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At least 865 records · Page 48Linked to original sources

Variation in age ratio of midcontinent greater white-fronted geese during fall migration

Annual productivity is an important parameter for the management of waterfowl populations. Fall age ratio (juveniles:total birds) is an index of productivity of the preceding breeding season. However, differences in the timing of migration between family groups and nonbreeding birds may bias age-ratio estimates. We examined temporal variation in age ratios of midcontinent greater white-fronted geese Anser albifrons frontalis from interior and northwestern Alaska at a northern autumn staging area near Delta Junction, Alaska. Photographic sampling conducted near Delta Junction resulted in an annual age ratio of 0.388 ± 0.004 (mean ± SE) in 2010 and 0.390 ± 0.001 in 2011. Our study demonstrated temporal variation in age ratios over the duration of the migration period during August and September. We recommend that sampling be conducted for 3-d periods at the beginning, middle, and end of the migration period to account for temporal variation in migration of family groups.

Journal of Fish and Wildlife Management↗

Grazing intensity effects on the breeding avifauna of North Dakota native grasslands

A breeding bird census and plant survey was conducted on 180 samples of lightly, moderately, and heavily grazed and hayed native grasslands in North Dakota in 1974. The ten most important cover plants on each of eight major physiographic landforms in three of the four regions (the Agassiz Lake Plain excluded) overlapped so extensively that only 19 species were involved: 13 grasses or sedges, four forbs, one shrub, and one clubmoss. Bird densities were generally highest in (i) regions and landforms containing numerous natural basin wetlands, (ii) flatter, glaciated landforms containing more fertile soils, and (iii) landforms of greater relief and high habitat heterogeneity. Avian species richness tended to decrease with increased grazing intensity, but total bird density increased due to higher populations of a few species, and hayland that had been mowed and raked during the previous growing season was highly attractive to some species.

Canadian Field-Naturalist↗

California condors

The California condor ( Gymnogyps californianus ) is a member of the vulture family. With a wingspan of about 3 m (9 ft) and weighing about 9 kg (20 lb), it spends much of its time in soaring flight visually seeking dead animals as food. The California condor has always been rare (Wilbur 1978; Pattee and Wilbur 1989). Although probably numbering in the thousands during the Pleistocene epoch in North America, its numbers likely declined dramatically with the extinction of most of North America's large mammals 10,000 years ago. Condors probably numbered in the hundreds and were nesting residents in British Columbia, Washington, Oregon, California, and Baja California around 1800. In 1939 the condor population was estimated at 60-100 birds, and its home range was reduced to the mountains and foothills of California, south of San Francisco and north of Los Angeles. Conservation to halt the condor's decline included establishing the Sisquoc (1937) and Sespe (1947) condor sanctuaries within the Los Padres National Forest, obtaining fully protected status under California Fish and Game Code (1953), placement on California's first state endangered species list (1971), and, finally, being listed by the federal government under the Endangered Species Act of 1973 (Wilbur 1978). The success of these efforts could not be judged, however, because verifiable status and trends data did not become available until 1982. By using these data, we confirmed the decline in condor numbers over the past 50 years was even greater than thought.

Book chapter↗

Changing land use: Problems and opportunities

Under the pressure of increasing human populations and expanding demands for food and fiber, native tropical and temperate habitats are becoming more restricted, and populations of many resident and migratory birds are declining. Mist net surveys of 111 forest and agricultural sites in Mexico, Belize, and Guatemala show that some migratory species use a wide variety of habitats during the non-breeding season; other migrants, especially ground-feeding insectivores that nest in temperate forests, are largely restricted to forest habitats during the northern winter. Most tropical residents are also scarce or absent in agricultural habitats; this is especially true of the suboscine families, which are an important component of tropical forests: Furnariidae, Dendrocolaptidae, Formicariidae, Tyrannidae, and Pipridae: Of the various agricultural habitats studied, arboreal crops, especially mature citrus and cacao, were used by a wide variety and relatively large number of migrants; at the other extreme, few birds were captured or observed in commercially grown allspice and platanos (bananas). Although habitat constraints on many species are increasing, the impact of these constraints can be reduced through research, management, legislation, and especially education. Long-range habitat management objectives that reduce forest fragmentation and promote retention of critical habitats for species can be realized if an informed and concerned public can be created

Book chapter↗

Invasive herbivory: resident Canada geese and the decline of wild rice along the tidal Patuxent River

While concern grows over the increasing numbers of exotic mute swans ( Cygnus olor ) on the Chesapeake Bay, less attention seems to be given to the highly familiar and native Canada goose ( Branta canadensis ) which has over time developed unprecedented nonmigratory, or resident, populations. Although nuisance flocks of Canada geese have been well advertised at city parks, athletic fields, and golf courses over the past three decades, recent expansion of populations to an estimated one million birds in the Atlantic Flyway, and to over 100,000 in Maryland, carries a threat of broader ecological consequences.

Maryland↗

Eastern Imperial Eagle Aquila heliaca

Eastern imperial eagles are a short-, medium-distance, partially-migratory, or even non-migratory, raptor that breeds at the forest-steppe interface in Eurasia and winters in Northern Africa, the Middle East or South Asia. Migratory strategies of imperial eagles are diverse. Eagles breeding in Central and Southeast Europe and south of the Black Sea usually are year-round residents or partial- or short- distance migrants that winter in the Balkan Peninsula, Northern Africa, or western parts of the Middle East. Eagles that summer to the east of the Black Sea are usually medium-distance migrants that winter in the Middle East or south Asia. Migration tends to follow topographic features, avoids water-crossings and, especially for young birds, may be intermittent and indirect. Populations of imperial eagles are small, in decline in some parts of the distribution, and the species faces a large number of threats including electrocution, persecution, and capture for sale in markets.

Book chapter↗

Framework for assessing and mitigating the impacts of offshore wind energy development on marine birds

Offshore wind energy development (OWED) is rapidly expanding globally and has the potential to contribute significantly to renewable energy portfolios. However, development of infrastructure in the marine environment presents risks to wildlife. Marine birds in particular have life history traits that amplify population impacts from displacement and collision with offshore wind infrastructure. Here, we present a broadly applicable framework to assess and mitigate the impacts of OWED on marine birds. We outline existing techniques to quantify impact via monitoring and modeling (e.g., collision risk models, population viability analysis), and present a robust mitigation framework to avoid, minimize, or compensate for OWED impacts. Our framework addresses impacts within the context of multiple stressors across multiple wind energy developments. We also present technological and methodological approaches that can improve impact estimation and mitigation. We highlight compensatory mitigation as a tool that can be incorporated into regulatory frameworks to mitigate impacts that cannot be avoided or minimized via siting decisions or alterations to OWED infrastructure or operation. Our framework is intended as a globally-relevant approach for assessing and mitigating OWED impacts on marine birds that may be adapted to existing regulatory frameworks in regions with existing or planned OWED.

Biological Conservation↗

A review of the ecology and conservation biology of Sali (Micronesian Starling, Aplonis opaca guami) on Guam

The accidental introduction of the Brown Treesnake (BTS, Boiga irregularis ) to Guam following World War II led to the extinction, extirpation, or severe decline of most of Guam’s native avifauna. One forest bird species that managed to persist is the cavity-nesting Såli (Micronesian Starling, Aplonis opaca guami ), a once-ubiquitous native omnivore whose current Guam distribution is restricted primarily to urbanized areas in the northern half of the island, with a second much smaller population on Cocos Island off southern Guam. Såli on Guam breed year-round and are habitat generalists with large home ranges. Besides arthropods and small vertebrates, they consume fruit from a wide range of native and introduced plant species and are key seed dispersers within the local ecosystem. Såli on Guam suffer extremely high post-fledging mortality due to predation by BTS and cats, leading to a population age structure strongly skewed towards adult birds. Other threats include habitat loss and degradation, extreme weather events, and sea level rise. Key management activities aimed at restoring this species include predator control, while non-native ungulate removal and vegetation management may be undertaken to support broader ecological recovery. This species account on Såli provides a review of the literature on this species, synthesizing information on habitat use, life history, and demographics to provide a biological foundation for future recovery efforts.

Micronesica↗

An assessment of arthropod prey resources at Nakula Natural Area Reserve, a potential site of reintroduction for Kiwikiu (Pseudonestor xanthophrys) and Maui `Alauahio (Parareomyza montana)

Hawaiian forest birds have declined dramatically since humans arrived in the archipelago. Birds from all foraging guilds have been affected but insectivorous species are currently at greatest risk of extinction. On the island of Maui, populations and ranges of the insectivorous kiwikiu (Maui parrotbill; Pseudonestor xanthophrys) and Maui ‘alauahio (Maui creeper; Paroreomyza montana) have declined significantly from historic levels primarily due to habitat loss, predation,disease, and food web disruption, leading to federal listings of endangered species and species of concern, respectively. Recovery plans for these birds include reestablishment of populations in parts of their former range. Nakula Natural Area Reserve on the leeward side of HaleakalāVolcano has been targeted for release of wild-caught or captive-bred individuals. The mesic, montane koa-‘ōhi‘a (Acacia koa-Metrosideros polymorpha) forest at Nakula has been heavily impacted through grazing by feral ungulates, but recent management actions to exclude these animals are promoting forest recovery. The objective of this study was to assess the arthropod prey base at Nakula in preparation for reintroductions of kiwikiu and Maui ‘alauahio. To accomplish that goal, we compared arthropod abundances at Nakula to those at Hanawi Natural Area Reserve and Waikamoi Preserve, areas where kiwikiu and Maui ‘alauahio are currently found. We also identified diets of kiwikiu and Maui ‘alauahio from fecal samples to better understand and evaluate the prey base at Nakula. Assessment methods included clipping branch tips to sample arthropods within the foliage of koa and ‘ōhi‘a, using traps to quantify arthropods on koa and ‘ōhi‘a bark surfaces, counting exit holes to quantify abundances of beetles (Coleoptera) within dead branches of koa, and measuring the density of arthropods within the stems of ‘ākala (Rubus hawaiiensis). The diet of kiwikiu was dominated by caterpillars (Lepidoptera larvae), which comprised 90% of all prey items for 50 adult birds and 98% of all prey for two nestlings. Caterpillars were also the most important prey for Maui ‘alauahio (43% for 104 adult birds) although spiders (Araneae, 16%), beetles (12%) and true bugs, planthoppers and psyllids (Hemiptera; 12%) were also important. Caterpillars were generally the most abundant type of arthropod in the foliage of koa and ‘ōhi‘a, although spiders, beetles and hemipterans were also common. Total arthropod biomass and caterpillar biomass at Nakula was as great, or greater, than that observed at Hanawi and Waikamoi per unit of foliage of both koa and ‘ōhi‘a. Spiders generally dominated the bark fauna on both koa and ‘ōhi‘a at all sites although isopods (Isopoda), millipedes (Myriapoda: Millipeda) and lacewings (Neuroptera) were also abundant at Waikamoi and Hanawi. Total arthropod biomass on bark, as well as the biomass of several individual taxa, was significantly lower at Nakula than the other sites. Our measurement of the density of beetle exit holes in dead koa branches found no difference between Nakula and Waikamoi. Finally, no difference existed in the abundance of arthropods (primarily caterpillars and moth pupae) within ‘ākala stems among sites. With the exception of bark surfaces, our results suggest that the arthropod prey base for birds on primary foraging substrates at Nakula is similar to that found at two sites within the current range of kiwikiu and Maui ‘alauahio. However, our results should be viewed with caution because they are limited to the scale of individual branch, tree, or ‘ākala stem. To complete the assessment, our results should be scaled up to the landscape level by determining the density of each substrate within each site. Key arthropod prey of kiwikiu and Maui ‘alauahio are available at Nakula and, as habitat restoration continues, food abundance should increase to the point at which populations of these birds can be supported.

Hawaii↗

Demographics, diet, movements, and survival of an isolated, unmanaged raccoon Procyon lotor (Procyonidae, Carnivora) population on the Outer Banks of North Carolina

Raccoons ( Procyon lotor ) are highly adaptable meso-carnivores that inhabit many environments, including the Atlantic barrier islands, where their role as predators of declining, beach-nesting bird and turtle species is of particular interest. Population models that improve our understanding of predator-prey dynamics are receiving increasing attention in the literature; however, their effective application requires site-specific information on population parameters. We studied an unharvested raccoon population on the Outer Banks of North Carolina and evaluated spatial and seasonal differences in a number of population/demographic factors of raccoons inhabiting areas of high and low human activity. Raccoons denned and foraged primarily in salt marsh habitats but shifted their movements in response to changes in seasonal resource conditions. The population was skewed toward older animals and exhibited delayed breeding, typical of populations at high density with few sources of mortality. Diet and movement analysis indicated shorebird and turtle predation was attributed to a small number of individual raccoons. Although seasonal resources appeared adequate to sustain a high population density of raccoons, poor body condition and low recruitment suggested a population near carrying capacity.

North Carolina↗

Adaptive harvest management for the Svalbard population of pink-footed geese: 2018 progress summary

This report describes progress on development and implementation of an adaptive harvest management program for maintaining the Svalbard population of Pink-footed Geese (Anser brachyrhynchus) near their target level (60,000) by providing sustainable harvests in Norway and Denmark. Specifically, this report provides an assessment of recent monitoring information and its implications for hunting seasons in 2018. An Adaptive Harvest Management (AHM) program requires specification of four elements: (a) A set of alternative population models, which bound the uncertainty about population dynamics; (b) A set of weights describing the relative credibility of the alternative models, which are updated each year based on a comparison of model predictions and monitoring information; (c) A set of alternative harvest quotas from which to choose; and (d) An objective function, by which alternative harvest strategies are evaluated and an optimal strategy chosen. The most current set of monitoring information was used to update model weights for the 1991-2017 period. Current model weights suggest little evidence for density-dependent survival and reproduction. These results suggest that the Pink-footed Goose population may have experienced a release from density-dependent mechanisms, corresponding to the period of rapid growth in population size. There is equivocal evidence for the effect of the number of days above freezing in May in Svalbard on survival, but the evidence for an effect on reproduction has been increasing in recent years. Since the 2016 hunting season, harvest quotas are prescribed on an annual basis rather than every three years because of the potential to better meet management objectives. Based on updated model probabilities, the November 2017 population size (72,000), the proportion of the population comprised of one-year-old birds (0.076), and days above freezing in Svalbard in May 2018 (27), the optimal harvest quota for the 2018 hunting season is approximately 27,000. With the agreed upon harvest allocation of 30% Norway and 70% Denmark, the national quotas are 8,100 and 18,900, respectively, which are higher than the harvests realized in previous years. In 2017 the quota for the two countries combined was 36,000, but only a harvest of about 11,400 was realized. The decrease in harvest quota for 2018 is largely attributable to the apparent decline in population size. We also describe the ongoing development of an Integrated Population Model (IPM), which uses all available demographic data for Pink-footed Geese in a single, unified analysis. IPM estimates of harvest rates of adult geese were variable and relatively low prior to the implementation of AHM (2013), and have been relatively high since. The increase in harvest rates has been accompanied by a decline in annual survival. The ratio of young-of-the-year to older birds just prior to the hunting season has been variable over time, and since about 2005 has been highly correlated with the number of days above freezing in May in Svalbard. IPM estimates of population size suggest that abundance of Pink-footed Geese has been relatively stable, or declining slightly, in recent years. Based on the IPM estimate of population size in November 2017 of 68,800 (95% credible interval: 58,200 – 79,400), the optimal harvest quota for the 2018 hunting season is 15,000. This is lower than that derived from the set of nine discrete models because the IPM estimate of November population size is lower than the November count, and because the IPM model does not consider May temperatures in Svalbard, but rather assumes reproductive success varies randomly about the mean.

Report↗

A critical look at national monitoring programs for birds and other wildlife species

Concerns?about declines in numerous taxa have created agreat deal of interest in survey development. Because birds have traditionally been monitored by a variety of methods, bird surveys form natural models for development of surveys for other taxa. Here I suggest that most bird surveys are not appropriate models for survey design. Most lack important design components associated with estimation of population parameters at sample sites or with sampling over space, leading to estimates that may be biased, I discuss the limitations of national bird monitoring programs designed to monitor population size. Although these surveys are often analyzed, careful consideration must be given to factors that may bias estimates but that cannot be evaluated within the survey. Bird surveys with appropriate designs have generally been developed as part of management programs that have specific information needs. Experiences gained from bird surveys provide important information for development of surveys for other taxa, and statistical developments in estimation of population sizes from counts provide new approaches to overcoming the limitations evident in many bird surveys. Design of surveys is a collaborative effort, requiring input from biologists, statisticians, and the managers who will use the information from the surveys.

Book chapter↗

Genetic structure, diversity, and interisland dispersal in the endangered Mariana Common Moorhen (Gallinula chloropus guami)

The Mariana Common Moorhen ( Gallinula chloropus guami ) is a highly endangered taxon, with fewer than 300 individuals estimated to occur in the wild. The subspecies is believed to have undergone population declines attributable to loss of wetland habitats on its native islands in the Mariana Islands. We analyzed mitochondrial DNA (mtDNA) sequences (control region and ND2 genes) and nuclear microsatellite loci in Mariana Common Moorhens from Guam and Saipan, the two most distal islands inhabited by the subspecies. Our analyses revealed similar nuclear genetic diversity and effective population size estimates on Saipan and Guam. Birds from Guam and Saipan were genetically differentiated (microsatellites: F ST = 0.152; control region: F ST = 0.736; ND2: F ST = 0.390); however, assignment tests revealed the presence of first-generation dispersers from Guam onto Saipan (1 of 27 sampled birds) and from Saipan onto Guam (2 of 28 sampled birds), suggesting the capability for long-distance interpopulation movements within the subspecies. The observed dispersal rate was consistent with long-term estimates of effective numbers of migrants per generation between islands, indicating that movement between islands has been an ongoing process in this system. Despite known population declines, bottleneck tests revealed no signature of historical bottleneck events, suggesting that the magnitude of past population declines may have been comparatively small relative to the severity of declines that can be detected using genetic data.

The Condor↗

The future demographic niche of a declining grassland bird fails to shift poleward in response to climate change

Context Temperate grasslands and their dependent species are exposed to high variability in weather and climate due to the lack of natural buffers such as forests. Grassland birds are particularly vulnerable to this variability, yet have failed to shift poleward in response to recent climate change like other bird species in North America. However, there have been few studies examining the effect of weather on grassland bird demography and consequent influence of climate change on population persistence and distributional shifts. Objectives The goal of this study was to estimate the vulnerability of Henslow’s Sparrow ( Ammodramus henslowii ), an obligate grassland bird that has been declining throughout much of its range, to past and future climatic variability. Methods We conducted a demographic meta-analysis from published studies and quantified the relationship between nest success rates and variability in breeding season climate. We projected the climate-demography relationships spatially, throughout the breeding range, and temporally, from 1981 to 2050. These projections were used to evaluate population dynamics by implementing a spatially explicit population model. Results We uncovered a climate-demography linkage for Henslow’s Sparrow with summer precipitation, and to a lesser degree, temperature positively affecting nest success. We found that future climatic conditions—primarily changes in precipitation—will likely contribute to reduced population persistence and a southwestward range contraction. Conclusions Future distributional shifts in response to climate change may not always be poleward and assessing projected changes in precipitation is critical for grassland bird conservation and climate change adaptation.

Landscape Ecology↗

Towards sustainable management of huntable migratory waterbirds in Europe

The EU Birds Directive and the African-Eurasian Waterbird Agreement provide an adequate legal framework for sustainable management of migratory waterbird populations. The main shortcoming of both instruments is that it leaves harvest decisions of a shared resource to individual Member States and Contracting Parties without providing a shared information base and mechanism to assess the impact of harvest and coordinate actions in relation to mutually agreed objectives. A recent update of the conservation status of waterbirds in the EU shows that almost half of the populations of species listed on Annex II of the Birds Directive have a declining short-term trend and over half of them are listed in Columns A and B of AEWA. This implies that their hunting could either only continue under the framework of an adaptive harvest management plan or their hunting should be regulated with the view of restoring them in favourable conservation status. We argue that a structured approach to decision-making (such as adaptive management) is needed, supported with adequate organisational structures at flyway scale. We review the experience with such an approach in North America and assess the applicability of a similar approach in the European context. We show there is no technical reason why adaptive harvest management could be not applied in the EU or even AEWA context. We demonstrate that an informed approach to setting allowable harvests does not require detailed demographic information. Essential to the process, however, are estimates of either the observed growth rate from a monitoring program or the growth rate expected under ideal conditions. In addition, periodic estimates of population size are needed, as well as either empirical information or reasonable assumptions about the form of density dependence. We show that such information exists for many populations, but improvements are needed to improve geographic coverage, reliability and timely data availability. We highlight the importance of the International Waterbird Census and specialised goose and seaduck monitoring in estimating population sizes and observed growth rate of the populations. We encourage further investments into the development of these schemes. We also recognise the importance of migration studies to improve our understanding of delineations of populations. We also highlight that, with a few exceptions, the available data does not allow the European Commission, competent authorities of the Members States or other AEWA Contracting Parties to assess levels of harvest and their sustainability and, therefore, regulate hunting accordingly. Therefore, we recommend that annual reporting on 2 harvest levels of waterbird populations would be gradually introduced in the EU and the AEWA region. We propose that future AEWA and EU action plans and management plans for Annex II species should apply the principles of adaptive harvest management framework and make provisions for setting up adequate monitoring and information management systems and organisational structures to manage the decision-making process. We suggest that internationally coordinated management structures are established to facilitate dialogue, learning and communication between stakeholders with different interests and cultural backgrounds.

Report↗

Adaptive harvest management for the Svalbard population of pink-footed geese (Anser brachyrhynchus)

This document describes progress to date on the development of an adaptive harvest-management strategy for maintaining the Svalbard population of pink-footed geese (Anser brachyrhynchus) near their target level (60,000) by providing for sustainable harvests in Norway and Denmark. Specifically, this report provides an assessment of the most recent monitoring information and its implications for the harvest management strategy. The development of an adaptive harvest management (AHM) strategy requires specification of four elements: (a) a set of alternative population models, which bound the uncertainty about effects of harvest and other relevant environmental factors; (b) a set of probabilities (or weights) describing the relative credibility of the alternative models, which are updated each year based on a comparison of model predictions and monitoring information; (c) a set of alternative harvest quotas from which to choose; and (d) a management objective function, by which alternative harvest strategies can be evaluated and a mathematically optimal strategy identified. By combining varying hypotheses about survival and reproduction, a suite of nine models were developed. Those models represent a wide range of possibilities concerning the extent to which demographic rates are density dependent, and the extent to which spring temperatures influence survival and reproduction. Five of the models incorporate density-dependent mechanisms that would maintain the population near a carrying capacity (i.e., in the absence of harvest) of 65,000 – 129,000 depending on the specific model. The remaining four models are density independent and predict an exponentially growing population even with moderate levels of harvest. The most current set of monitoring information was used to update model weights for the period 1991 – 2016. Current model weights suggest little evidence for density-dependent survival and reproduction. These results suggest that the pink-footed goose population may have recently experienced a release from density-dependent mechanisms, corresponding to the period of most rapid growth in population size. There is equivocal evidence for the effect of May temperature days in Svalbard (number of days with temperatures above freezing) on survival and reproduction. Beginning with the 2016 hunting season, harvest quotas are chosen on an annual basis rather than every three years because of the potential to better meet management objectives. The optimal harvest strategy, however, remains “knife-edged,” meaning that small changes in resource status can precipitate large changes in the annual harvest quota. This potential outcome is likely to be of concern to hunters, and we are investigating ways in which large swings in harvest quotas might be dampened. Based on updated model probabilities, the recent observations of record-high population size (88,000), the aboveaverage proportion of the population comprised of one-year-old birds (0.196), and temperature days in Svalbard (4), the suggested harvest quota for the 2017 hunting season is 36,000. Last year the quota was 25,000, yet a harvest of only 16,143 was realized. We are increasingly concerned that with the return of average spring temperatures in Svalbard, the population will continue to grow beyond managers’ ability to control it, as is the case with many goose populations in Europe and North America.

EGMP Technical Report↗

Distribution of non-breeding great lakes piping plovers (Charadrius melodus) along Atlantic and Gulf of Mexico coastlines: Ten years of band sightings

In 1993, a mark-recapture effort was initiated to band annually all Great Lakes Piping Plover nesting adults and offspring. With voluntary reporting by observers, >430 sightings of 154 individually-marked Great Lakes banded birds were documented on the wintering grounds during 19952005. This paper reports non-breeding distribution and site-fidelity and identifies Critical Habitat units used by this population during the winter. Information obtained through banded bird sightings indicates that the winter range of Great Lakes Piping Plovers extends from North Carolina to Texas, and the Bahamas, with the majority (75%) of reported individuals wintering in Georgia and Florida. About 95% of sightings were near or within federally-designated winter Critical Habitat for Piping Plovers. Within season (52%) and between-year (62%) site fidelity was documented for resightings within 3.5 km of initial sighting. Although breeding pairs do not winter in close association, there is some evidence to suggest that offspring winter closer to the male rather than the female parent (P-value = 0.03), and adult males and females appear to exhibit latitudinal segregation (P-value < 0.001). Females reach the winter grounds before males, arriving in July and staying through April (???9 months) or 75% of the annual cycle. The study is the first to identify winter distribution for the Great Lakes Piping Plover population. The significant proportion of the annual cycle spent on the wintering grounds emphasizes the importance of habitat protection during the non-breeding season for this federally-listed population.

Waterbirds↗