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Long Term Resource Monitoring Program procedures: fish monitoring

This manual constitutes the second revision of the U.S. Army Corps of Engineers’ Upper Mississippi River Restoration-Environmental Management Program (UMRR-EMP) Long Term Resource Monitoring Program (LTRMP) element Fish Procedures Manual. The original (1988) manual merged and expanded on ideas and recommendations related to Upper Mississippi River fish sampling presented in several early documents. The first revision to the manual was made in 1995 reflecting important protocol changes, such as the adoption of a stratified random sampling design. The 1995 procedures manual has been an important document through the years and has been cited in many reports and scientific manuscripts. The resulting data collected by the LTRMP fish component represent the largest dataset on fish within the Upper Mississippi River System (UMRS) with more than 44,000 collections of approximately 5.7 million fish. The goal of this revision of the procedures manual is to document changes in LTRMP fish sampling procedures since 1995. Refinements to sampling methods become necessary as monitoring programs mature. Possible refinements are identified through field experiences (e.g., sampling techniques and safety protocols), data analysis (e.g., planned and studied gear efficiencies and reallocations of effort), and technological advances (e.g., electronic data entry). Other changes may be required because of financial necessity (i.e., unplanned effort reductions). This version of the LTRMP fish monitoring manual describes the most current (2014) procedures of the LTRMP fish component.

Illinois;Iowa;Minnesota;Montana;Wisconsin↗

Inaccuracies in sediment budgets arising from estimations of tributary sediment inputs: an example from a monitoring network on the southern Colorado plateau

Sediment budgets are an important tool for understanding how riverine ecosystems respond to perturbations. Changes in the quantity and grain-size distribution of sediment within river systems affect the channel morphology and related habitat resources. It is therefore important for resource managers to know if a channel reach is in a state of sediment accumulation, deficit or stasis. Many studies have estimated sediment loads from ungaged tributaries using regional sediment-yield equations or other similar techniques. While these approaches may be valid in regions where rainfall and geology are uniform over large areas, use of sediment-yield equations may lead to poor estimations of sediment loads in semi-arid climates, where rainfall events, contributing geology, and vegetation have large spatial variability.

Arizona↗

Crocodylus acutus (American Crocodile). Long distance juvenile movement

Crocodylus acutus (American Crocodile) is the most widely distributed New World crocodilian species with its range extending from Peru in the south to the southern tip of peninsular Florida in the north. Crocodylus acutus occupies primarily coastal brackish water habitat, however it also occurs in freshwater to hypersaline habitats (Thorbjarnarson 2010. In Crocodiles. Status Survey and Conservation Action Plan. [Third Edition], American Crocodile Crocodylus acutus, pp. 46–53 S.C. Manolis and C. Stevenson. Crocodile Specialist Group, Darwin). There is limited literature on long distance movements of juvenile crocodilians worldwide and no literature on juvenile crocodiles in Florida. However, adult C. acutus in Florida have been documented to make seasonal movements of 5–15 km from preferred foraging habitat to nesting beaches (Mazzotti 1983. The Ecology of Crocodylus acutus in Florida. PhD Dissertation. The Pennsylvania State University, University Park, Pennsylvania. 161pp), and one adult was documented making a 35 km trip from her nest site to preferred foraging habitat (Cherkiss et. al. 2006. Herpetol. Rev. 38:72–73). Rodda (1984. Herpetologica 40:444–451) reported on juvenile C. acutus movement in Gatun Lake, Panama, and found that juveniles stayed within 1 km of their nest site for the first month. Movements of juvenile Crocodylus porosus (Saltwater Crocodile) in a river system in Northern Australia showed a maximum movement of 38.9 km from a known nest site, with the majority of the crocodiles staying within 15.6 km downstream to 6.8 km upstream (Webb and Messel 1978. Aust. Wildlife Res. 5:263–283). Juvenile movement of Crocodylus niloticus (Nile Crocodile) in Lake Ngezi, Zimbabwe showed crocodiles restricted their movements from 1.0 km up to 4.5 km through the wet and dry seasons (Hutton 1989. Am. Zool. 29:1033–1049). Long distance movements of alligators were recorded for sizes ranging from 28 cm to 361 cm in a coastal refuge in Louisiana, where the distance traveled ranged from 0.3 km to 90.2 km. The data showed that the smaller alligators moved greater distance than larger ones (Lance et al. 2011. Southeast Nat. 10:389–398). An ongoing 30 year mark and recapture study for Crocodylus acutus in Florida allowed us to look at long distance movement (>30 km) of juveniles (30km). Initial and most recent captures as a juvenile were used to analyze distances moved (Fig. 1). These distances were measured linearly between capture locations. Maximum linear distances of 76.3 km and 69.6 km were recorded for animals 4838 and 6662. All crocodiles moved from nesting habitat through potentially optimal nursery habitat prior to reaching their recapture locations. These juvenile long distance movements could be due to larger crocodiles facilitating their dispersal from the nest location (Lance et al. 2011. op. cit.). These data (Table 1.) support that there is exchange of individuals among the nesting colonies and our ongoing efforts to monitor this threatened species allow us to make observations of how juvenile crocodiles are moving throughout the landscape in an ecosystem currently undergoing restoration.

Herpetological Review↗

Elwha River dam removal: A major opportunity for salmon and steelhead recolonization

In this in-depth paper, authors George R. Pess, Gary A. Winans and Timothy J. Beechie of the NOAA Fisheries, Northwest Fisheries Science Center in Seattle, Samuel J. Brenkman of the National Park Service, Olympic National Park, Michael L. McHenry of the Lower Elwha Klallam Tribe and Jeffrey J. Duda of the U.S. Geological Survey, Western Fisheries Research Center in Seattle, provide an historical overview of the Elwha River system, and its native anadromous fish runs and the prospect of their recolonization after the Elwha and Glines Canyon dams are removed.

Washington↗

Evaluation of Alaskan wetlands for waterfowl

Few studies have focused specifically on use of Alaskan wetlands by waterfowl. However, substantial information on the values of wetlands is available from studies on individual species or that were conducted for other purposes. Most investigators have found it most effective to classify habitat use on the basis of observed distribution patterns of waterfowl in relation to local physiographic features and vegetation types. Major variations in habitat among regions result from differences in precipitation, summer temperature, length of frost free period, and presence of permafrost. Large areas of intertidal habitat, usually associated with river deltas, are of key importance wherever they occur. Lakes subject to extensive periods of draw-down or that fluctuate with river systems are more productive than those with more stable water levels.

Conference Paper↗

Understanding aquatic animal virus survival and trafficking and its role in risk assessment

The stability of infectious agents in different media and under different physical and chemical environments has been extensively studied for some viruses and virtually ignored for others. Gaps in our knowledge are due in part to difficulties in reproducing virus «life cycles» and determination if the agent is in fact inactive. Additionally, isolation of the agent under certain conditions can present significant challenges. Studies on the susceptibility of viruses to different physical or chemical parameters have often been conducted under artificial conditions and quantitative data on the rate of inactivation are lacking for many agents. Using infectious hematopoietic necrosis virus (IHNV) as an example, survival was assessed under different environmental conditions. Three IHNV isolates that exhibited antigenic and genetic differences were diluted either in freshwater collected from a spring, after it passed through a fish farm, or the river that received water from the fish farm. Each treatment was incubated at 15o C in a water bath and samples were removed daily. Virus concentrations were determined by plaque assay on EPC cells. Virus suspended in spring water survived longer than virus incubated in water obtained from a fish farm or the river. Virus suspended in river water exhibited a 99.99% reduction in virus concentrations in less than 24 h. Survival of IHNV was also evaluated at different temperatures. A 1982 isolate appeared to be less temperature sensitive than isolates collected in 1990. A preliminary study was also conducted to determine the genetic similarity of IHNV isolates present downstream in a river system from the state of Idaho. Isolates were analyzed using the RNase protection assay (RPA) and by nucleotide sequencing of RT-PCR products of specific isolates. Genetic typing of IHNV allows monitoring of virus traffic and may provide insight into the epizootiology and mechanisms of virus spread. These results illustrate the complexity in evaluating virus survival and trafficing and using this sort of information in risk assessment.

Conference Paper↗

Non-lethal estimation of body composition of Yukon River salmon

Because of the importance of Chinook salmon to commercial and subsistence fisheries on the Yukon River, further study of the factors that may affect the success of this species and our ability to manage the fisheries is warranted. Critical to these studies is the determination of the amount of lipids (fat) stored and available to the fish as its primary energy source for migration and spawning. Recent developments of Bioelectrical Impedance Analysis (BIA) promise a simple, non-lethal means of estimating proximate composition (e.g. fat, protein, water content) for field applications with fish. The goal of the project was to develop BIA models for Chinook salmon from the Yukon River watershed that would permit the non-lethal estimation of body proximate composition for use in field studies. Our results clearly demonstrated that BIA can be used to estimate proximate composition and energy density of salmon. While some minor refinements were suggested, the methodology can be used in a wide variety of field applications. For instance, application of the BIA models to predict energy levels of fish during their migration will allow evaluation of management programs, while also yielding data that can be used to evaluate energy use along the migratory path. Correlations of energy level with ongoing tagging, radio-tracking, and genetic studies also have the potential to allow managers and scientists to understand the relationship between fat content and distance to spawning location. These models have the potential for application to this species in other river systems. They also provide tools for a variety of other scientific investigation such as: 1) differences in energy stores in spawning and recruitment success; 2) effects of global warming on migratory salmonid stocks; and 3) differences in annual flow and temperature regiments upon migratory energy costs and resulting recruitment success.

Alaska, Yukon↗

To sell or not sell: Assessments of Bangladesh hydrocarbons

A decision by the government of Bangladesh to sell or not sell some of its natural gas reserves to neighboring countries in South Asia will be important in determining the economic future of Bangladesh, a country with an area about equal to Wisconsin. Bangladesh is a country of 150 million people, many of whom live at or below the poverty line. It is situated almost entirely on the great low-lying delta of the Ganges and Brahmaputra River systems. Folded Tertiary strata that form hill tracts in easternmost Bangladesh, adjacent India, and Myanmar provide a little relief above a monotonous deltaic terrain (Fig. 1).

Oil & Gas Journal↗

Instream flows to assist the recovery of endangered fishes of the upper Colorado River basin

The riverine landscape of the upper Colorado River basin has been extensively modified by dams, diversions, revetments, and water abstractions. These changes, probably coupled with the introduction of many nonnative fishes, have compromised the existence of four of the native fishes (Colorado River squawfish Ptychocheilus lucius, humpback chub Gila cypha, bonytail chub Gila elegans, and razorback sucker Xyrauchen texanus) of the river system. Efforts to recover these endangered fishes have emphasized reregulation of flows to provide better habitat conditions than existed during the last half century, when ranges and abundances of the fishes declined significantly. Contention emerged, however, with regard to the efficacy of methods used by the U.S. Fish and Wildlife Service to justify flow recommendations to protect the endangered fishes. The purpose of this study was to review the science pertaining to the issue of flow provision, to identify critical uncertainties, and to provide recommendations for determining the instream flow needs of the endangered fishes.

Biological Report↗

Mapping the thermal landscape of the Upper Mississippi River

Temperature has a fundamental influence on physical, chemical and biological processing in aquatic ecosystems. River temperatures respond to a diverse array of drivers including air temperature, streamflow, and thermal inputs, but the physical template has been shown to play a significant role in structuring spatial and temporal variation in water temperature. How these factors interact to affect water temperature in complex floodplain river habitats such as those present in the Upper Mississippi River System (UMRS) is not well-studied. We used a combination of airborne thermal imagery and continuous temperature loggers deployed across aquatic area types to evaluate spatial and temporal patterns in water temperature in Navigation Pool 8 during the summer and fall of 2017. The mid-wave infrared thermal camera available for this study is not commonly used for thermal imagery acquisition over water, so we discuss accommodations that were made to account for potential interferences and describe considerations for future users interested in using the technology. We quantified thermal metrics from imagery and continuous loggers (e.g., mean, coefficient of variation, range) and compared those to hydrogeomorphic variability across aquatic areas using a Geographic Information System (GIS) dataset. Our findings showed that both temporal and spatial temperature patterns were linked to variation in depth and connectivity of aquatic areas across the pool. Despite some of the technical challenges associated with acquiring this imagery, the method shows promise for characterizing spatial variation in surface temperatures in the UMRS associated with complex physical features such as habitat rehabilitation and enhancement projects.

Iowa, Illinois, Minnesota, Wisconsin↗

Scale-specific metrics for adaptive generalization and geomorphic classification of stream features

The Richardson plot has been used to illustrate fractal dimension of naturally occurring landscape features that are sensitive to changes in scale or resolution, such as coastlines and river channels. The Richardson method estimates the length of a path by traversing (i.e., “walking”) the path with a specific stride length. Fractal dimension is determined as the slope of the Richardson plot, which shows path length over a range of stride lengths graphed on log-log axes. This paper describes a variant of the Richardson plot referred to as the Scale-Specific Sinuosity (S 3 ) plot. S 3 is defined as negative one times the slope of the Richardson plot for a given stride length. A plot of S 3 against stride length offers a frequency distribution whose area under the curve reflects total sinuosity, and whose points mark the amount of sinuosity contributed to the total sinuosity at each stride length. Mathematical relations of S 3 with fractal dimension and sinuosity for linear features are described. The S 3 metric is demonstrated and discussed for several linear stream features distributed over the conterminous United States. The S 3 metric can help guide the preservation of stream feature sinuosity during cartographic generalization and may assist automated geomorphic classification of river systems.

Conference Paper↗

Assessing the state of water resource knowledge and tools for future planning in the lower Rio Grande-Rio Bravo Basin

The Rio Grande/Rio Bravo Basin (hereinafter referred to as the Rio Grande) is a transboundary basin, with the Rio Grande forming the border between the United States and Mexico for approximately 2,034 km. The waters of the Rio Grande serve as a critical drinking source for 13 million people, connecting numerous population centers representing diverse backgrounds and cultures along its length. Cross-border ecosystems and communities make water management strategies particularly challenging. With different regulations and societal interests in the two countries, developing effective water-management strategies is challenging and requires the coordination of diverse interested parties representing different government agencies, institutions, and stakeholder groups with varying and sometimes conflicting objectives. To better evaluate the human and environmental water needs (environmental flows) of this constrained river system, an improved understanding of past and present water management objectives, policies, allocation practices, and water use is needed.

Lower Rio Grande-Río Bravo Basin↗

Coastal observations of alligator snapping turtles in the Florida Panhandle

The genus Macrochelys (alligator snapping turtles) inhabits river systems that empty into the Gulf of Mexico from Florida to Texas and contains the largest freshwater turtles in North America (Pritchard 2006). This paper details observations of Macrochelys on the coast and barrier islands in Franklin and Gulf counties, Florida, that contribute to our knowledge of their movements in brackish or saltwater ecosystems and occurrence on islands. Thomas et al. (2014) described turtles from this geographical area as a new species, Apalachicola alligator snapping turtle ( M. apalachicolae ), but Folt and Guyer (2015) disputed this designation, and many authorities still consider them to be alligator snapping turtles ( M. temminckii ; e.g., Crother 2017, Turtle Taxonomy Working Group 2017).

Florida Field Naturalist↗

Life zone investigations in Wyoming

Wyoming is among the foremost of our States in its wealth of natural scenery, culminating in the grandeur of Yellowstone National Park, one of the wonders of the world. In addition to this distinction it posseses vast open plains and lofty mountains whence flow the headwaters of mighty river systems emptying far away to the west into the Pacific Ocean, to the southeast into the Gulf of Mexico, and to the southwest into the Gulf of California. The various slope exposures of its mountain ranges, the fertility of its intervening valleys or basins, and the aridity of its desert spaces present a study of geographic and vertical distribution of wild life that is in many particulars unique. The study of geographic and vertical distribution of life with the governing factors and attendant problems is valuable as a matter of scientific research and in the attainment of practical knowledge. The Biological Survey has been making detailed investigations of the transcontinental life belts, or zones, of North America for some years, and this work has been carried on with special reference to their practical value. It has become increasingly evident that life zones furnish a fairly accurate index to average climatic conditions and, therefore, are useful as marking the limits of agricultural possibilities, so far as these are dependent upon climate. The knowledge thus gained has been published and made available as the investigations have progressed and the life zones have been mapped. 1

North American Fauna↗

Groundwater-flow model for the Wood River Valley aquifer system, south-central Idaho

A three-dimensional numerical model of groundwater flow was developed for the Wood River Valley (WRV) aquifer system, Idaho, to evaluate groundwater and surface-water availability at the regional scale. This mountain valley is located in Blaine County and has a drainage area of about 2,300 square kilometers (888 square miles). The model described in this report can serve as a tool for water-rights administration and water-resource management and planning. The model was completed with support from the Idaho Department of Water Resources, and is part of an ongoing U.S. Geological Survey effort to characterize the groundwater resources of the WRV. A highly reproducible approach was taken for constructing the WRV groundwater-flow model. The collection of datasets, source code, and processing instructions used to construct and analyze the model was distributed as an R statistical-computing and graphics package. Flow in the WRV aquifer was simulated using the MODFLOW-USG groundwater flow model. The transient flow model simulates groundwater flow between 1995 and 2010. The model uses a 100-meter (328-feet) uniform grid spacing with 54,922 active model cells distributed over three model layers. A confining unit in the south-central part of the Bellevue fan necessitated the use of a multi-layer model. Specified-flow boundaries were used to simulate the groundwater inflows from each of the major tributary basins (also known as tributary basin underflow) and the areal recharge of precipitation and applied irrigation. Head‑dependent flow boundaries were used to simulate the stream-aquifer flow exchange in river reaches and the groundwater discharge at the outlet boundaries of Stanton Crossing and Silver Creek. The model was calibrated by adjusting aquifer hydraulic properties to match simulated and measured water levels and stream-aquifer flow exchange, using the parameter-estimation program PEST. The model reasonably simulated the measured water-table elevation, orientation, and gradients. Stream-aquifer flow exchange along river reaches also was reasonably simulated by the model. Inflow into the WRV aquifer system originates from three sources (from largest to smallest): Streamflow loss from the Big Wood River and Silver Creek, Areal recharge of precipitation and applied irrigation, and Tributary basin underflow. Outflow from the WRV aquifer system originates from five sources (from largest to smallest): Aquifer discharge into the Big Wood River and Silver Creek, Production-well pumping, Discharge across the water table into the vadose zone, Subsurface outflow beneath Silver Creek near Picabo, and Subsurface outflow beneath the Big Wood River near Stanton Crossing. Temporal changes in aquifer storage are most affected by areal recharge and groundwater pumping, and also contribute to changes in streamflow gains.

Idaho↗

Mathematical model of the Tesuque aquifer system underlying Pojoaque River basin and vicinity, New Mexico

A three-dimensional digital model of ground-water flow was constructed to represent the dipping anisotropic beds of the Tesuque aquifer system underlying the Pojoaque River basin and vicinity, New Mexico. Simulations of steady-state conditions and historical ground-water withdrawals were consistent with observed data. The model was used to simulate the response of the aquifer system to an irrigation development plan in the Pojoaque River basin. Storage is the main source of water as 34.05 cubic feet per second (86 percent of the withdrawal rate) was simulated to be withdrawn from storage after 50 years of withdrawals for irrigation development; the maximum simulated water-level decline was 334 feet; and net simulated streamflow capture from the Rio Grande, Santa Cruz, Pojoaque, and Santa Fe Rivers was 5.63 cubic feet per second (14 percent of the withdrawal rate). The sensitivity of the model was tested by varying aquifer characteristics to the limits of the plausible range. Change in hydraulic head in Pojoaque River basin is most sensitive to hydraulic conductivity. In all simulations, after 50 years of withdrawals, the maximum simulated decline in hydraulic head ranged between 210 and 474 feet, storage in the aquifer system was the source of 80 to 90 percent of the water withdrawn from wells, and streamflow capture from the Rio Grande and its tributaries plus irrigation diversions from the tributaries of the Pojoaque River simulated a decrease in the flow of the Rio Grande of between 17.13 and 21.11 cubic feet per second.

New Mexico↗

Detailed measured sections, cross sections, and paleogeographic reconstructions of the upper cretaceous and lower tertiary nonmarine interval, Wind River Basin, Wyoming: Chapter 10 in Petroleum systems and geologic assessment of oil and gas resources in the Wind River Basin Province, Wyoming

Detailed measured sections and regional stratigraphic cross sections are used to reconstruct facies maps and interpret paleogeographic settings for the interval from the base of Upper Cretaceous Mesaverde Formation to top of lower member of the Paleocene Fort Union Formation in the Wind River Basin, Wyoming. The Mesaverde Formation spans the time during which the Upper Cretaceous seaway retreated eastward out of central Wyoming in Campanian time and the initial stages of the Lewis transgression in earliest Maastrichtian time. This retreat stalled for a considerable period of time during deposition of the lower part of the Mesaverde, creating a thick buildup of marginal marine sandstones and coaly coastal plain deposits across the western part of the basin. The Lewis sea transgressed into the northeast part of Wind River Basin, beginning in early Maastrichtian time during deposition of the Teapot Sandstone Member of the Mesaverde Formation. The Meeteetse Formation, which overlies the Teapot, was deposited in a poorly-drained coastal plain setting southwest of the Lewis seaway. The Lewis seaway, at maximum transgression, covered much of the northeast half of the Wind River Basin area but was clearly deflected around the present site of the Wind River Range, southwest of the basin, providing the first direct evidence of Laramide uplift on that range. Uplift of the Wind River Range continued during deposition of the overlying Maastrichtian Lance Formation. The Granite Mountains south of the basin also became a positive feature during this time. A rapidly subsiding trough during the Maastrichtian time formed near the presentday trough of the Wind River Basin in which more than 6,000 feet of Lance was deposited. The development of this trough appears to have begun before the adjacent Owl Creek Mountains to the north started to rise; however, a muddy facies in the upper part of Lance in the deep subsurface, just to the south, might be interpreted to indicate that the Cretaceous Cody Shale was being eroded off a rising Owl Creek Mountains in latest Cretaceous time. The Paleocene Fort Union Formation unconformably overlies older units but with only slight angular discordance around much of the margins of the Wind River Basin. Pre-Fort Union erosion was most pronounced toward the Wind River Range to the southwest, where the Fort Union ultimately overlies strata as old as the upper part of the Cretaceous Cody Shale. The unconformity appears to die out toward the basin center. Coal-forming mires developed throughout the western part of the basin near the beginning of the Paleocene. River systems entering the basin from the Wind River Range to the southwest and the Granite Mountains to the south produced areas of sandy fluvial deposition along mountain fronts. A major river system appears to have entered the basin from about the same spot along the Wind River Range throughout much of the Paleocene, probably because it became incised and could not migrate laterally. The muddy floodplain facies that developed along the deep basin trough during latest Cretaceous time, expanded during the early part of the Paleocene. Coal-forming mires that characterize part of the lower Fort Union Formation reached maximum extent near the beginning of the late Paleocene and just prior to the initial transgression of Lake Waltman. From the time of initial flooding, Lake Waltman expanded rapidly, drowning the coal-forming mires in the central part of the basin and spreading to near basin margins. Outcrop studies along the south margin of the basin document that once maximum transgression was reached, the lake was rapidly pushed basinward and replaced by fluvial environments.

Wyoming↗

Estimates of ground-water recharge to the Yakima River Basin aquifer system, Washington, for predevelopment and current land-use and land-cover conditions

Two models were used to estimate ground-water recharge to the Yakima River Basin aquifer system, Washington for predevelopment (estimate of natural conditions) and current (a multi-year, 1995-2004, composite) land-use and land-cover conditions. The models were the Precipitation-Runoff Modeling System (PRMS) and the Deep Percolation Model (DPM) that are contained in the U.S. Geological Survey's Modular Modeling System. Daily values of recharge were estimated for water years 1950-98 using previously developed PRMS-watershed models for four mainly forested upland areas, and for water years 1950-2003 using DPM applied to 17 semiarid to arid areas in the basin. The mean annual recharge under predevelopment conditions was estimated to be about 11.9 in. or 5,450 ft 3 /s (about 3.9 million acre-ft) for the 6,207 mi 2 in the modeled area. In the modeled areas, recharge ranged from 0.08 in. (1.2 ft 3 /s) to 34 in. (2,825 ft 3 /s). About 97 percent of the recharge occurred in the 3,667 mi 2 area included in the upland-area models, but much of this quantity is not available to recharge the bedrock hydrogeologic units. Only about 1.0 in., or 187 ft 3 /s (about 0.14 million acre-ft), was estimated to occur in the 2,540 mi 2 area included in the semiarid to arid lowland modeled areas. The mean annual recharge to the aquifer system under current conditions was estimated to be about 15.6 in., or 7,149 ft 3 /s (about 5.2 million acre-ft). The increase in recharge is due to the application of irrigation water to croplands. The annual quantity of irrigation was more than five times the annual precipitation for some of the modeled areas. Mean annual actual evapotranspiration was estimated to have increased from predevelopment conditions by more than 1,700 ft 3 /s (about 1.2 million acre-ft) due to irrigation.

Washington↗