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At least 955 records · Page 53Linked to original sources

Increasing seasonal variation in the extent of rivers and lakes from 1984 to 2022

Knowledge of the spatial and temporal distribution of surface water is important for water resource management, flood risk assessment, monitoring ecosystem health, constraining estimates of biogeochemical cycles and understanding our climate. While global-scale spatiotemporal change detection of surface water has significantly improved in recent years due to planetary-scale remote sensing and computing, it has remained challenging to distinguish the changing characteristics of rivers and lakes. Here we analyze the spatial extent of permanent and seasonal rivers and lakes globally over the past 38 years based on new data of river system extents and surface water trends. Results show that while the total permanent surface area of both rivers and lakes has remained relatively constant, the areas with intermittent seasonal coverage have increased by 12 % and 27 % for rivers and lakes, respectively. The increase is statistically significant in over 84 % of global water catchments based on Spearman's rank correlations (rho) above 0.05 and p values less than 0.05. The seasonal river extent is nearly 32 % larger than the previously observed annual mean river extent, suggesting large seasonal variations that impact not only ecosystem health but also estimations of terrestrial biogeochemical cycles of carbon. The outcomes of our analysis are shared as the Surface Area of Rivers and Lakes (SARL) database, serving as a valuable resource for monitoring and research of hydrological cycles, ecosystem accounting, and water management.

Hydrology and Earth System Sciences↗

Warming in the upper San Francisco Estuary: Patterns of water temperature change from five decades of data

Temperature is a key controlling variable from subcellular to ecosystem scales. Thus, climatic warming is expected to have broad impacts, especially in economically and ecologically valuable systems such as estuaries. The heavily managed upper San Francisco Estuary supplies water to millions of people and is home to fish species of high conservation, commercial, and recreational interest. Despite a long monitoring record (> 50 yr), we do not yet know how water temperatures have already changed or how trends vary spatially or seasonally. We fit generalized additive models on an integrated database of discrete water temperature observations to estimate long-term trends with spatio-seasonal variability. We found that water temperatures have increased 0.017°C yr −1 on average over the past 50 yr. Rates of temperature change have varied over time, but warming was predominant. Temperature increases were most widespread in the late-fall to winter (November to February) and mid-spring (April to June), coinciding with the winter development of juvenile Chinook salmon and spring spawning window of the endangered delta smelt. Warming was fastest in the northern regions, a key fish migration corridor with important tidal wetland habitat. However, no long-term temperature trends were detected in October and were only observed in some regions in May, July, and August. These results can help identify optimal areas for restoration or refugia to buffer the effects of a warming climate, and the methods can be leveraged to understand the spatiotemporal variability in climate warming patterns in other aquatic systems.

California↗

Comparative phylogeography of a coevolved community: Concerted population expansions in Joshua trees and four Yucca moths

Comparative phylogeographic studies have had mixed success in identifying common phylogeographic patterns among co-distributed organisms. Whereas some have found broadly similar patterns across a diverse array of taxa, others have found that the histories of different species are more idiosyncratic than congruent. The variation in the results of comparative phylogeographic studies could indicate that the extent to which sympatrically-distributed organisms share common biogeographic histories varies depending on the strength and specificity of ecological interactions between them. To test this hypothesis, we examined demographic and phylogeographic patterns in a highly specialized, coevolved community - Joshua trees (Yucca brevifolia) and their associated yucca moths. This tightly-integrated, mutually interdependent community is known to have experienced significant range changes at the end of the last glacial period, so there is a strong a priori expectation that these organisms will show common signatures of demographic and distributional changes over time. Using a database of >5000 GPS records for Joshua trees, and multi-locus DNA sequence data from the Joshua tree and four species of yucca moth, we combined paleaodistribution modeling with coalescent-based analyses of demographic and phylgeographic history. We extensively evaluated the power of our methods to infer past population size and distributional changes by evaluating the effect of different inference procedures on our results, comparing our palaeodistribution models to Pleistocene-aged packrat midden records, and simulating DNA sequence data under a variety of alternative demographic histories. Together the results indicate that these organisms have shared a common history of population expansion, and that these expansions were broadly coincident in time. However, contrary to our expectations, none of our analyses indicated significant range or population size reductions at the end of the last glacial period, and the inferred demographic changes substantially predate Holocene climate changes.

PLoS ONE↗

Deposition, exhumation, and paleoclimate of an ancient lake deposit, Gale crater, Mars

The landforms of northern Gale crater on Mars expose thick sequences of sedimentary rocks. Based on images obtained by the Curiosity rover, we interpret these outcrops as evidence for past fluvial, deltaic, and lacustrine environments. Degradation of the crater wall and rim probably supplied these sediments, which advanced inward from the wall, infilling both the crater and an internal lake basin to a thickness of at least 75 meters. This intracrater lake system probably existed intermittently for thousands to millions of years, implying a relatively wet climate that supplied moisture to the crater rim and transported sediment via streams into the lake basin. The deposits in Gale crater were then exhumed, probably by wind-driven erosion, creating Aeolis Mons (Mount Sharp).

Science↗

Hydroclimate and fire paleorecords across the southern Rockies and Colorado Plateau over the common era

The southwestern US has been experiencing a severe drought and increased fire activity over the past two decades, affecting people’s health, homes, and businesses. Many individual fires occurring in the Southwest are the most severe in recorded history both in terms of dollars of damages as well as in fire extent. It is essential to be able to place the present drought and fires into the context of the current changing climate as well as in the framework of climate variability and human activity over centennial to millennial timescales. Dendrochronology can determine both the seasonality of fires as well as changes in precipitation. Charcoal, pollen, and the biomarkers levoglucosan, mannosan, and galactosan can help determine fire activity and the type of burned vegetation. People have controlled and utilized fire in the southwestern US for thousands of years. While proxies cannot yet determine if people ignited a specific fire, fecal sterols can determine if people were within an individual watershed. Here, we examine the strengths and weaknesses of using high-resolution tree-ring data in conjunction with biomarkers in ice and lake cores from the southwestern US to study interactions between changes in hydroclimate, fires, and human activity.

Conference Paper↗

Pleistocene porcupine (Erethizontidae) records in arid southwestern North America and comparisons with the modern distribution of Erethizon dorsatum in southern California and Arizona

he North American porcupine ( Erethizon dorsatum ) is widely distributed throughout many ecosystems on the continent from northern Mexico to the Arctic Ocean, except for the southeastern portion of the United States. Habi­tats include the arid Desert Southwest region where modern records are generally sparse. The paleogeographic range in the Desert Southwest is not unlike the modern distribution of Erethizon , with some exceptions. Although there are early Pleistocene records of Erethizon for southern California, no late Pleistocene records (Rancholabrean North American Land Mammal Age [NALMA]) are known for Imperial, Inyo, Riverside, or San Bernardino Counties in southern California or in Mexico. This is surprising given late-Pleistocene records in Arizona, Nevada, and New Mexico and modern records from southern California and Mexico. Pleistocene habitats for porcupines were similar to those occupied today, just at different elevations due to differing climates. Modern preferred habitats in the Desert Southwest include conifer-clad high-elevation areas and associated pinyon pine / juniper belts, and occasionally riparian corridors in desert environments. The overall rarity of porcupines today in arid parts of the Desert Southwest is likely due to the combination of increasing aridity in the region during the Holocene, past persecution by humans, and increasing predator populations.

Arizona, California↗

Fire and aquatic ecosystems in forested biomes of North America

Synthesis of the literature suggests that physical, chemical, and biological elements of a watershed interact with long‐term climate to influence fire regime, and that these factors, in concordance with the postfire vegetation mosaic, combine with local‐scale weather to govern the trajectory and magnitude of change following a fire event. Perturbation associated with hydrological processes is probably the primary factor influencing postfire persistence of fishes, benthic macroinvertebrates, and diatoms in fluvial systems. It is apparent that salmonids have evolved strategies to survive perturbations occurring at the frequency of wildland fires (10°–10 2 years), but local populations of a species may be more ephemeral. Habitat alteration probably has the greatest impact on individual organisms and local populations that are the least mobile, and reinvasion will be most rapid by aquatic organisms with high mobility. It is becoming increasingly apparent that during the past century fire suppression has altered fire regimes in some vegetation types, and consequently, the probability of large stand‐replacing fires has increased in those areas. Current evidence suggests, however, that even in the case of extensive high‐severity fires, local extirpation of fishes is patchy, and recolonization is rapid. Lasting detrimental effects on fish populations have been limited to areas where native populations have declined and become increasingly isolated because of anthropogenic activities. A strategy of protecting robust aquatic communities and restoring aquatic habitat structure and life history complexity in degraded areas may be the most effective means for insuring the persistence of native biota where the probability of large‐scale fires has increased.

Transactions of the American Fisheries Society↗

Climate change and Arctic ecosystems: 2. Modeling, paleodata-model comparisons, and future projections

Large variations in the composition, structure, and function of Arctic ecosystems are determined by climatic gradients, especially of growing-season warmth, soil moisture, and snow cover. A unified circumpolar classification recognizing five types of tundra was developed. The geographic distributions of vegetation types north of 55°N, including the position of the forest limit and the distributions of the tundra types, could be predicted from climatology using a small set of plant functional types embedded in the biogeochemistry-biogeography model BIOME4. Several palaeoclimate simulations for the last glacial maximum (LGM) and mid-Holocene were used to explore the possibility of simulating past vegetation patterns, which are independently known based on pollen data. The broad outlines of observed changes in vegetation were captured. LGM simulations showed the major reduction of forest, the great extension of graminoid and forb tundra, and the restriction of low- and high-shrub tundra (although not all models produced sufficiently dry conditions to mimic the full observed change). Mid-Holocene simulations reproduced the contrast between northward forest extension in western and central Siberia and stability of the forest limit in Beringia. Projection of the effect of a continued exponential increase in atmospheric CO 2 concentration, based on a transient ocean-atmosphere simulation including sulfate aerosol effects, suggests a potential for larger changes in Arctic ecosystems during the 21st century than have occurred between mid-Holocene and present. Simulated physiological effects of the CO 2 increase (to >700 ppm) at high latitudes were slight compared with the effects of the change in climate.

Journal of Geophysical Research D: Atmospheres↗

Northern forest winters have lost cold, snowy conditions that are important for ecosystems and human communities

Winter is an understudied but key period for the socio-ecological systems of northeastern North American forests. A growing awareness of the importance of the winter season to forest ecosystems and surrounding communities has inspired several decades of research, both across the northern forest and at other mid- and high-latitude ecosystems around the globe. Despite these efforts, we lack a synthetic understanding of how winter climate change may impact hydrological and biogeochemical processes and the social and economic activities they support. Here we take advantage of 100 years of meteorological observations across the northern forest region of the northeastern U.S. and eastern Canada to develop a suite of indicators that enable a cross-cutting understanding of 1) how winter temperatures and snow cover have been changing and 2) how these shifts may impact both ecosystems and surrounding human communities. We show that cold and snow-covered conditions have generally decreased over the past 100 years. These trends suggest positive outcomes for tree health as related to reduced fine root mortality and nutrient loss associated with winter frost but negative outcomes as related to the northward advancement and proliferation of forest insect pests. In addition to effects on vegetation, reductions in cold temperatures and snow cover are likely to have negative impacts on the ecology of the northern forest through impacts on water, soils, and wildlife. The overall loss of coldness and snow cover may also have negative consequences for logging and forest products, vector-borne diseases and human health, recreation and tourism, and cultural practices, which together represent important social and economic dimensions for the northern forest region. These findings advance our understanding of how our changing winters may transform the socio- ecological system of a region that has been defined by the contrasting rhythm of the seasons. Our research also identifies a trajectory of change that informs our expectations for the future as the climate continues to warm.

Ecological Applications↗

A model of strength

In her AAAS News & Notes piece "Can the Southwest manage its thirst?" (26 July, p. 362), K. Wren quotes Ajay Kalra, who advocates a particular method for predicting Colorado River streamflow "because it eschews complex physical climate models for a statistical data-driven modeling approach." A preference for data-driven models may be appropriate in this individual situation, but it is not so generally, Data-driven models often come with a warning against extrapolating beyond the range of the data used to develop the models. When the future is like the past, data-driven models can work well for prediction, but it is easy to over-model local or transient phenomena, often leading to predictive inaccuracy (1). Mechanistic models are built on established knowledge of the process that connects the response variables with the predictors, using information obtained outside of an extant data set. One may shy away from a mechanistic approach when the underlying process is judged to be too complicated, but good predictive models can be constructed with statistical components that account for ingredients missing in the mechanistic analysis. Models with sound mechanistic components are more generally applicable and robust than data-driven models.

Science↗

The tundra phenology database: More than two decades of tundra phenology responses to climate change

Observations of changes in phenology have provided some of the strongest signals of the effects of climate change on terrestrial ecosystems. The International Tundra Experiment (ITEX), initiated in the early 1990s, established a common protocol to measure plant phenology in tundra study areas across the globe. Today, this valuable collection of phenology measurements depicts the responses of plants at the colder extremes of our planet to experimental and ambient changes in temperature over the past decades. The database contains 150 434 phenology observations of 278 plant species taken at 28 study areas for periods of 1–26 years. Here we describe the full data set to increase the visibility and use of these data in global analyses and to invite phenology data contributions from underrepresented tundra locations. Portions of this tundra phenology database have been used in three recent syntheses, some data sets are expanded, others are from entirely new study areas, and the entirety of these data are now available at the Polar Data Catalogue ( https://doi.org/10.21963/13215 ).

Arctic Science↗

Data Report: Intra-annual variability of the diatom assemblages at Hole 1034B (Saanich Inlet) near 9 ka

Ocean Drilling Program (ODP) Site 1034 (48°38.000´N, 123°30.000´W) was drilled at a water depth of 200 m in the Saanich Inlet, an anoxic fjord on the southeastern coast of Vancouver Island, British Columbia, to a depth of 118.2 meters below seafloor (mbsf). The uppermost 50 m consists of very well-laminated (triplet varves) diatomaceous muds deposited over the past 7000 yr. Below, sediments become progressively less distinctly laminated and reflect better oxygenated bottom-water conditions. The oldest sediments recovered at Site 1034 were dated as 14 to 15 ka (see Shipboard Scientific Party, 1998). Varved sediments recovered during Leg 169S in the Saanich Inlet offer an excellent opportunity to compare the interannual variability of recent climatic elements (e.g., sea-surface temperature, salinity, precipitation) with those of late Quaternary periods that were characterized by climates appreciably different from those of today. A large amount of recent diatom data from Saanich Inlet exists, which can serve as a baseline for such comparisons: Sancetta (1989a, 1989b, 1990) investigated modern processes controlling the accumulation of diatoms and spacial and temporal trends of diatom flux in the Inlet; Sancetta and Calvert (1988) documented the annual cycle of sedimentation in the fjord. McQuoid (1995) and McQuoid and Hobson (1997) studied the modern pattern of diatom succession in the Saanich Inlet and analyzed the diatoms in laminae couplets in frozen sediment cores for the years 1900 to 1991 A.D. During the hypsithermal warming of the early Holocene (~10-6 ka), climatic conditions throughout much of northern North America were warmer and drier than those of the present (Pielou, 1991; Hebda and Whitlock, 1997), largely as a result of increased solar insolation, which peaked between 10 and 9 ka at 65°N (Berger and Loutre, 1991). Temperatures are estimated to have been 2° to 4°C warmer than today for most of this interval, reaching a maximum between ~9 and 7 ka (Hebda and Whitlock, 1997). According to Heusser (1983) and Heusser (1985), rapid warming occurred at ~10 ka in southwestern British Columbia with summer conditions that were drier and as warm or warmer than today lasting until ~6 ka. Clague and Mathewes (1989) report that treeline elevation in the southeastern coast mountains of British Columbia reached elevations that were between 60 and 130 m higher than today between 9.1 and 8.2 ka. Thompson et al. (1993) argue that the driest conditions (period of maximum summer drought) of the Holocene were reached in western North America at 9 ka. The warmer and drier conditions of this Holocene thermal maximum were gradually replaced by cooler and wetter conditions (Hebda, 1995; Hebda and Whitlock, 1997). This report describes the intra-annual variability of the diatom assemblages at Site 1034, during a 8-yr interval near 9 ka. Pollen and dinoflagellates are being studied from the same samples by R. Hebda and P. Mudie (unpubl. data).

British Columbia↗

Reconnaissance hydrology of Portage Glacier basin, Alaska

Early reports of conditions in Portage Pass, Alaska, provide evidence that Portage Glacier was formerly larger and thicker. Past conditions, recent history, current retreat, and possible future changes are summarized from an analysis of reports, photographs of the glacier (1939, 1950, and annually since about 1960), and data on snow and ice balance and bathymetry (1972). Between 1900 and 1972, the glacier terminus retreated 3.4 kilometers, and the lower part of the glacier thinned 200 meters. Climatic change controlled the retreat until about 1930; since then deep water at the terminus has influenced the calving retreat. The calving rate and present terminus position cannot be sustained by current climatic conditions and rate of snow accumulation. Thus the glacier will continue to recede until the terminus stabilizes in shallower water, probably about 1.5 kilometers upvalley from the present terminus and in about year 2020, assuming no change in present climatic conditions and calving rate. Possible small climatic changes could cause a shift in the point at which annual snow accumulation equals annual ablation (500 meters) and a corresponding change in terminus behavior. Potential natural hazards include avalanches, outburst floods from ice-dammed lakes, and unstable icebergs.

Alaska↗

Informing our successors: What botanical information for Santa Cruz Island will researchers and conservation managers in the century ahead need the most?

Climate changes are predicted to drive changes in plant species composition and vegetation cover around the world. Preserved specimens and other botanical information that we gather today—a period future practitioners may look back on as an early stage of modern anthropogenic climate change—will be of value to conservation managers and conservation biologists in the decades and centuries ahead. Here, we present suggestions for the systematic collection, long-term curation (in museums, herbaria, and other research institutions), and maintenance of plant specimens, along with associated data and analyses on the plants and vegetation present today and in the past. The primary aim of this systematic survey is to provide information of high value to conservation researchers and managers both in the near term (the next several years) and through the century to come. Such a systematic survey would build on a strong foundation of research and adaptive management on the island. It would fill gaps in less well-studied groups of organisms and identify environmental, ecological, and cultural factors related to current patterns of distribution. It would also archive previously collected data, photographs, and other materials which would otherwise gradually degrade and become inaccessible. As a case study, we use Santa Cruz Island, California, which is managed for conservation. We are confident that the same approach may be applied to other lands and waters around the world. We argue that there is a particular need to collect and archive herbarium specimens and seeds from today's populations, activities largely overlooked in recent decades. We encourage conservation researchers and managers to consider what information will be most important for future managers and to help launch studies, monitoring programs, and collections to prepare their successors for success.

Western North American Naturalist↗

Past and predicted future effects of housing growth on open space conservation opportunity areas and habitat connectivity around National Wildlife Refuges

Context Housing growth can alter suitability of matrix habitats around protected areas, strongly affecting movements of organisms and, consequently, threatening connectivity of protected area networks. Objectives Our goal was to quantify distribution and growth of housing around the U.S. Fish and Wildlife Service National Wildlife Refuge System. This is important information for conservation planning, particularly given promotion of habitat connectivity as a climate change adaptation measure. Methods We quantified housing growth from 1940 to 2000 and projected future growth to 2030 within three distances from refuges, identifying very low housing density open space, “opportunity areas” (contiguous areas with <6.17 houses/km 2 ), both nationally and by USFWS administrative region. Additionally, we quantified number and area of habitat corridors within these opportunity areas in 2000. Results Our results indicated that the number and area of open space opportunity areas generally decreased with increasing distance from refuges and with the passage of time. Furthermore, total area in habitat corridors was much lower than in opportunity areas. In addition, the number of corridors sometimes exceeded number of opportunity areas as a result of habitat fragmentation, indicating corridors are likely vulnerable to land use change. Finally, regional differences were strong and indicated some refuges may have experienced so much housing growth already that they are effectively too isolated to adapt to climate change, while others may require extensive habitat restoration work. Conclusions Wildlife refuges are increasingly isolated by residential housing development, potentially constraining the movement of wildlife and, therefore, their ability to adapt to a changing climate.

Landscape Ecology↗

Satellite-based evidence of recent decline in global forest recovery rate from tree mortality events

Climate-driven forest mortality events have been extensively observed in recent decades, prompting the question of how quickly these affected forests can recover their functionality following such events. Here we assessed forest recovery in vegetation greenness (normalized difference vegetation index) and canopy water content (normalized difference infrared index) for 1,699 well-documented forest mortality events across 1,600 sites worldwide. By analysing 158,427 Landsat surface reflectance images sampled from these sites, we provided a global assessment on the time required for impacted forests to return to their pre-mortality state (recovery time). Our findings reveal a consistent decline in global forest recovery rate over the past decades indicated by both greenness and canopy water content. This decline is particularly noticeable since the 1990s. Further analysis on underlying mechanisms suggests that this reduction in global forest recovery rates is primarily associated with rising temperatures and increased water scarcity, while the escalation in the severity of forest mortality contributes only partially to this reduction. Moreover, our global-scale analysis reveals that the recovery of forest canopy water content lags significantly behind that of vegetation greenness, implying that vegetation indices based solely on greenness can overestimate post-mortality recovery rates globally. Our findings underscore the increasing vulnerability of forest ecosystems to future warming and water insufficiency, accentuating the need to prioritize forest conservation and restoration as an integral component of efforts to mitigate climate change impacts.

Nature Plants↗

Past, present and future of a meandering river in the Bolivian Amazon basin

Field observations on small rivers of the Amazon basin are less common due to their remote location and difficult accessibility. Here we show, through remote sensing analysis and field works, the planform evolution and riverbed topography of a small river located in the upper foreland Amazon basin, the Ichilo River. By tracking planform changes over 30 years, we identified the factors that control meander migration rates in the Ichilo River: cutoffs, climate and human interventions. The data suggest that neck cutoffs are the main controls in the Ichilo River, with an annual density of 0.022 cutoffs/km. In addition, climate controls have been identified in the form of high-precipitation events that may have promoted cutoffs, an increase in meander migration rate and channel widening. The width distribution of the Ichilo River is well represented by general extreme value and inverse Gaussian distributions. The spatiotemporal variability of meandering migration rates in the Ichilo River is analysed in two locations where neck cutoffs are expected. Analysing the distance across the neck in these two points, we predict the occurrence of a new cutoff. The combined methodology of bathymetric surveys and structure from motion photogrammetry shows us the Ichilo riverbed topography and banks at high resolution, where two scour holes were identified. Finally, we discuss the impact of planform changes of the Ichilo River on communities that are established along its riverbanks.

Earth Surface Processes and Landforms↗

Pluvial lakes in the Great Basin of the western United States: a view from the outcrop

Paleo-lakes in the western United States provide geomorphic and hydrologic records of climate and drainage-basin change at multiple time scales extending back to the Miocene. Recent reviews and studies of paleo-lake records have focused on interpretations of proxies in lake sediment cores from the northern and central parts of the Great Basin. In this review, emphasis is placed on equally important studies of lake history during the past ∼30 years that were derived from outcrop exposures and geomorphology, in some cases combined with cores. Outcrop and core records have different strengths and weaknesses that must be recognized and exploited in the interpretation of paleohydrology and paleoclimate. Outcrops and landforms can yield direct evidence of lake level, facies changes that record details of lake-level fluctuations, and geologic events such as catastrophic floods, drainage-basin changes, and isostatic rebound. Cores can potentially yield continuous records when sampled in stable parts of lake basins and can provide proxies for changes in lake level, water temperature and chemistry, and ecological conditions in the surrounding landscape. However, proxies such as stable isotopes may be influenced by several competing factors the relative effects of which may be difficult to assess, and interpretations may be confounded by geologic events within the drainage basin that were unrecorded or not recognized in a core. The best evidence for documenting absolute lake-level changes lies within the shore, nearshore, and deltaic sediments that were deposited across piedmonts and at the mouths of streams as lake level rose and fell. We review the different shorezone environments and resulting deposits used in such reconstructions and discuss potential estimation errors. Lake-level studies based on deposits and landforms have provided paleohydrologic records ranging from general changes during the past million years to centennial-scale details of fluctuations during the late Pleistocene and Holocene. Outcrop studies have documented the integration histories of several important drainage basins, including the Humboldt, Amargosa, Owens, and Mojave river systems, that have evolved since the Miocene within the active tectonic setting of the Great Basin; these histories have influenced lake levels in terminal basins. Many pre-late Pleistocene lakes in the western Great Basin were significantly larger and record wetter conditions than the youngest lakes. Outcrop-based lake-level data provide important checks on core-based proxy interpretations; we discuss four such comparisons. In some cases, such as for Lakes Owens and Manix, outcrop and core data synthesis yields stronger and more complete records; in other cases, such as for Bonneville and Lahontan, conflicts point toward reconsideration of confounding factors in interpretation of core-based proxies.

Great Basin↗