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Semi-arid vegetation response to antecedent climate and water balance windows

Questions Can we improve understanding of vegetation response to water availability on monthly time scales in semi-arid environments using remote sensing methods? What climatic or water balance variables and antecedent windows of time associated with these variables best relate to the condition of vegetation? Can we develop credible near-term forecasts from climate data that can be used to prepare for future climate change effects on vegetation? Location Semi-arid grasslands in Capitol Reef National Park, Utah, USA. Methods We built vegetation response models by relating the normalized difference vegetation index (NDVI) from MODIS imagery in Mar–Nov 2000–2013 to antecedent climate and water balance variables preceding the monthly NDVI observations. We compared how climate and water balance variables explained vegetation greenness and then used a multi-model ensemble of climate and water balance models to forecast monthly NDVI for three holdout years. Results Water balance variables explained vegetation greenness to a greater degree than climate variables for most growing season months. Seasonally important variables included measures of antecedent water input and storage in spring, switching to indicators of drought, input or use in summer, followed by antecedent moisture availability in autumn. In spite of similar climates, there was evidence the grazed grassland showed a response to drying conditions 1 mo sooner than the ungrazed grassland. Lead times were generally short early in the growing season and antecedent window durations increased from 3 mo early in the growing season to 1 yr or more as the growing season progressed. Forecast accuracy for three holdout years using a multi-model ensemble of climate and water balance variables outperformed forecasts made with a naïve NDVI climatology. Conclusions We determined the influence of climate and water balance on vegetation at a fine temporal scale, which presents an opportunity to forecast vegetation response with short lead times. This understanding was obtained through high-frequency vegetation monitoring using remote sensing, which reduces the costs and time necessary for field measurements and can lead to more rapid detection of vegetation changes that could help managers take appropriate actions.

Applied Vegetation Science↗

From salmon to shad: Shifting sources of marine-derived nutrients in the Columbia River Basin

Like Pacific salmon ( Oncorhynchus spp.), nonnative American shad ( Alosa sapidissima ) have the potential to convey large quantities of nutrients between the Pacific Ocean and freshwater spawning areas in the Columbia River Basin (CRB). American shad are now the most numerous anadromous fish in the CRB, yet the magnitude of the resulting nutrient flux owing to the shift from salmon to shad is unknown. Nutrient flux models revealed that American shad conveyed over 15,000 kg of nitrogen (N) and 3,000 kg of phosphorus (P) annually to John Day Reservoir, the largest mainstem reservoir in the lower Columbia River. Shad were net importers of N, with juveniles and postspawners exporting just 31% of the N imported by adults. Shad were usually net importers of P, with juveniles and postspawners exporting 46% of the P imported by adults on average. American shad contributed <0.2% of the total annual P load into John Day Reservoir, but during June when most adult shad are migrating into John Day Reservoir, they contributed as much as 2.0% of the P load. Nutrient inputs by American shad were similar to current but far less than historical inputs of Pacific salmon owing to their smaller size. Given the relatively high background P levels and low retention times in lower Columbia River reservoirs, it is unlikely that shad marine-derived nutrients affect nutrient balances or food web productivity through autotrophic pathways. However, a better understanding of shad spawning aggregations in the CRB is needed.

Columbia River, John Day Reservoir↗

Growth of a threatened desert fish becomes resource dependent when released from temperature limitation in a regulated river

1. Growth of ectotherms typically positively correlates with increasing temperature towards an optimal temperature range, followed by a decline in growth when temperatures exceed the optimal temperature range for that organism. When temperature is within or close to the optimal range, food availability and other environmental factors can play important roles in modifying growth rates. River regulation often alters both environmental conditions (e.g., flow, turbidity and temperature) and aquatic food webs, modifying the quality and quantity of resources available to consumers. Here we aim to understand how food availability, water temperature and other environmental factors are associated with fish growth (defined here as a change in length) in a large, regulated desert river. 2. We measured growth of humpback chub (Gila cypha) at two river reaches located 240 km apart in the Colorado River within the Grand Canyon, Arizona, USA. We then fit Bayesian state-space models of growth that account for variable time at large to test environmental predictors such as water temperature, flow, turbidity (which increases during tributary flooding and can be a proxy for allochthonous inputs) and gross primary productivity (a proxy for autochthonous food) using an inclusion parameter approach. Both river reaches are located below Glen Canyon Dam, which regulates flow, alters seasonal water temperatures and traps sediment, nutrients, and organic carbon; however, merging tributaries and a greater distance from the dam lead to differences in environmental conditions. 3. In the upriver reach, where warm season water temperatures are below the optimal range for humpback chub most of the time, temperature was the primary driver of variation in growth according to inclusion parameter values and the strength of the standardised effect size. Turbidity was also included in the final model for the upper reach, but the inclusion parameter values and standardised effect size were smaller than for temperature. In the downriver, warmer reach, temperature still had the highest inclusion parameter value, but GPP had the highest standardised effect and nearly as high an inclusion parameter value. 4. We conclude that temperature is the primary limiting factor on growth of humpback chub in the Grand Canyon; however, as water temperatures increase, other factors, especially GPP, are associated with temporal variation in growth rates. Over the past two decades, declines in reservoir elevations and the volume of summer releases have increased downstream water temperatures. If warm temperatures persist into the future, other factors may have increasing roles in regulating interannual variability in chub growth. 5. Field studies of ectotherm growth often emphasise the role of water temperature; however, the role of resource availability is also increasingly recognised, especially in bioenergetics studies. Quantifying resource availability directly can be difficult, especially in large, remote rivers. Here, we show here that using proxies for resource availability (i.e., GPP as a proxy for autochthonous inputs) can yield useful insights regarding resource limitations with implications for management of federally listed fish species in a larger regulated river system.

Arizona↗

The response of soil organic carbon of a rich fen peatland in interior Alaska to projected climate change

It is important to understand the fate of carbon in boreal peatland soils in response to climate change because a substantial change in release of this carbon as CO 2 and CH 4 could influence the climate system. The goal of this research was to synthesize the results of a field water table manipulation experiment conducted in a boreal rich fen into a process-based model to understand how soil organic carbon (SOC) of the rich fen might respond to projected climate change. This model, the peatland version of the dynamic organic soil Terrestrial Ecosystem Model (peatland DOS-TEM), was calibrated with data collected during 2005–2011 from the control treatment of a boreal rich fen in the Alaska Peatland Experiment (APEX). The performance of the model was validated with the experimental data measured from the raised and lowered water-table treatments of APEX during the same period. The model was then applied to simulate future SOC dynamics of the rich fen control site under various CO 2 emission scenarios. The results across these emissions scenarios suggest that the rate of SOC sequestration in the rich fen will increase between year 2012 and 2061 because the effects of warming increase heterotrophic respiration less than they increase carbon inputs via production. However, after 2061, the rate of SOC sequestration will be weakened and, as a result, the rich fen will likely become a carbon source to the atmosphere between 2062 and 2099. During this period, the effects of projected warming increase respiration so that it is greater than carbon inputs via production. Although changes in precipitation alone had relatively little effect on the dynamics of SOC, changes in precipitation did interact with warming to influence SOC dynamics for some climate scenarios.

Alaska↗

Nitrogen deposition weakens soil carbon control of nitrogen dynamics across the contiguous United States

Anthropogenic nitrogen (N) deposition is unequally distributed across space and time, with inputs to terrestrial ecosystems impacted by industry regulations and variations in human activity. Soil carbon (C) content normally controls the fraction of mineralized N that is nitrified ( ƒ nitrified ), affecting N bioavailability for plants and microbes. However, it is unknown whether N deposition has modified the relationships among soil C, net N mineralization, and net nitrification. To test whether N deposition alters the relationship between soil C and net N transformations, we collected soils from coniferous and deciduous forests, grasslands, and residential yards in 14 regions across the contiguous United States that vary in N deposition rates. We quantified rates of net nitrification and N mineralization, soil chemistry (soil C, N, and pH), and microbial biomass and function (as beta-glucosidase (BG) and N -acetylglucosaminidase (NAG) activity) across these regions. Following expectations, soil C was a driver of ƒ nitrified across regions, whereby increasing soil C resulted in a decline in net nitrification and ƒ nitrified . The ƒ nitrified value increased with lower microbial enzymatic investment in N acquisition (increasing BG:NAG ratio) and lower active microbial biomass, providing some evidence that heterotrophic microbial N demand controls the ammonium pool for nitrifiers. However, higher total N deposition increased ƒ nitrified , including for high soil C sites predicted to have low ƒ nitrified , which decreased the role of soil C as a predictor of ƒ nitrified . Notably, the drop in contemporary atmospheric N deposition rates during the 2020 COVID-19 pandemic did not weaken the effect of N deposition on relationships between soil C and ƒ nitrified . Our results suggest that N deposition can disrupt the relationship between soil C and net N transformations, with this change potentially explained by weaker microbial competition for N. Therefore, past N inputs and soil C should be used together to predict N dynamics across terrestrial ecosystems.

contiguous United States↗

Simulating water-quality trends in public-supply wells in transient flow systems

Models need not be complex to be useful. An existing groundwater-flow model of Salt Lake Valley, Utah, was adapted for use with convolution-based advective particle tracking to explain broad spatial trends in dissolved solids. This model supports the hypothesis that water produced from wells is increasingly younger with higher proportions of surface sources as pumping changes in the basin over time. At individual wells, however, predicting specific water-quality changes remains challenging. The influence of pumping-induced transient groundwater flow on changes in mean age and source areas is significant. Mean age and source areas were mapped across the model domain to extend the results from observation wells to the entire aquifer to see where changes in concentrations of dissolved solids are expected to occur. The timing of these changes depends on accurate estimates of groundwater velocity. Calibration to tritium concentrations was used to estimate effective porosity and improve correlation between source area changes, age changes, and measured dissolved solids trends. Uncertainty in the model is due in part to spatial and temporal variations in tracer inputs, estimated tracer transport parameters, and in pumping stresses at sampling points. For tracers such as tritium, the presence of two-limbed input curves can be problematic because a single concentration can be associated with multiple disparate travel times. These shortcomings can be ameliorated by adding hydrologic and geologic detail to the model and by adding additional calibration data. However, the Salt Lake Valley model is useful even without such small-scale detail.

Utah↗

A new groundwater energy transport model for the MODFLOW hydrologic simulator

Heat transport in the subsurface is an important aspect of research related to the effects of a warming climate on ecological services (i.e., cold-water refugia); the development of geothermal resources for energy banking schemes (i.e., aquifer thermal energy storage [ATES]); and the effects of temperature on other aspects of groundwater quality, such as nutrient cycling. Historically, simulation of heat transport using the MODFLOW groundwater simulator and related codes was performed by scaling the input parameters of a solute-transport model to emulate heat transport. However, that approach required additional pre- and post-processing of input and output and could not account for the variation in effective thermal storage and transport properties during transient, unsaturated flow, for example. True heat-transport capabilities in the context of MODFLOW were first introduced in a variant called USG-Transport. More recently, a new groundwater energy-transport (GWE) model type has been added to MODFLOW 6, the core version of the MODFLOW hydrologic simulator. GWE supports the simulation of heat transport on structured or unstructured grids as well as within and between features of advanced packages that represent streams, lakes, multi-aquifer wells, and the unsaturated zone. GWE is integrated within MODFLOW 6 and is accessible through the FloPy Python package and the MODFLOW 6 application programming interface (API). An example simulation demonstrates conduction between grid cells through both the water and the solid aquifer material, including thermal bleeding from saturated overburden cells into a groundwater flow field.

Groundwater↗

Long-term increases in young-of-the-year growth of Arctic cisco Coregonus autumnalis and environmental influences

Arctic cisco Coregonus autumnalis young-of-year (YOY) growth was used as a proxy to examine the long-term response of a high-latitude fish population to changing climate from 1978 to 2004. YOY growth increased over time (r2 = 0&middot;29) and was correlated with monthly averages of the Arctic oscillation index, air temperature, east wind speed, sea-ice concentration and river discharge with and without time lags. Overall, the most prevalent correlates to YOY growth were sea-ice concentration lagged 1 year (significant correlations in 7 months; r 2 = 0&middot;14-0&middot;31) and Mackenzie River discharge lagged 2 years (significant correlations in 8 months; r 2 = 0&middot;13-0&middot;50). The results suggest that decreased sea-ice concentrations and increased river discharge fuel primary production and that life cycles of prey species linking increased primary production to fish growth are responsible for the time lag. Oceanographic studies also suggest that sea ice concentration and fluvial inputs from the Mackenzie River are key factors influencing productivity in the Beaufort Sea. Future research should assess the possible mechanism relating sea ice concentration and river discharge to productivity at upper trophic levels.

Journal of Fish Biology↗

Population structure, persistence, and seasonality of autochthonous Escherichia coli in temperate, coastal forest soil from a Great Lakes watershed

The common occurrence of Escherichia coli in temperate soils has previously been reported, however, there are few studies to date to characterize its source, distribution, persistent capability and genetic diversity. In this study, undisturbed, forest soils within six randomly selected 0.5 m2 exclosure plots (covered by netting of 2.3 mm2 mesh size) were monitored from March to October 2003 for E. coli in order to describe its numerical and population characteristics. Culturable E. coli occurred in 88% of the samples collected, with overall mean counts of 16 MPN g-1, ranging from <1 to 1657 (n = 66). Escherichia coli counts did not correlate with substrate moisture content, air, or soil temperatures, suggesting that seasonality were not a strong factor in population density control. Mean E. coli counts in soil samples (n = 60) were significantly higher inside than immediately outside the exclosures; E. coli distribution within the exclosures was patchy. Repetitive extragenic palindromic polymerase chain reaction (Rep-PCR) demonstrated genetic heterogeneity of E. coli within and among exclosure sites, and the soil strains were genetically distinct from animal (E. coli) strains tested (i.e. gulls, terns, deer and most geese). These results suggest that E. coli can occur and persist for extended periods in undisturbed temperate forest soils independent of recent allochthonous input and season, and that the soil E. coli populations formed a cohesive phylogenetic group in comparison to the set of fecal strains with which they were compared. Thus, in assessing E. coli sources within a stream, it is important to differentiate background soil loadings from inputs derived from animal and human fecal contamination. ?? 2005 Society for Applied Microbiology and Blackwell Publishing Ltd.

Environmental Microbiology↗

Caribbean mangroves adjust to rising sea level through biotic controls on change in soil elevation

Aim The long-term stability of coastal ecosystems such as mangroves and salt marshes depends upon the maintenance of soil elevations within the intertidal habitat as sea level changes. We examined the rates and processes of peat formation by mangroves of the Caribbean Region to better understand biological controls on habitat stability. Location Mangrove-dominated islands on the Caribbean coasts of Belize, Honduras and Panama were selected as study sites. Methods Biological processes controlling mangrove peat formation were manipulated (in Belize) by the addition of nutrients (nitrogen or phosphorus) to Rhizophora mangle (red mangrove), and the effects on the dynamics of soil elevation were determined over a 3-year period using rod surface elevation tables (RSET) and marker horizons. Peat composition and geological accretion rates were determined at all sites using radiocarbon-dated cores. Results The addition of nutrients to mangroves caused significant changes in rates of mangrove root accumulation, which influenced both the rate and direction of change in elevation. Areas with low root input lost elevation and those with high rates gained elevation. These findings were consistent with peat analyses at multiple Caribbean sites showing that deposits (up to 10 m in depth) were composed primarily of mangrove root matter. Comparison of radiocarbon-dated cores at the study sites with a sea-level curve for the western Atlantic indicated a tight coupling between peat building in Caribbean mangroves and sea-level rise over the Holocene. Main conclusions Mangroves common to the Caribbean region have adjusted to changing sea level mainly through subsurface accumulation of refractory mangrove roots. Without root and other organic inputs, submergence of these tidal forests is inevitable due to peat decomposition, physical compaction and eustatic sea-level rise. These findings have relevance for predicting the effects of sea-level rise and biophysical processes on tropical mangrove ecosystems.

Caribbean Sea↗

Relations between fish abundances, summer temperatures, and forest harvest in a northern Minnesota stream system from 1997 to 2007

Short‐term effects of forest harvest on fish habitat have been well documented, including sediment inputs, leaf litter reductions, and stream warming. However, few studies have considered changes in local climate when examining postlogging changes in fish communities. To address this need, we examined fish abundances between 1997 and 2007 in a basin in a northern hardwood forest. Streams in the basin were subjected to experimental riparian forest harvest in fall 1997. We noted a significant decrease for fish index of biotic integrity and abundance of Salvelinus fontinalis and Phoxinus eos over the study period. However, for P. eos and Culaea inconstans , the temporal patterns in abundances were related more to summer air temperatures than to fine sediment or spring precipitation when examined using multiple regressions. Univariate regressions suggested that summer air temperatures influenced temporal patterns in fish communities more than fine sediment or spring precipitation.

Minnesota↗

Simulation of cylindrical flow to a well using the U.S. Geological Survey Modular Finite-Difference Ground-Water Flow Model

Cylindrical (axasymmetric) flow to a well is an important specialized topic of ground-water hydraulics and has been applied by many investigators to determine aquifer properties and determine heads and flows in the vicinity of the well. A recent modification to the U.S. Geological Survey Modular Three-Dimensional Finite-Difference Ground-Water Flow Model provides the opportunity to simulate axisymmetric flow to a well. The theory involves the conceptualization of a system of concentric shells that are capable of reproducing the large variations in gradient in the vicinity of the well by decreasing their area in the direction of the well. The computer program presented serves as a preprocessor to the U.S. Geological Survey model by creating the input data file needed to implement the axisymmetric conceptualization. Data input requirements to this preprocessor are described, and a comparison with a known analytical solution indicates that the model functions appropriately.

Groundwater↗

The ratio method of estimating water resistivity and TDS from resistivity logs

It is a general belief that useful estimates of total dissolved solids concentrations of ground water cannot be made from borehole geophysical logs. A case study of estimating total dissolved solids concentration of ground water in the local area using the ratio method yielded estimates with an average error of less than 25 percent. The results do not support the hypothesis that useful estimates of total dissolved solids concentration cannot be made from borehole geophysical logs. The case study included a comparison of estimates of total dissolved solids concentration utilizing a resistivity of the mud input versus using resistivity of the mud filtrate input. Estimates made using resistivity of mud had a correlation coefficient of 0.97 whereas estimates using resistivity of mud filtrate had a correlation coefficient of only 0.27. The results from the case study suggest that at least in some cases the resistivity of the mud (Rm) produce a better estimate of the resistivity of water (Rw) in the fully flushed zone than an estimate using the resistivity of the mud filtrate Rmf. The ratio method can be easily used to estimate ground-water resistivity and total dissolved solids concentration of the formation water based only on data from resistivity logs. The advantage of the method is that data on porosity, cementation exponent, temperature, and volume of clay are not required. The method, which has been used by the oil industry to crudely estimate water resistivity, is based in part on the ratio of the resistivity of a fully water-saturated formation to the resistivity of the fully flushed zone adjacent to the annulus in a mud-filled borehole. The method, which is very robust, requires only an estimate of the resistivity of a fully water-saturated formation from a deep looking induction or resistivity log, an estimate of the resistivity of the fully flushed zone from a microresistivity or short normal log, and a measurement of resistivity of the mud or mud filtrate and its temperature.

Ground Water↗

Estimating background and threshold nitrate concentrations using probability graphs

Because of the ubiquitous nature of anthropogenic nitrate (NO 3-) in many parts of the world, determining background concentrations of NO3- in shallow ground water from natural sources is probably impossible in most environments. Present-day background must now include diffuse sources of NO3- such as disruption of soils and oxidation of organic matter, and atmospheric inputs from products of combustion and evaporation of ammonia from fertilizer and livestock waste. Anomalies can be defined as NO3- derived from nitrogen (N) inputs to the environment from anthropogenic activities, including synthetic fertilizers, livestock waste, and septic effluent. Cumulative probability graphs were used to identify threshold concentrations separating background and anomalous NO3-N concentrations and to assist in the determination of sources of N contamination for 232 spring water samples and 200 well water samples from karst aquifers. Thresholds were 0.4, 2.5, and 6.7 mg/L for spring water samples, and 0.1, 2.1, and 17 mg/L for well water samples. The 0.4 and 0.1 mg/L values are assumed to represent thresholds for present-day precipitation. Thresholds at 2.5 and 2.1 mg/L are interpreted to represent present-day background concentrations of NO3-N. The population of spring water samples with concentrations between 2.5 and 6.7 mg/L represents an amalgam of all sources of NO3- in the ground water basins that feed each spring; concentrations >6.7 mg/L were typically samples collected soon after springtime application of synthetic fertilizer. The 17 mg/L threshold (adjusted to 15 mg/L) for well water samples is interpreted as the level above which livestock wastes dominate the N sources. Copyright ?? 2006 The Author(s).

Ground Water↗

MODFLOW-style parameters in underdetermined parameter estimation

In this article, we discuss the use of MODFLOW-Style parameters in the numerical codes MODFLOW_2005 and MODFLOW_2005-Adjoint for the definition of variables in the Layer Property Flow package. Parameters are a useful tool to represent aquifer properties in both codes and are the only option available in the adjoint version. Moreover, for overdetermined parameter estimation problems, the parameter approach for model input can make data input easier. We found that if each estimable parameter is defined by one parameter , the codes require a large computational effort and substantial gains in efficiency are achieved by removing logical comparison of character strings that represent the names and types of the parameters. An alternative formulation already available in the current implementation of the code can also alleviate the efficiency degradation due to character comparisons in the special case of distributed parameters defined through multiplication matrices. The authors also hope that lessons learned in analyzing the performance of the MODFLOW family codes will be enlightening to developers of other Fortran implementations of numerical codes.

Groundwater↗

Oxidized Nitrogen in Precipitation, Throughfall, and Streamfall from a Forested Watershed in Oklahoma

Oxidized nitrogen (nitrite plus nitrate N) concentrations were measured from bulk precipitation, bulk throughfall, and screamflow in a 7. 86 hectare forested watershed in southeastern Oklahoma during the wet season from March through June 1983. Oxidized nitrogen inputs comparable to results of other studies were recorded during the 19 rainstorms sampled. Oxidized nitrogen concentrations appeared to increase after rainfall interacted with the pine and hardwood canopies and were inversely related to both rainfall and throughfall depth. Oxidized N concentrations in streamflow were greatest during the rising limb of storm flow with subsequent decreases during the falling limb of storm hydrographs and lowest during base flow. The oxidized N inputs from bulk precipitation were considerably greater than outputs from streamflow resulting in a net retention of oxidized nitrogen within the watershed during the study period.

Water Resources Bulletin↗

Effects of agricultural nutrient management on nitrogen fate and transport in Lancaster County, Pennsylvania

Nitrogen inputs to, and outputs from, a 55-acre site in Lancaster County, Pennsylvania, were estimated to determine the pathways and relative magnitude of loads of nitrogen entering and leaving the site, and to compare the loads of nitrogen before and after the implementation of nutrient management. Inputs of nitrogen to the site were manure fertilizer, commercial fertilizer, nitrogen in precipitation, and nitrogen in ground-water inflow; and these sources averaged 93, 4, 2, and 1 percent of average annual nitrogen additions, respectively. Outputs of nitrogen from the site were nitrogen in harvested crops, loads of nitrogen in surface runoff, volatilization of nitrogen, and loads of nitrogen in ground-water discharge, which averaged 37, less than 1,25, and 38 percent of average annual nitrogen removals from the site, respectively. Virtually all of the nitrogen leaving the site that was not removed in harvested crops or by volatilization was discharged in the ground water. Applications of manure and fertilizer nitrogen to 47.5 acres of cropped fields decreased about 33 percent, from an average of 22,700 pounds per year (480 pounds per acre per year) before nutrient management to 15,175 pounds of nitrogen per year (320 pounds per acre per year) after the implementation of nutrient management practices. Nitrogen loads in ground-water discharged from the site decreased about 30 percent, from an average of 292 pounds of nitrogen per million gallons of ground water before nutrient management to an average of 203 pounds of nitrogen per million gallons as a result of the decreased manure and commercial fertilizer applications. Reductions in manure and commercial fertilizer applications caused a reduction of approximately 11,000 pounds (3,760 pounds per year, 70 pounds per acre per year) in the load of nitrogen discharged in ground water from the 55-acre site during the three-year period 1987-1990.

Water Resources Bulletin↗

Biogeochemical effects of global change on U.S. National Parks

Federal parks and other public lands have unique mandates and rules regulating their use and conservation. Because of variation in their response to local, regional, and global-scale disturbance, development of mitigation strategies requires substantial research in the context of long-term inventory and monitoring. In 1982, the National Park Service began long-term, watershed-level studies in a series of national parks. The objective was to provide a more comprehensive database against which the effects of global change and other issues could be quantified. A subset of five sites in North Carolina, Texas, Washington, Michigan, and Alaska, is examined here. During the last 50 years, temperatures have declined at the southern sites and increased at the northern sites with the greatest increase in Alaska. Only the most southern site has shown an increase in precipitation amount. The net effect of these trends, especially for the most northern and southern sites, would likely be an increase in the growing season and especially the time soil processes could continue without moisture or temperature limitations. During the last 18 years, there were few trends in atmospheric ion inputs. The most evident was the decline in SO 4 2 deposition. There were no significant relationships between ion input and stream water output. This finding suggests other factors as modification of precipitation or canopy throughfall by soil processes, hydrologic flow path, and snowmelt rates are major processes regulating stream water chemical outputs.

Alaska, Michigan, North Carolina, Texas, Washingto↗