USGS Science⌕ Search

SEARCH · USGS Science

Results for “Conservation Ecology”

Search indexed USGS publications on groundwater, aquifers, geologic maps, mineral resources and earthquakes. Explore source records by subject and place.

Quote a phrase for an exact phrase match. Source license links do not imply unrestricted reuse.

At least 1,171 records · Page 65Linked to original sources

The effects of human-caused mortality on mammalian cooperative breeders: A synthesis

Human-caused mortality can be pervasive and even highly selective for individuals in groups of cooperative breeders. Many studies of cooperative breeders, however, do not address human-caused mortality. Similarly, studies focused on the effects of human-caused mortality on wildlife populations often do not consider the ecology of cooperative breeders. We searched the literature and identified 58 studies where human-caused mortality affected a group characteristic, vital rate, or population state of a cooperative breeder. Of studies reporting population growth or decline, 80% reported a link between human-caused mortality and population declines in cooperative breeders. Such studies often did not identify the mechanism behind population declines, but 28% identified concurrent declines in adult survival and another 21% reported concurrent declines in recruitment or reproduction. There was little overlap between the cooperative breeding and human-caused mortality literatures, limiting our ability to accrue knowledge. Future work would be beneficial if it ( i ) identified the vital rate(s) causing population declines, ( ii ) leveraged management actions such as lethal removal to ask questions about the ecology of group-living in cooperative breeders, and ( iii ) used insights from cooperative breeding theory to inform management actions and conservation of group-living species.

Biological Reviews↗

Microsatellite loci for distinguishing spotted owls (Strix occidentalis), barred owls (Strix varia), and their hybrids

We identified four diagnostic microsatellite loci that distinguish spotted owls ( Strix occidentalis ), barred owls ( Strix varia ), F 1 hybrids and backcrosses. Thirty-four out of 52 loci tested (65.4%) successfully amplified, and four of these loci (11.8%) had allele sizes that did not overlap between spotted and barred owls. The probability of correctly identifying a backcross with these four loci is 0.875. Genotyping potential hybrid owls with these markers revealed that field identifications were often wrong. Given the difficulty of identifying hybrids in the field, these markers will be useful for hybrid identification, law enforcement and spotted owl conservation.

Molecular Ecology Resources↗

Multi-scale assessment of roost selection by ‘ōpe‘ape‘a, the Hawaiian hoary bat (Lasiurus semotus)

The Hawaiian hoary bat ( Lasiurus semotus ; Chiroptera: Vespertilionidae), commonly and locally known as ‘ōpe‘ape‘a, is a solitary, insectivorous, and foliage-roosting species distributed across a wide range of habitats in lowland and montane environments. The species, as with many others in the Hawaiian archipelago, are facing a suite of challenges due to habitat loss and degradation, introduced predators and pests, and climate change. An understanding of the roost requirements of foliage-roosting tree bats is critical to their conservation as these habitats provide several important benefits to survival and reproduction. Because little is known about ‘ōpe‘ape‘a roost ecology and considerable effort is needed to capture and track bats to roost locations, we examined resource selection at multiple spatial scales—perch location within a roost tree, roost tree, and forest stand. We used a discrete choice modeling approach to investigate day-roost selection and describe attributes of roost trees including those used as maternity roosts. ‘Ōpe‘ape‘a were found roosting in 19 tree species and in an assortment of landcover types including native and non-native habitats. Our results are largely consistent with findings of other studies of foliage-roosting, insectivorous tree bats where bats selected roost locations that may offer protection and thermoregulatory benefits.

Hawaii↗

Linking demographic effects of habitat fragmentation across landscapes to continental source-sink dynamics

Forest fragmentation may cause increased brood parasitism and nest predation of breeding birds. In North America, nest parasitism and predation are expected to increase closer to forest edges because the brood-parasitic Brown-headed Cowbird (Molothrus ater) and generalist nest predators often enter the forest from adjoining developed (largely agricultural) habitats. Yet the abundance of brood parasites and nest predators at the patch scale may be strongly constrained by the total area of developed habitat at landscape scales. The scale and extent of landscape effects are unclear, however, because past studies were mostly conducted within local landscapes rather than across independent landscapes. We report replicated studies from 30 independent landscapes across 17 states of the United States that show that nest parasitism is strongly affected by fragmentation at a 20 km radius scale, equivalent to the maximum foraging range of cowbirds. Nest predation is influenced by both edge and landscape effects, and increases with fragmentation at a 10 km radius scale. Predation is additive to parasitism mortality, and the two together yield decreased population growth potential with increasing forest fragmentation at a 10 km radius scale for 20 of 22 bird species. Mapping of population growth potential across continental landscapes displays broad impacts of fragmentation on population viability and allows geographic prioritization for conservation.

Ecological Applications↗

Temporal and maternal effects on reproductive ecology of the giant gartersnake ( Thamnophis gigas )

We used mixed-effects models to examine relationships of reproductive characteristics of the giant gartersnake ( Thamnophis gigas ) to improve population modeling and conservation planning for this species. Neonates from larger litters had lower mass, and mass of neonates also was affected by random variation among mothers. Length of mother did not affect relative mass of litters; however, our data suggest that longer mothers expended less reproductive effort per offspring than shorter mothers. We detected random variation in length of neonates among mothers, but these lengths were not related to length of mother or size of litter. Mean size of litter varied among years, but little evidence existed for a relationship between size of litter or mass of litter and length of mother. Sex ratios of neonates did not differ from 1:1.

Southwestern Naturalist↗

Occurrence of a suite of stream-obligate amphibians in timberlands of Mendocino County, California, examined using environmental DNA

Stream-obligate amphibians are important indicators of ecosystem health in the Pacific Northwest, but distributional information to improve forest management is lacking in many regions. We analyzed archived DNA extracted from water samples in 60 pools in streams on private timberlands in Mendocino County, California, for 3 California Species of Special Concern—Coastal Tailed Frogs ( Ascaphus truei ), Foothill Yellow-legged Frogs ( Rana boylii ), and Southern Torrent Salamanders ( Rhyacotriton variegatus )—to better understand their distributions in the region. Detection probabilities for eDNA of Foothill Yellow-legged Frogs and Coastal Tailed Frogs were positively influenced by water temperature. eDNA occurrence for both frogs was affected by whether silt or organic matter was a dominant substrate in the sampled pool, and Foothill Yellow-legged Frog eDNA occurrence was also affected by water temperature. Foothill Yellow-legged Frog eDNA occurrence had a strong, positive association with water temperature, with occurrence unlikely below 14°C and very likely above 16°C, and a positive association with silt or organic substrates in pools, which was likely an indicator of higher-order stream reaches. In contrast, Coastal Tailed Frogs had a negative association with silt or organic substrates. Historical visual detections were generally congruent with findings using eDNA, but differences highlight important areas for further study. We did not detect Southern Torrent Salamanders using eDNA at any sites. Our study reinforces that ecological relationships of these species are varied, and shows the importance of maintaining the integrity of streams with diverse characteristics for conserving stream amphibians.

California↗

Creating a Coastal National Elevation Database (CoNED) for science and conservation applications

The U.S. Geological Survey is creating the Coastal National Elevation Database, an expanding set of topobathymetric elevation models that extend seamlessly across coastal regions of high societal or ecological significance in the United States that are undergoing rapid change or are threatened by inundation hazards. Topobathymetric elevation models are raster datasets useful for inundation prediction and other earth science applications, such as the development of sediment-transport and storm surge models. These topobathymetric elevation models are being constructed by the broad regional assimilation of numerous topographic and bathymetric datasets, and are intended to fulfill the pressing needs of decision makers establishing policies for hazard mitigation and emergency preparedness, coastal managers tasked with coastal planning compatible with predictions of inundation due to sea-level rise, and scientists investigating processes of coastal geomorphic change. A key priority of this coastal elevation mapping effort is to foster collaborative lidar acquisitions that meet the standards of the USGS National Geospatial Program's 3D Elevation Program, a nationwide initiative to systematically collect high-quality elevation data. The focus regions are located in highly dynamic environments, for example in areas subject to shoreline change, rapid wetland loss, hurricane impacts such as overwash and wave scouring, and/or human-induced changes to coastal topography.

Journal of Coastal Research↗

Feeding preferences of West Indian manatees in Florida, Belize, and Puerto Rico as indicated by stable isotope analysis

The endangered West Indian manatee Trichechus manatus has 2 recognized subspecies: the Florida T. m. latirostris and Antillean T. m. manatus manatee, both of which are found in freshwater, estuarine, and marine habitats. A better understanding of manatee feeding preferences and habitat use is essential to establish criteria on which conservation plans can be based. Skin from manatees in Florida, Belize, and Puerto Rico, as well as aquatic vegetation from their presumed diet, were analyzed for stable carbon and nitrogen isotope ratios. This is the first application of stable isotope analysis to Antillean manatees. Stable isotope ratios for aquatic vegetation differed by plant type (freshwater, estuarine, and marine), collection location, and in one instance, season. Carbon and nitrogen isotope ratios for manatee skin differed between collection location and in one instance, season, but did not differ between sex or age class. Signatures in the skin of manatees sampled in Belize and Puerto Rico indicated a diet composed primarily of seagrasses, whereas those of Florida manatees exhibited greater regional variation. Mixing model results indicated that manatees sampled from Crystal River and Homosassa Springs (Florida, USA) ate primarily freshwater vegetation, whereas manatees sampled from Big Bend Power Plant, Ten Thousand Islands, and Warm Mineral Springs (Florida) fed primarily on seagrasses. Possible diet–tissue discrimination values for 15 N were estimated to range from 1.0 to 1.5‰. Stable isotope analysis can be used to interpret manatee feeding behavior over a long period of time, specifically the use of freshwater vegetation versus seagrasses, and can aid in identifying critical habitats and improving conservation efforts.

Florida↗

Manatee grazing impacts on a mixed species seagrass bed

The endangered manatee Trichechus manatus is one of few large grazers in seagrass systems. To assess the long-term impacts of repeated grazing on seagrasses, we selected a study site within Kennedy Space Center in the northern Banana River, Brevard County, Florida, that was typically grazed by large numbers of manatees in spring. Two 13x13 m manatee exclosures and 2 paired open plots of equal size were established at the study site in October 1990. Shoot counts, biomass, and species composition of the co-dominant seagrass species, Syringodium filiforme and Halodule wrightii , were sampled 3 times per year in all 4 plots between October 1990 and October 1994. We used a Bayesian modelling approach, accounting for the influence of depth, to detect treatment (exclosed vs. open) effects. S. filiforme shoot counts, total biomass, and frequency of occurrence significantly increased in the exclosures. By July 1993, mean biomass values in the exclosures (167 g dry wt m -2 ) greatly exceeded those in the open plots (28 g dry wt m -2 ). H. wrightii decreased in the exclosures by 1994. Initially, both S. filiforme and H. wrightii responded positively to release from manatee grazing pressure. As S. filiforme continued to become denser in the exclosures, it gradually replaced H. wrightii . Our findings may be helpful to biologists and managers interested in predicting seagrass recovery and manatee carrying capacity of repeatedly grazed seagrass beds in areas of special significance to manatees and seagrass conservation.

Marine Ecology Progress Series↗

Shared habitat use by juveniles of three sea turtle species

The first step in understanding how sympatric species share habitat is defining spatial boundaries. While home range data for juvenile sea turtles exists, few studies have examined spatial overlap of multiple species in foraging habitat. Using satellite tracking technology, we define home ranges for juveniles of 3 sea turtle species (loggerhead, Kemp’s ridley, and green; n = 21) captured at 2 adjacent foraging sites in the northern Gulf of Mexico. In these areas, green turtles are known to be primarily herbivorous, whereas Kemp’s ridley turtles forage predominately on crabs, and loggerhead turtles on various hard-shelled benthic invertebrates. No differences in home range size or characteristics, such as water depth and distance to shore, were observed among species, although fine-scale foraging patches were not examined in this study. A high degree of overlap in habitat-use among all 3 species was documented in summer at both sites. Seasonal movements, triggered by colder winter temperatures, were documented and appeared to differ among species, with Kemp’s ridley and loggerhead turtles leaving bays, and green turtles overwintering inside bays. By identifying shared habitat-use by juvenile sea turtles, we have created a foundation for further fine-scale studies on resource partitioning that will aid in habitat management and conservation of these threatened and endangered species.

Marine Ecology Progress Series↗

Synthesizing professional opinion of Lake Whitefish and Cisco recruitment drivers across the Great Lakes

Disentangling the suite of ecological drivers that explain recruitment variability for Lake Whitefish Coregonus clupeaformis and Cisco C. artedi is of critical importance for their conservation, management, and stewardship in the Laurentian Great Lakes. However, recruitment is inherently variable and can be regulated by many interacting processes, the relative importance of which can vary spatially, temporally, and ontogenetically. Given this complexity, comparisons across lakes and species that identify overarching hypotheses could efficiently guide future research. Using facilitated deliberations among fishery professionals (n = 57) with expertise in Great Lakes Coregonus spp., we synthesized current knowledge regarding (1) which biophysical processes are most important for driving contemporary recruitment between species, among lakes, and across life stages and (2) mechanisms by which those drivers regulate recruitment at key life stages. Participants affirmed the hypothesis that many drivers interact in complex ways to regulate Lake Whitefish and Cisco recruitment. Large-scale climatic processes affecting early life-stage growth and survival were consistently considered important. Other drivers were only deemed influential in certain lakes, highlighting perceived context-dependent recruitment dynamics. Notably, recruitment in Lakes Superior, Michigan, and Huron was considered limited during larval and early juvenile life stages by low productivity, whereas spawning-habitat degradation and reduced metapopulation diversity were hypothesized to limit recruitment during embryonic and larval stages in Lakes Erie and Ontario. Several drivers were hypothesized to similarly impact Lake Whitefish and Cisco during early life stages, while drivers acting on post-larval life stages were typically distinct between species. The hypotheses synthesized herein can guide future research on Lake Whitefish and Cisco recruitment dynamics in the Great Lakes.

Great Lakes↗

Case Study 4: NABat acoustic monitoring allows inferences about bat populations at multiple scales

North American bats face unprecedented risks from continuing and emerging threats including white-nose syndrome, wind energy development, and habitat loss. Many species of bats are thought to be recently experiencing unparalleled population declines unlike any previously observed (O’Shea et al. 2016). The North American Bat Monitoring Program (NABat) was conceived to better understand the true ecological consequences of these large-scale population reductions (Loeb et al. 2015). NABat aims is to improve the state of conservation science for the 47 species of bats shared by Canada, United States, and Mexico. To meet this objective, NABat offers standardize protocols and a unifying sample design facilitating a multi-agency, multinational, collaborative monitoring effort. A key element of NABat is cross-boundary partner coordination and sharing of limited resources for the collection of bat echolocation data. Here we provide three compelling examples of how NABat provides a convenient framework for using acoustic data to assess the potential impacts of current and future threats to North American bats across multiple spatial scales.

Oregon↗

Klamath-Siskiyou herpetofauna: Biogeographic patterns and conservation strategies

The Klamath-Siskiyou region of southwest Oregon and northwest California (USA) has some of the most complex landscape mosaics and plant communities in western North America, reflecting its marked diversity of precipitation and topography. With 38 native species of amphibians and reptiles, the Klamath-Siskiyou region has the most species-rich herpetofauna of any similarly sized mountain range in the Pacific Northwest. Although it is a biodiversity 'hot spot,' there are only two endemic species, both salamanders, in the Klamath-Siskiyou region. High diversity is due to the overlap of two major biogeographic groups: the Arcto- (= northern) and Madro- (= southern) Tertiary herpetofaunas. Many of the amphibians in the Klamath-Siskiyou region are restricted to specialized habitats. Much of our knowledge about the biology of the regional fauna is based on studies elsewhere. Distributional surveys and ecological research are needed to address how the herpetofauna responds to timber harvest and other human activities that may reduce populations and increase fragmentation of suitable habitats. Conservation of the region's diverse herpetofauna should emphasize strategies directed at habitat specialists and species at the latitudinal limits of their ranges.

California, Oregon↗

Improving conservation outcomes with a new paradigm for understanding species’ fundamental and realized adaptive capacity

Worldwide, many species are responding to ongoing climate change with shifts in distribution, abundance, phenology, or behavior. Consequently, natural-resource managers face increasingly urgent conservation questions related to biodiversity loss, expansion of invasive species, and deteriorating ecosystem services. We argue that our ability to address these questions is hampered by the lack of explicit consideration of species’ adaptive capacity (AC). AC is the ability of a species or population to cope with climatic changes and is characterized by three fundamental components: phenotypic plasticity, dispersal ability, and genetic diversity. However, few studies simultaneously address all elements; often, AC is confused with sensitivity or omitted altogether from climate-change vulnerability assessments. Improved understanding, consistent definition, and comprehensive evaluations of AC are needed. Using classic ecological-niche theory as an analogy, we propose a new paradigm that considers fundamental and realized AC: the former reflects aspects inherent to species, whereas the latter denotes how extrinsic factors constrain AC to what is actually expressed or observed. Through this conceptualization, we identify ecological attributes contributing to AC, outline areas of research necessary to advance understanding of AC, and provide examples demonstrating how the inclusion of AC can better inform conservation and natural-resource management.

Conservation Letters↗

Southeast Regional Assessment Project for the National Climate Change and Wildlife Science Center, U.S. Geological Survey

The Southeastern United States spans a broad range of physiographic settings and maintains exceptionally high levels of faunal diversity. Unfortunately, many of these ecosystems are increasingly under threat due to rapid human development, and management agencies are increasingly aware of the potential effects that climate change will have on these ecosystems. Natural resource managers and conservation planners can be effective at preserving ecosystems in the face of these stressors only if they can adapt current conservation efforts to increase the overall resilience of the system. Climate change, in particular, challenges many of the basic assumptions used by conservation planners and managers. Previous conservation planning efforts identified and prioritized areas for conservation based on the current environmental conditions, such as habitat quality, and assumed that conditions in conservation lands would be largely controlled by management actions (including no action). Climate change, however, will likely alter important system drivers (temperature, precipitation, and sea-level rise) and make it difficult, if not impossible, to maintain recent historic conditions in conservation lands into the future. Climate change will also influence the future conservation potential of non-conservation lands, further complicating conservation planning. Therefore, there is a need to develop and adapt effective conservation strategies to cope with the effects of climate and landscape change on future environmental conditions. Congress recognized this important issue and authorized the U.S. Geological Survey (USGS) National Climate Change and Wildlife Science Center (NCCWSC; http://nccw.usgs.gov/) in the Fiscal Year 2008. The NCCWSC will produce science that will help resource management agencies anticipate and adapt to climate change impacts to fish, wildlife, and their habitats. With the release of Secretarial Order 3289 on September 14, 2009, the mandate of the NCCWSC was expanded to address climate change-related impacts on all Department of the Interior (DOI) resources. The NCCWSC will establish a network of eight DOI Regional Climate Science Centers (RCSCs) that will work with a variety of partners to provide natural resource managers with tools and information that will help them anticipate and adapt conservation planning and design for projected climate change. The forecasting products produced by the RCSCs will aid fish, wildlife, and land managers in designing suitable adaptive management approaches for their programs. The DOI also is developing Landscape Conservation Cooperatives (LCCs) as science and conservation action partnerships at subregional scales. The USGS is working with the Southeast Region of the U.S. Fish and Wildlife Service (FWS) to develop science collaboration between the future Southeast RCSC and future LCCs. The NCCWSC Southeast Regional Assessment Project (SERAP) will begin to develop regional downscaled climate models, land cover change models, regional ecological models, regional watershed models, and other science tools. Models and data produced by SERAP will be used in a collaborative process between the USGS, the FWS (LCCs), State and federal partners, nongovernmental organizations, and academia to produce science at appropriate scales to answer resource management questions. The SERAP will produce an assessment of climate change, and impacts on land cover, ecosystems, and priority species in the region. The predictive tools developed by the SERAP project team will allow end users to better understand potential impacts of climate change and sea level rise on terrestrial and aquatic populations in the Southeastern United States. The SERAP capitalizes on the integration of five existing projects: (1) the Multi-State Conservation Grants Program project "Designing Sustainable Landscapes," (2) the USGS multidisciplinary Science Thrust project "Water Availability for Ecological Needs," (3) the USGS Southeast Pilot Project "Climate Change in the Southeastern U.S. and its Impacts on Bird Distributions and Habitats," (4) a sea-level rise impacts study envisioned jointly with the National Oceanic and Atmospheric Administration (NOAA), and (5) two USGS sea-level rise impact assessment projects that address inundation hazards and provide probabilistic forecasts of coastal geomorphic change. The SERAP will expand on these existing projects and include the following tasks, which were initiated in summer 2009: * Regionally downscaled probabilistic climate-change projections * Integrated coastal assessment * Integrated terrestrial assessment * Multi-resolution assessment of potential climate change effects on biological resources: aquatic and hydrologic dynamics * Optimal conservation strategies to cope with climate change The SERAP seeks to formally integrate these tasks to aid conservation planning and design so that ecosystem management decisions can be optimized for providing desirable outcomes across a range of species and environments. The following chapters detail SERAP's efforts in providing a suite of regional climate, watershed, and landscape-change analyses and develop the interdisciplinary framework required for the biological planning phases of adaptive management and strategic conservation. The planning phase will include the identification of conservation alternatives, development of predictive models and decision support tools, and development of a template to address similar challenges and goals in other regions. The project teams will explore and develop ways to link the various ecological models arising from each component. The SERAP project team also will work closely with members of the LCCs and other partnerships throughout the life of the project to ensure that the objectives of the project meet resources mangers needs in the Southeast.

Open-File Report↗

Length in assessing status of freshwater fish populations: A review

Objective Effective policy formulation regarding the conservation of freshwater fish necessitates an understanding of water‐specific prevailing conditions and trends. Assessing fish populations in inland waters is difficult and expensive because there are many independent systems that need to be evaluated. Therefore, numerous freshwater systems are beset by insufficient data and the lack of systematic assessments of their status. To alleviate this deficiency, the objective of this study was to review length‐based metrics that may have utility in evaluating the well‐being of freshwater fish populations. Methods Length measurements can serve as proxies for a range of ecological and population dynamics attributes that are essential for the effective management of fish and associated fisheries. A review of the historical development of length measurements in fish conservation is provided, along with an examination of the potential biases that may arise from the use of lengths in practical contexts. In addition, we examine techniques that enable the spatial and temporal visualization of length data sets, as well as a range of indices and metrics that can be computed using length measurements. Result Building populations assessments around length may be a cost‐effective strategy that allows a first cut at managing a large number of waters. Length‐based assessments can signal if management intervention is necessary, if management policies are yielding the intended outcome, or if surveys beyond mere length are necessary. Conclusion Our review indicates that length offers a straightforward and efficient approach to evaluate the status of fish populations in inland systems. We encourage pursuing additional study and to this end propose specific areas for investigation.

North American Journal of Fisheries Management↗

Top 40 priorities for science to inform conservation and management policy in the United States

To maximize the utility of research to decisionmaking, especially given limited financial resources, scientists must set priorities for their efforts. We present a list of the top 40 high-priority, multidisciplinary research questions directed toward informing some of the most important current and future decisions about management of species, communities, and ecological processes in the United States. The questions were generated by an open, inclusive process that included personal interviews with decisionmakers, broad solicitation of research needs from scientists and policymakers, and an intensive workshop that included scientifically oriented individuals responsible for managing and developing policy related to natural resources. The process differed from previous efforts to set priorities for conservation research in its focus on the engagement of decisionmakers in addition to researchers. The research priorities emphasized the importance of addressing societal context and exploration of trade-offs among alternative policies and actions, as well as more traditional questions related to ecological processes and functions.

BioScience↗

Projected climate-induced faunal change in the Western Hemisphere

Climate change is predicted to be one of the greatest drivers of ecological change in the coming century. Increases in temperature over the last century have clearly been linked to shifts in species distributions. Given the magnitude of projected future climatic changes, we can expect even larger range shifts in the coming century. These changes will, in turn, alter ecological communities and the functioning of ecosystems. Despite the seriousness of predicted climate change, the uncertainty in climate-change projections makes it difficult for conservation managers and planners to proactively respond to climate stresses. To address one aspect of this uncertainty, we identified predictions of faunal change for which a high level of consensus was exhibited by different climate models. Specifically, we assessed the potential effects of 30 coupled atmosphere-ocean general circulation model (AOGCM) future-climate simulations on the geographic ranges of 2954 species of birds, mammals, and amphibians in the Western Hemisphere. Eighty percent of the climate projections based on a relatively low greenhouse-gas emissions scenario result in the local loss of at least 10% of the vertebrate fauna over much of North and South America. The largest changes in fauna are predicted for the tundra, Central America, and the Andes Mountains where, assuming no dispersal constraints, specific areas are likely to experience over 90% turnover, so that faunal distributions in the future will bear little resemblance to those of today. ?? 2009 by the Ecological Society of America.

Ecology↗