundwater can move between Detrital Valley and Black Canyon. Thus, the development of groundwater resources in Detrital Valley may not result in a decrease in discharge from springs in Black Canyon.

Arizona, Nevada↗

Studying the Earth from space

Space age technology contains a key to increased knowledge about the Earth's resources; this key is remote sensing detecting the nature or condition of something without actually touching it. An early and still most useful form of remote sensing is photography which records the scene, as man sees it, on film sensitive to that part of the electromagnetic spectrum called visible energy. Electromagnetic energy travels in waves of various lengths; most are invisible to the human eye. Wavelengths progressively longer than those that the eye can see are infrared and microwave. Wavelengths progressively shorter than the eye can see are ultraviolet, X-rays, and gamma rays.

General Information Product↗

High-resolution seismic-reflection profiling data from the inner continental shelf of southeastern Massachusetts

Six hundred-seventy kilometers of closely spaced high-resolution seismic-­reflection data have been collected from eastern Rhode Island Sound and Vineyard Sound, Mass, by the U.S. Geological Survey in cooperation with the Massachusetts Department of Public Works. These data were obtained during the June 1975 cruise of the R/V ASTERIAS as part of a continuing regional study of the Massachusetts offshore area to assess potential mineral resources, to evaluate environmental impact of mining of resources and of offshore disposal of solid waste and harbor dredge-spoil materials, and to map the offshore geology and shallow structure. The data were obtained by using a surface-towed EG&G Unit Pulse Boomer* (300 joules: 400 Hz-8kHz frequency) sound source. Reflected acoustic energy was detected by a 4.6-m, a-element hydrophone array, was amplified, was actively filtered (400 Hz-4kHz bandpass), and was graphically displayed on an EPC* dry paper recorder at a 0.25-second sweep rate. System resolution was generally 1 to 1.5 m. Navigational control was provided by Loran C (posi­tional accuracy within 0.2 km) and was supplemented by radar and visual fixes. Positional information was logged at 15-minute intervals and at major course changes. The original records may be examined at the Data Library, U.S. Geological Survey, Woods Hole, MA 02543. Microfilm copies of the data are available for purchase from the National Geophysical and Solar-Terrestrial Data Center (NGSDC), Boulder, CO 80302.

Open-File Report↗

Assessment of dreissenid biodeposits as a potential food resource for invasive Asian carp

Silver carp ( Hypophthalmichthys molitrix ) and bighead carp ( H. nobilis ) are poised to invade the Laurentian Great Lakes. Zebra mussels ( Dreissena polymorpha ) and quagga mussels ( D. rostriformis bugensis ) have shifted nutrient pathways towards the benthos, partly through deposition of feces and rejected food particles called biodeposits. When biodeposit material was fed to bighead and silver carp, they fed on the material, but on average lost weight. Energy density between fed and unfed fish did not differ, but a few individual fish did gain weight on the biodeposits diet. Our results demonstrate that biodeposits might be considered a supplemental food for bigheaded carps.

Michigan↗

Petroleum systems succeed play basis in Appalachian basin resource estimate

The US Geological Survey (USGS) periodically conducts subjective probabilistic assessments of the technically recoverable undiscovered hydrocarbon resources of the US and of the world. In addition, the USGS prepares forecasts of that portion of the technically recoverable resources that may be economic under specified conditions of supply, demand, and price. Depending on priorities, regional hydrocarbon assessments of the US are revised every 5 to 10 years. These assessments of undiscovered hydrocarbons supplement the data on hydrocarbon reserves that are reported annually by the US Department of Energy, Energy Information Administration. In between assessments, USGS assessment geologists conduct research and compile geologic and production data that may be used to improve future assessments. This new information commonly effects changes in the way the USGS defines "plays" or "assessment units" from assessment to assessment. Furthermore, USGS assessment methodology is in a constant state of evolution and changes to some degree from assessment to assessment.

Oil & Gas Journal↗

Summary appraisals of the nation's ground-water resources – Pacific Northwest region

The Pacific Northwest Region's ground-water reservoirs are capable of providing large additional fresh-water supplies; these reservoirs become more important as undeveloped surface-storage sites and unapportioned surface-water supplies dwindle. Withdrawals of fresh water from all surface and underground sources are increasing; they may rise from the rate of 30 billion gallons per day in 1970 to about 60 billion gallons per day in 2020. By 1975 the withdrawal of ground water had increased 70 percent over the 1970 rate and accounted for 22 percent of total fresh-water withdrawal. Substantial increases in ground-water withdrawal must continue if projected water demands are to be met in the future. Large variations exist in the availability of and needs for ground water in the region, largely because of the great variety of landforms, climate, and earth materials. More than one-half the region is underlain by rock materials capable of yielding ground water to wells at moderate to large rates; six extensive areas are identified as major ground-water reservoirs. The most significant current problems pertaining to one or more of these ground-water reservoirs are: (1) Progressively declining water levels; (2) water-quality problems; (3) waterlogging; (4) inadequate information; and (5) competition for available supplies. In the future these same problems are expected to persist and generally worsen (especially water-quality deterioration) in most of the major reservoir areas. Management opportunities in the region include: (1) Development of new supplies and additional uses of ground water; (2) protection and enhancement of water quality; (3) reduction of waterlogging; (4) energy development from some ground-water reservoirs; (5) improving access to the ground water; (6) increased use of underground space for storage and disposal; and (7) greater use of advanced management and conservation techniques. Conjunctive use of surface and ground water to provide greater available supplies probably is the most promising water-management opportunity. However, if the full potential of the ground-water resources is to be realized, important constraints, including present water-right structures and serious deficiencies in information, must be overcome.

Idaho, Montana, Oregon, Washington↗

Water quality and landscape processes of four watersheds in eastern Puerto Rico

Humid tropical regions occupy about a quarter of Earth's land surface, yet they contribute a substantially higher fraction of the water, solutes, and sediment discharged to the world's oceans. Nearly half of Earth's population lives in the tropics, and development stresses can potentially harm soil resources, water quality, and water supply and in addition increase landslide and flood hazards. Owing to Puerto Rico's steep topography, low water storage capacity, and dependence on trade-wind precipitation, the island's people, ecosystems, and water supply are vulnerable to extreme weather such as hurricanes, floods, and droughts. Eastern Puerto Rico offers a natural laboratory for separating geologic and land-cover influences from regional- and global-scale influences because of its various bedrock types and the changing land cover surrounding intact, mature forest of the Luquillo Experimental Forest. Accordingly, a multiyear assessment of hydrological and biogeochemical processes was designed to develop an understanding of the effects of these differences on local climate, streamflow, water quality, and ecosystems, and to form the basis for a long-term and event-based program of climate and hydrologic monitoring. Because infrequent, large storms play a major role in this landscape, we focused on high-runoff events, sampling 263 storms, including all major hurricanes from 1991 through 2005. The largest storms have profound geomorphic consequences, such as landslides, debris flows, deep gullying on deforested lands, excavation and suspension of sediment in stream channels, and delivery of a substantial fraction of annual stream sediment load. Large storms sometimes entrain ocean foam and spray causing high concentrations of seasalt-derived constituents in stream waters during the storm. Past deforestation and agricultural activities in the Cayaguás and Canóvanas watersheds accelerated erosion and soil loss, and this material continues to be remobilized during large storms. Nearly 5,000 routine and event samples were analyzed for parameters that allow determination of denudation rates based on suspended and dissolved loads; 860 of these samples were analyzed for a comprehensive suite of chemical constituents. The rivers studied are generally similar in water-quality characteristics, and windward or leeward aspect appears to exert a stronger influence on water quality than geology or land cover. Of samples analyzed for comprehensive chemistry and for sediment, 543 were collected at runoff rates greater than 1 millimeter per hour, 256 at rates exceeding 10 millimeters per hour, and 3 at rates exceeding 90 millimeters per hour. Streams have rarely been sampled during events with such high runoff rates. Rates of physical and chemical weathering are especially high, and physical denudation rates, forested watersheds included, are considerably greater than is expected for a steady-state system. The elevated physical erosion drives an increased particulate organic carbon flux, one that is large, important to the carbon cycle, and sustainable, because soil-carbon regeneration is rapid. The 15-year Water, Energy, and Biogeochemical Budget dataset, which includes discharge, field parameters, suspended sediment, major cations and anions, and nutrients, is available from the U.S. Geological Survey's National Water Information System (http://waterdata.usgs.gov/nwis). The dataset provides a baseline for characterizing future environmental change and will improve our understanding of the interdependencies of land, water, and biological resources and their responses to changes in climate and land use. Because eastern Puerto Rico resembles many tropical regions in terms of geology and patterns of development, implications from this study are transferable to other tropical regions facing deforestation, rapid land-use change, and climate change.

Professional Paper↗

Are waterfowl food resources limited during spring migration? A bioenergetic assessment of playas in Nebraska's rainwater basin

Accurate bioenergetic carrying capacity estimates of wetlands on public and private lands, as well as those managed for crop production are important for managing waterfowl populations and habitats. Given the importance of wetlands in the Rainwater Basin region of Nebraska for spring migrating waterfowl, we quantified and compared seed and aquatic invertebrate biomass and true metabolizable energy (TME) at three wetland types; public wetlands, wetlands enrolled in the Wetlands Reserve Program (WRP), and cropped wetlands. Median seed biomass estimates at public, WRP, and cropped wetlands were 593kg/ha, 561kg/ha, and 419kg/ha respectively. Cumulative TME varied among wetland type, with greater TME at cropped wetlands (2,431kcal/kg) than public (1,740kcal/kg) and WRP wetlands (1,781kcal/kg). Seed biomass estimates from this study were statistically greater than those currently used for management planning in the RWB, however, TME estimates were statistically lower than estimates currently assumed for WRP and public wetlands. Our estimates for aquatic invertebrate biomass were approximately 40-fold less than seed biomass estimates. Based on spring ponding frequency at wetlands in Nebraska’s Rainwater Basin, and the caloric estimates derived for each wetland type, we concluded that the regions wetlands meet the energetic demand of spring migrating waterfowl during 10% of years.

Nebraska↗

Water-resources data collected in the Devils Hole area, Nye County, Nevada, 1977-1978

This report presents water data collected at Devils Hole and vicinity for July 1977 through June 1978 to observe the effects of irrigation pumping on the pool level in Devils Hole. The pool contains the endangered species of pupfish, Cyprinodon diabolis . The report presents data on fluctuations in water level and springflow in graphic form and tabulates electrical energy consumed by irrigation wells as an index of the pumpage.

Nevada↗

Rejoinder: Challenge and opportunity in the study of ungulate migration amid environmental change

Increasingly, animals that migrate long distances to exploit seasonal habitats must traverse political boundaries capable of altering the very ecological gradients that promote migratory behavior. This transboundary aspect of migration presents many new challenges and opportunities for research and conservation (e.g., Bolger et al. 2008, Taillon et al. 2012). Work to date has often focused on physical barriers to movement (roads, fences,and housing and energy development) that can threaten migratory populations to varying degrees (Holdo et al. 2011, Sawyer et al. 2013). However, even in the absence of conspicuous barriers, political and jurisdictional boundaries can bring dramatic differences in land use and conservation policy. What happens to migratory populations when these boundaries alter the resources and refuges that they seek on their seasonal journeys?

Ecology↗

Chemical analysis of 659 coal samples from the eastern United States

This report includes the analytical data on 659 coal samples from 8 States east of the Mississippi River. The number of samples from each State are: Pennsylvania, 132; Ohio, 101; Maryland, 25; West Virginia, 254; Virginia, 33; Indiana, 49; Massachusetts, 5; and Rhode Island, 10. The analytic data are filed in the National Coal Resource Data System of the U.S. Geological Survey. The U.S. Geological Survey has quantitatively determined 35 major, minor, and trace elements in each sample. Tests were also made for 35 other trace elements using semi-quantitative spectrographic methods. In addition, the Coal Analysis Section of the U.S. Department of Energy has determined proximate and ultimate analyses, 3cu, forms of sulfur, free-swelling indexes, and ash-fusion temperatures on 563 of the 659 samples. Comparison of the geometric means of the 644 bituminous coal samples of this report with an additional 968 bituminous coal samples of the eastern United States reported by Swanson and others in 1976 and by Zubovic and others in 1979 is shown in tables 2a-c. As shown by the means for ultimate and proximate analyses, small differences exist between the two sets of data; however, only the moisture and oxygen contents are significantly different. The forms of sulfur, ash-fusion temperature and heat of combustion are similar. The means for the major and minor oxides in ash are similar for SiO 2 , K 2 O, Fe 2 O 3 , TiO 2 , MgO, and P 2 O 5 are lower, whereas, MnO is higher in the analyses of the samples of this report. Eighteen of the means for the trace elements in the coal are about equal for the two sets of samples. Only As, Co, La, Li, and Pb are significantly higher, whereas, Ag, B, Ba, Be, Cd, Cs, F, Ge, Mo, Nb, and Nd are lower in the in the 64L,bituminous coal samples of this report.

Indiana, Maryland, Massachusetts, Ohio, Pennsylvan↗

Survival and reproduction in Arctic caribou are associated with summer forage and insect harassment

Investigators have speculated that the climate-driven “greening of the Arctic” may benefit barren-ground caribou populations, but paradoxically many populations have declined in recent years. This pattern has raised concerns about the influence of summer habitat conditions on caribou demographic rates, and how populations may be impacted in the future. The short Arctic summer provides caribou with important forage resources but is also the time they are exposed to intense harassment by insects, factors which are both being altered by longer, warmer growing seasons. To better understand the effects of summer forage and insect activity on Arctic caribou demographic rates, we investigated the influence of estimated forage biomass, digestible energy (DE), digestible nitrogen (DN), and mosquito activity on the reproductive success and survival of adult females in the Central Arctic Herd on the North Slope of Alaska. We tested the hypotheses that greater early summer DN would increase subsequent reproduction (parturition and late June calving success) while greater biomass and DE would increase adult survival (September–May), and that elevated mosquito activity would reduce both demographic rates. Because the period when abundant forage DN is limited and overlaps with the period of mosquito harassment, we also expected years with low DN and high harassment to synergistically reduce caribou reproductive success. Examining these relationships at the individual-level, using GPS-collared females, and at the population-level, using long-term monitoring data, we generally found support for our expectations. Greater early summer DN was associated with increased subsequent calving success, while greater summer biomass was associated with increased adult survival. Mosquito activity was associated with reductions in adult female parturition, late June calving success, and survival, and in years with low DN, had compounding effects on subsequent late June calving success. Our findings indicate that summer nutrition and mosquito activity collectively influence the demographic rates of Arctic caribou, and may impact the dynamics of populations in the future under changing environmental conditions.

Alaska↗