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Forage and habitat for pollinators in the northern Great Plains—Implications for U.S. Department of Agriculture conservation programs

Managed and wild pollinators are critical components of agricultural and natural systems. Despite the well-known value of insect pollinators to U.S. agriculture, Apis mellifera (Linnaeus, 1758; honey bees) and wild bees currently face numerous stressors that have resulted in declining health. These declines have engendered support for pollinator conservation efforts across all levels of government, private businesses, and nongovernmental organizations. In 2014, the U.S. Department of Agriculture (USDA) and the U.S. Geological Survey initiated an interagency agreement to evaluate honey bee forage across multiple States in the northern Great Plains and upper Midwest. The long-term goal of this study was to provide an empirical evaluation of floral resources used by honey bees, and the relative contribution of multiple land covers and USDA conservation programs to bee health and productivity. Our multi-State analysis of land-use change from 2006 to 2016 revealed loss of grassland and increases in corn and soybean area in North and South Dakota, representing a significant loss of bee-friendly land covers in areas that support the highest density of summer bee yards in the entire United States. Our landscape models demonstrate the importance of the Conservation Reserve Program in providing safe locations for beekeepers to keep honey bees during the summer and highlights how land use in the northern Great Plains has a lasting effect on the health of honey bee colonies during almond pollination the subsequent spring. Our multiseason, multi-State genetic analysis of honey bee-collected pollen revealed Melilotus spp., Asteraceae, Trifolium spp., Fabaceae, Sonchus arvensis , Symphyotrichum cordifolium , and Solidago spp. were the top taxa detected; Melilotus spp. represented 42 percent of all detected taxa. Symphyotrichum cordifolium , Solidago spp., and Grindelia spp. were the top native forbs detected in honey bee-collected pollen. We also conducted plant and bee surveys on private lands enrolled in the Conservation Reserve Program and Environmental Quality Incentives Program. In general, we found significant variability in floral resources and pollinator utilization across USDA programs and practices. On average, greater than 75 percent of honey bee flower observations on private lands enrolled in a USDA conservation program were on non-native forbs, whereas 33 percent of wild bee flower observations were on non-native forbs. Melilotus officinalis and Medicago sativa were the most visited by honey bees, wherease Medicago sativa and Helianthus maximiliani were the most visited by wild bees. Our analysis of nectar dearth periods in June and September for honey bees revealed that although Melilotus officinalis and Medicago sativa were highly visited, less common native forb species such as Ratibida columnifera , Agastache foeniculum , and Gaillardia aristata were preferred species. However, these preferred species were relatively rare on the landscape and are, therefore, unlikely to make up a sizable part of the honey bee diet. In addition to our empirical results, we also showcase how the U.S. Geological Survey Pollinator Library, a decision-support tool for natural resource managers, can be used to design cost-effective seeding mixes for pollinators. Collectively, the results of this research will assist USDA with maximizing the ecological impact and cost-effectiveness of their conservation programs on pollinators in the northern Great Plains.

Minnesota, North Dakota, South Dakota↗

Potential effects of climate change on Appalachian stoneflies ( Remenus kirchneri , Acroneuria kosztarabi , and Tallaperla lobata )

Plecoptera (stoneflies) are an order of insects where most species rely on clean, fast-moving freshwater for an aquatic larval stage followed by a short terrestrial adult stage. Most species of Plecoptera seem to be restricted to specific stream types and thermal regimes. Climate-driven changes are likely to alter stream temperatures and flow, resulting in physiological stress, reduced reproductive success, and possibly latitudinal or elevational distribution shifts. This report focuses on climate projections and the resulting ecological effect for three species of Appalachian stoneflies: Remenus kirchneri , Acroneuria kosztarabi , and Tallaperla lobata . Although species-specific information is sparse for these three species, climate studies for other Plecoptera spp. are applicable. In the focal region, temperature is increasing and likely leading to increased stream temperatures. In response, Plecoptera spp. will likely experience physiological stress from increasing metabolic rates and energy demands concurrent with changing food quality and access. Warming temperatures and decreased larval energy stores are likely to contribute to lower adult body size and longevity, thus decreasing reproductive success. Whereas projected changes to precipitation and runoff are less certain, under drier future climate projections, decreased streamflow may further stress larval Plecoptera. Remenus kirchneri , A. kosztarabi , and T. lobata will likely retain stable permanent stream habitats for the analyzed future (2006–99). Changing climate is of particular concern for mountaintop species R. kirchneri and T. lobata because they may be unable to track shifts in suitable climate and habitat.

North Carolina, Tennessee, Virginia↗

Guide for benthic invertebrate studies in support of Natural Resource Damage Assessment and Restoration

This guide is intended to assist with characterizing injury to freshwater benthic macroinvertebrates (BMIs) in Natural Resource Damage Assessment and Restoration (NRDAR) cases. The contents are narrowly focused on insects, crustaceans, snails, and other invertebrate fauna that are typically considered part of BMI communities and are not intended to address studies of injury to larger benthic taxa such as freshwater mussels, crayfish, or benthic fishes or amphibians. Although some percentage of the community functions as predators, BMIs are predominantly primary consumers (for example, scrapers, shredders, and filterer/gatherer feeding groups) that play an essential role in converting carbon and nitrogen from plant tissues into animal biomass for higher-order consumers, especially in flowing waters. Aquatic contaminants can disrupt the quantity and quality of energy transferred (ecosystem function) by reducing invertebrate biomass and diversity. Additionally, the accumulation of toxic residues in invertebrate tissues may be a source of exposure leading to adverse effects in higher trophic levels. The goal of NRDAR BMI assessments is to establish direct linkages of contaminant exposure to injuries reflected by changes in community structure (for example, reduced density and taxa richness) or by effects at the individual population level (for example, survival, growth, and reproduction). BMIs are infrequently the U.S. Department of Interior (DOI)-managed resource in a NRDAR case, with managed resources more frequently including migratory birds, fish, or other insectivorous vertebrates. Therefore, it is critical to have clearly defined objectives for evaluating BMIs and an understanding of how invertebrate data relate to the quantification of injuries to the DOI-managed resource. This guide is intended to assist decisions on whether or not to proceed with BMI studies, use of existing information and data for screening purposes, and what types of studies can support a BMI-injury determination. This document is intended to provide general considerations and best practices for assessing BMIs. Relevant guidance and references are listed throughout the report as sources for specific methods and analysis.

Open-File Report↗

Non-negligible near-term risk of extinction to the eastern migratory population of monarch butterflies—An updated assessment (2006–22)

The eastern migratory population of monarch butterflies ( Danaus plexippus ) started declining as early as the mid-1970s and seemed to stop declining by the early 2000s; the population now (about 2022) persists at a much-reduced abundance. Stochastic variation in abundance, at levels typical of monarch butterflies and other insects, was assessed to determine whether this population is at heightened risk of quasi-extinction, a level of abundance below which recovery of the migratory behavior is uncertain. Using previously published Bayesian state-space modeling methods it was determined roughly equivalent risk of quasi-extinction as was reported in 2016 for the species (28.7 percent [1.9–81.0 credible interval] and 52.0 percent [3.2–97.7 credible interval] at the 10- and 20-year marks, respectively). Though highly uncertain, the risk is non-negligibly positive. Warning signal analysis indicates the current dynamic is dominated by stochastic variation, which seems to be heightening risk with the passage of time. Increasing breeding opportunities through restoration of milkweed in its northern breeding locations seems to be the most promising means of mitigating extinction risk for this species.

Open-File Report↗

Hydrobiological characteristics of Shark River estuary, Everglades National Park, Florida

Water quality in the Shark River estuary was strongly influenced by seasonal patterns of rainfall, water level and temperature. During the rainy season (summer and early fall) the salinity in the 20-mile long estuary ranged from that of fresh water to half that of sea water while concentrations of dissolved oxygen were low, 2-5 milligrams per liter (mg/l) presumably because, among other factors, microbial activity and respiration were accelerated by high temperatures (30-33 degrees C). During the dry season (late fall through spring) the salinity ranged from 18 grams per liter (g/l) in the headwaters to 36 g/l at the Gulf during a dry year such as 1967 and from 1 to 25 g/l during a wet year such as 1969. Concentrations of dissolved oxygen increased from 2-3 mg/l in the summer of 1967 to 4-7 mg/l in the winter of 1968, and temperature decreased from an average of about 30 degrees C in summer to 20 degrees C in winter. Water level declined 5 to 10 decimeters in the headwaters during the dry season, and salinity and tidal action increased. Large amounts of submerged vegetation died in some headwater creeks at the end of the dry season, presumably killed by salinities above 3 g/l. The decaying organic matter and the decrease in photosynthesis resulted in low dissolved oxygen (1-2 mg/l). Fish died at this time probably as a result of the low dissolved oxygen. Trace elements, heavy metals and insecticides occurred in the waters of the estuary in concentrations below those indicated as harmful for aquatic life by current standards established by the Federal Water Pollution Control Administration (1968). The insecticides detected were concentrated in sediment and in various organisms. The patterns of distribution of planktonic and small nektonic animals in the estuary were related to salinity. Copepods (Arcatia tonsa, Labidocera aestiva, Pseudodiaptomus coronatus), cumaceans (Cyclaspis sp.), chaetognaths (Sagitta hispida), bay anchovies (Anchoa mitchilli), and scaled sardines (Harengula pensacolae) were the dominant animals collected in the higher salinities (10-25 g/l) near the mouth of the estuary. Amphipods (Corophium sp. and Grandidierella sp.), mysids (Mysidopsis almyra and Gastrosaccus dissimilis), crab larvae, and the young anchovies, sardines, and related fish were the dominant forms in the brackish water (1-10 g/l) of the mid-estuary. The presence of large numbers of juvenile and young animals and young animals indicated the importance of these brackish waters as nursery grounds. Aquatic insects, cyclopoid copepods (Macrocyclops sp.), cladocerans, mysids (Taphromysis bowmani), ostracods (Cypridopsis sp. ), fresh-water prawns (Palaemonetes paludosus), and various marshfish were dominant in the ?fresh? headwaters. The amount of plant detritus collected in the estuary averaged about ten times that of the zooplankton. The estimated mean wet-weight of the zooplankton was 65 milligrams per cubic meter (mg/m?) and ranged from 1 to 173 mg/m?, with the smallest amounts occurring in the ?fresh? headwaters. Nekton, consisting of small fish and prawns, ranged from 3 to 214 mg/m? in weight and had a mean of 30 mg/m?. Largest catches were made in the headwaters at the end of the dry season, where the weight of the standing crop increased more than 15 times during the sampling period. The small fish and prawns, which were concentrated in the headwaters at the water level dropped, served as a rich source of food for predatory marine fish and birds.

Open-File Report↗

Surface-water-quality assessment of the Yakima River basin in Washington: major-and minor-element data for sediment, water, and aquatic biota, 1987-91

Major- and minor-element concentrations are presented for streambed and suspended sediment, filtered- and unfiltered-water, and aquatic-biota samples collected during 1987-91 from the Yakima River Basin in south-central Washington. The samples were collected as part of the U.S. Geological Survey's National Water-quality Assessment (NAWQA) program which is designed to provide results that are useful in understanding and managing the Nation's water resources. This report includes the sampling approach, field collection and processing techniques, and methods of chemical analysis, as well as a compilation of chemical data, statistical summaries, and quality- control data. These data may be used by scientists and resource managers to describe (1) spatial distribution of selected major and minor elements in sediment, water, and aquatic biota of the Yakima River Basin; (2) temporal variation for element concentrations in filtered water and in suspended sediment at selected sites; (3) suita- bility of surface water for preservation of aquatic life and protection of human health; and (4) major natural and anthropogenic sources of major and minor elements in the Yakima River Basin that affect observed water-quality conditions. Streambed-sediment samples were collected once from 27 sites in the basin during 1987-91. Suspended-sediment and filtered-water samples were collected monthly and during hydrologic events (including snowmelt and winter rainstorms) at seven sites, and filtered-water samples were collected at least once at an additional 37 sites during synoptic samplings. Unfiltered-water samples were collected at seven sites on a quarterly basis during 1987 only. Samples of aquatic plants were collected once in 1989, and aquatic insects, fish, and clams were collected from 34 sites three times during 1989-90.

Washington↗

Water-quality, bed-sediment, and biological data (October 1992 through September 1993) and statistical summaries of water-quality data (March 1985 through September 1993) for streams in the upper Clark Fork basin, Montana

Water, bed sediment, and biota were sampled in streams from Butte to below Missoula as part of a program to characterize aquatic resources in the upper Clark Fork basin of western Montana. Water-quality data were obtained periodically at 16 stations during October 1992 through September 1993 (water year 1993); daily suspended-sediment data were obtained at six of these stations. Bed-sediment and biological data were obtained at 11 stations in August 1993. Sampling stations were located on the Clark Fork and major tributaries. The primary constituents analyzed were trace elements associated with mine tailings from historic mining and smelting activities. Water-quality data include concentra- tions of major ions, trace elements, and suspended sediment in samples collected periodically during water year 1993. A statistical summary of water- quality data is provided for the period of record at each station since 1985. Daily values of streamflow, suspended-sediment concentration, and suspended-sediment discharge are given for six stations. Bed-sediment data include trace- element concentrations in the fine and bulk fractions. Biological data include trace-element concentrations in whole-body tissue of aquatic benthic insects. Quality-assurance data are reported for analytical results of water, bed sediment, and biota.

Open-File Report↗

Biological and associated water-quality data for lower Olmos Creek and upper San Antonio River, San Antonio, Texas, March-October 1990

Biological and associated water-quality data were collected from lower Olmos Creek and upper San Antonio River in San Antonio, Texas, during March-October 1990, the second year of a multiyear data-collection program. The data will be used to document water-quality conditions prior to implementation of a proposal to reuse treated wastewater to irrigate city properties in Olmos Basin and Brackenridge Parks and to augment flows in the Olmos Creek/San Antonio River system. Benthic macroinvertebrate, periphyton, and phy toplankton communities were sampled at three sites along a 4.2-kilometer reach of the Olmos Creek/San Antonio River system. Total mean densities of benthic macroinvertebrates for the three sites ranged from 1,000 to 15,000 organisms per square meter. The most abundant macroinvertebrates were the class Insecta (insects). Total densities of periphyton ranged from 700 to 86,000 cells per square millimeter. Cyanophyta (blue-green algae) and Bacillariophyta (diatoms) were the predominant periphyton organisms. Total densities of phytoplankton ranged from 940 to 21,000 cells per milliliter. Diatoms and Chlorophyta (green algae) made up most of the rest of the phytoplankton. Mean diversity index for the three sites ranged from 1.0 to 2.1 for benthic macroinvertebrates. The diversity index for all sites ranged from 1.5 to 3.1 for periphyton and 0.9 to 2.5 for phytoplankton. Hardness ranged from 220 to 280 milligrams per liter as calcium carbonate, and alkalinity ranged from 173 to 213 milligrams per liter as calcium carbonate. The largest dissolved nitrate concentration was 1.68 milligrams per liter. Dissolved ammonia nitrogen generally was less than 0.100 milligram per liter. The largest total phosphorus concentration was 0.080 milligram per liter, over one-half of which was dissolved orthophosphate. Total aluminum and total iron were the only trace elements in water to exceed the reporting threshold by large concentrations. Total aluminum concentrations ranged from 100 to 250 micrograms per liter, and total iron concentrations ranged from 70 to 280 micrograms per liter. Lead was the most prominent trace element in bottommaterial samples with concentrations ranging from 40 to 190 micrograms per gram. The maximum copper concentration in the bottom material was 60 micrograms per gram at site 3 (the most downstream site). Depths and velocities, measured at sample points after benthic macroinvertebrate sampling, ranged from 0.03 to 0.30 meter and from 0.06 to 1.2 meters per second, respectively. Measurable stream discharge ranged from 0.01 to 0.27 cubic meter per second. During two of the sampling periods, no flow was at site 1.

Open-File Report↗

Biological and associated water-quality data for lower Olmos Creek and upper San Antonio River, San Antonio, Texas, April - September 1989

Biological and associated water-quality data were collected from lower Olmos Creek and upper San Antonio River in San Antonio, Texas, during April-September 1989. Benthic macroinvertebrate, periphyton, and phytoplankton communities were sampled at three sites along the Olmos Creek/San Antonio River system. Total mean densities of benthic macroinvertebrates for the three sites ranged from 670 to 10,000 organisms per square meter. The most abundant macroinvertebrates were the class Insecta (insects). Total densities of periphyton ranged from 2,900 to 110,000 cells per square millimeter. Cyanophyta (blue-green algae) and Bacillariophyta (diatoms) were the predominant periphyton organisms. Total densities of phyto- plankton ranged from 5,000 to 47,000 cells per square milliliter. Blue-green algae accounted for more than one- half of the phytoplankton in each sample. Hardness ranged from 160 to 250 milligrams per liter as calcium carbonate, and alkalinity ranged from 130 to 220 milligrams per liter as calcium carbonate. The largest dissolved nitrite concentration was 0.038 milligram per liter. The largest total phosphorus concentration was 0.150 milligram per liter, over one-half of which was dissolved orthophosphate. Total aluminum and total iron were the only trace elements in water to exceed the reporting threshold by large concen- trations. Total aluminum concentrations ranged from 70 to 280 micrograms per liter, and total iron concentrations ranged from 70 to 340 micrograms per liter. Lead was the most prominent trace element in bottom-material samples, with concentrations ranging from 30 to 230 micrograms per gram.

Open-File Report↗

Water-quality, bed-sediment, and biological data (October 1993 through September 1994) and statistical summaries of data for streams in the Upper Clark Fork basin, Montana

Water, bed sediment, and biota were sampled in streams from Butte to below Missoula as part of a program to characterize aquatic resources in the upper Clark Fork basin of western Montana. Water- quality data were obtained periodically at 16 stations during October 1993 through September 1994 (water year 1994); daily suspended-sediment data were obtained at six of these stations. Bed-sediment and biological data were obtained at 11 stations in August 1994. Sampling stations were located on the Clark Fork and major tributaries. The primary constituents analyzed were trace elements associated with mine tailings from historical mining and smelting activities. Water-quality data include concentrations of major ions, trace elements, and suspended sediment in samples collected periodically during water year 1994. Daily values of streamflow, suspended-sediment concentration, and suspended- sediment discharge are given for six stations. Bed- sediment data include trace-element concentrations in the fine and bulk fractions. Biological data include trace-element concentrations in whole-body tissue of aquatic benthic insects. Quality-assurance data are reported for analytical results of water, bed sediment, and biota. Statistical summaries of bed sediment, and biological data are provided for the period of record at each station since 1985.

Open-File Report↗

Water-quality, bed-sediment, and biological data (October 1994 through September 1995) and statistical summaries of data for streams in the upper Clark Fork basin, Montana

Water, bed sediment, and biota were sampled in streams from Butte to below Missoula as part of a program to characterize aquatic resources in the upper Clark Fork basin of western Montana. Sampling stations were located on the Clark Fork and major tributaries. Water-quality data were obtained periodically at 16 stations during October 1994 through September 1995 (water year 1995). Data for twelve bed-sediment and eleven biological stations were obtained in August 1995. The primary constituents analyzed were trace elements associated with mine tailings from historical mining and smelting activities. Water-quality data include concentrations of major ions, trace element, and suspended sediment in stream samples collected periodically during water year 1995. Daily values of streamflow, suspended- sediment concentrations in the fine-grained and bulk fractions. Biological data include trace- element concentrations in whole-body tissue of aquatic benthic insects. Quality-assurance data are reported for analytical results of water, bed sediment, and biota. Statistical summaries of water-quality, bed-sediment, and biological data are provided for the period of record at each station since 1985.

Montana↗

Transport of diazinon in the San Joaquin River basin, California

Most of the application of the organophosphate insecticide diazinon in the San Joaquin River Basin occurs in winter to control wood boring insects in dormant almond orchards. A federal-state collaborative study found that diazinon accounted for most of the observed toxicity of San Joaquin River water to water fleas in February 1993. Previous studies focussed mainly on west-side inputs to the San Joaquin River. In this 1994 study, the three major east-side tributaries to the San Joaquin River, the Merced, Tuolumne, and Stanislaus Rivers, and a downstream site on the San Joaquin River were sampled throughout the hydrographs of a late January and an early February storm. In both storms, the Tuolumne River had the highest concentrations of diazinon and transported the largest load of the three tributaries. The Stanislaus River was a small source in both storms. On the basis of previous storm sampling and estimated traveltimes, ephemeral west-side creeks were probably the main diazinon source early in the storms, while the Tuolumne and Merced Rivers and east-side drainage directly to the San Joaquin River were the main sources later. Although 74 percent of diazinon transport in the San Joaquin River during 199193 occurred in January and February, transport during each of the two 1994 storms was only 0.05 percent of the amount applied during preceeding dry periods. Nevertheless, some of the diazinon concentrations in the San Joaquin River during the January storm exceeded 0.35 micrograms per liter, a concentration shown to be acutely toxic to water fleas. Diazinon concentrations were highly variable during the storms and frequent sampling was required to adequately describe the concentration curves and to estimate loads.

Open-File Report↗

Transport of sediment-bound organochlorine pesticides to the San Joaquin River, California

Most of the application of the organophosphate insecticide diazinon in the San Joaquin River Basin occurs in winter to control wood boring insects in dormant almond orchards. A federal-state collaborative study found that diazinon accounted for most of the observed toxicity of San Joaquin River water to water fleas in February 1993. Previous studies focused mainly on west-side inputs to the San Joaquin River. In this 1994 study, the three major east-side tributaries to the San Joaquin River, the Merced, Tuolumne, and Stanislaus Rivers, and a downstream site on the San Joaquin River were sampled throughout the hydrographs of a late January and an early February storm. In both storms, the Tuolumne River had the highest concentrations of diazinon and transported the largest load of the three tributaries. The Stanislaus River was a small source in both storms. On the basis of previous storm sampling and estimated traveltimes, ephemeral west-side creeks probably were the main diazinon source early in the storms, whereas the Tuolumne and Merced Rivers and east-side drainages directly to the San Joaquin River were the main sources later. Although 74 percent of diazinon transport in the San Joaquin River during 1991-1993 occurred in January and February, transport during each of the two 1994 storms was only 0.05 percent of the amount applied during preceding dry periods. Nevertheless, some of the diazinon concentrations in the San Joaquin River during the January storm exceeded 0.35 micrograms per liter, a concentration shown to be acutely toxic to water fleas. Diazinon concentrations were highly variable during the storms and frequent sampling was required to adequately describe the concentration curves and to estimate loads.

Open-File Report↗

Deglaciation and postglacial treeline fluctuation in the northern San Juan Mountains, Colorado

The San Juan Mountains of southwestern Colorado contain numerous lakes and bogs at and above treeline. In June 1978, Lake Emma, a tarn above present-day treeline, was suddenly drained by the collapse of underground mine workings. This study was initiated because the draining exposed a well-preserved archive of subfossil coniferous wood fragments that provided a unique opportunity to further our understanding of the paleoclimatic history of this region. These paleoclimatic studies-coniferous macrofossil identification in conjunction with radiocarbon dating, deuterium analysis of the dated conifer fragments, as well as pollen and fossil insect analyses-yielded new information regarding Holocene climate and accompanying treeline changes in the northern San Juan Mountains. This report synthesizes previously published reports by the author and other investigators, and unpublished information of the author bearing on late Pleistocene and Holocene treeline and climate in this region. Retreat of the glacier that occupied the upper Animas River valley from its Pinedale terminal position began about 19.4 + or - 1.5 10 Be thousands of years ago and was essentially complete by about 12.3 + or - 1.0 10 Be thousands of years ago. Two sets of late Pleistocene cirque moraines were identified in the northern San Juan Mountains. The older set is widespread and probably correlates with the Younger Dryas (11,000-10,000 radiocarbon years before present; 12,800-11,500 calendar years). The younger set is found only in the Grenadier Range and represents remnant glacier ice lying in well-shaded niches in a mountain range undergoing rapid deglaciation. A snowbank at the northern base of this range appears to be fronted by a Little Ice Age moraine. Soon after deglaciation the average July temperature is estimated to have been about 5°C cooler and timberline about 650 meters lower than at present. However, timberline (and treeline) responded rapidly to the postglacial warming and reached higher-than-present elevations by the early Holocene. A comparison of recently obtained accelerator mass spectrometry radiocarbon ages of coniferous wood fragments from Lake Emma, previously dated by conventional radiocarbon methods during the 1980s, led to a slight modification of previously published ages of Holocene treeline fluctuations. As early as 9,200 radiocarbon years before present (about 10,400 calendar years) and probably to about 5,400 radiocarbon years before present (about 6,200 calendar years), treeline was at least 80 meters higher than at present (about 3,660 meters). Furthermore, a large conifer fragment with a complacent annual ring record suggests that timberline may have been at least 140 meters higher than present (about 3,600 meters) about 8,000 radiocarbon years before present (about 8,900 calendar years). These past elevations of treeline and timberline suggest that growing-season temperatures were at least 0.5°-0.9°C warmer than at present. Deuterium data from the Lake Emma wood samples suggests that the maximum average temperature change from about 9,000 to 5,400 radiocarbon years before present (about 10,150 to 6,200 calendar years) was about 4°C. Owing to these warmer temperatures the summer monsoon circulation, which currently brings a large part of the annual precipitation to the San Juan Mountains, probably was more intense during the early and middle Holocene than it is today. Between about 5,400 and 3,500 radiocarbon years before present (about 6,200 and 3,770 calendar years) it appears that treeline was near its present-day limit. After 3,500 radiocarbon years before present (about 3,770 calendar years), evidence of treeline position is very sparse, suggesting that treeline lay at, or below, its present-day elevation. However, a spruce krummholz fragment from the Lake Emma site provided two radiocarbon ages of about 3,100 radiocarbon years before present (about 3,300 cal

Colorado↗

Willamette Valley Ecoregion: Chapter 3 in Status and trends of land change in the Western United States--1973 to 2000

The Willamette Valley Ecoregion (as defined by Omernik, 1987; U.S. Environmental Protection Agency, 1997) covers approximately 14,458 km² (5,582 mi 2 ), making it one of the smallest ecoregions in the conterminous United States. The long, alluvial Willamette Valley, which stretches north to south more than 193 km and ranges from 32 to 64 km wide, is nestled between the sedimentary and metamorphic Coast Ranges (Coast Range Ecoregion) to the west and the basaltic Cascade Range (Cascades Ecoregion) to the east (fig. 1). The Lewis and Columbia Rivers converge at the ecoregion’s northern boundary in Washington state; however, the majority of the ecoregion falls within northwestern Oregon. Interstate 5 runs the length of the valley to its southern boundary with the Klamath Mountains Ecoregion. Topography here is relatively flat, with elevations ranging from sea level to 122 m. This even terrain, coupled with mild, wet winters, warm, dry summers, and nutrient-rich soil, makes the Willamette Valley the most important agricultural region in Oregon. Population centers are concentrated along the valley floor. According to estimates from the Oregon Department of Fish and Wildlife (2006), over 2.3 million people lived in Willamette Valley in 2000. Portland, Oregon, is the largest city, with 529,121 residents (U.S. Census Bureau, 2000). Other sizable cities include Eugene, Oregon; Salem (Oregon’s state capital); and Vancouver, Washington. Despite the large urban areas dotting the length of the Willamette Valley Ecoregion, agriculture and forestry products are its economic foundation (figs. 2,3). The valley is a major producer of grass seed, ornamental plants, fruits, nuts, vegetables, and grains, as well as poultry, beef, and dairy products. The forestry and logging industries also are primary employers of the valley’s rural residents (Rooney, 2008). These activities have affected the watershed significantly, with forestry and agricultural runoff contributing to river sedimentation and decreased water quality in the Willamette River and its tributary streams (Oregon Department of Fish and Wildlife, 2006). Recent years have seen a marked decline in forest health related to the increased frequency of multiyear droughts. Insect damage and other diseases also are present; however, drought- related water stress is the primary factor in coniferous-tree mortality (Oregon Department of Forestry, 2008). Trees most at risk include Douglas-fir ( Pseudotsuga menziesii ), grand fir ( Abies grandis ), and western red cedar ( Thuja plicata ). Overstocking by timber companies and planting on sites with poor conditions increase susceptibility. Over time, these problems may lead to changes in planting practices and the use of more drought-tolerant species such as ponderosa pine ( Pinus ponderosa ).

Oregon↗

Montana Valley and Foothill Prairies Ecoregion: Chapter 6 in Status and trends of land change in the Western United States--1973 to 2000

The Montana Valley and Foothill Prairies Ecoregion comprises numerous intermountain valleys and low-elevation foothill prairies spread across the western half of Montana, on both sides of the Continental Divide (Omernik, 1987; U.S. Environmental Protection Agency, 1997). The ecoregion, which covers approximately 64,658 km 2 (24,965 mi 2 ), includes the Flathead Valley and the valleys surrounding Helena, Missoula, Bozeman, Billings, Anaconda, Dillon, and Lewistown (fig. 1). These valleys are generally characterized by shortgrass prairie vegetation and are flanked by forested mountains (Woods and others, 1999); thus, the valleys’ biotas with regards to fish and insects are comparable. In many cases, the valleys are conduits for some of the largest rivers in the state, including Clark Fork and the Missouri, Jefferson, Madison, Flathead, Yellowstone, Gallatin, Smith, Big Hole, Bitterroot, and Blackfoot Rivers (fig. 2). The Montana Valley and Foothill Prairies Ecoregion also includes the “Rocky Mountain front,” an area of prairies along the eastern slope of the northern Rocky Mountains. Principal land uses within the ecoregion include farming, grazing, and mining. The valleys serve as major transportation and utility corridors and also contain the majority of Montana’s human population. The Montana Valley and Foothill Prairies Ecoregion extends into 17 mostly rural counties throughout western Montana. Only three of the counties—Carbon, Yellowstone, and Missoula—are part of a metropolitan statistical area with contiguous built-up areas tied to an employment center. Nearly two-thirds of Montana residents live in nonmetropolitan counties (Albrecht, 2008). Ten of the counties within the ecoregion had population growth rates greater than national averages (9–13 percent) between 1970 and 2000 (table 1). Ravalli and Gallatin Counties had the highest growth rates. Population growth was largely due to amenity-related inmigration and an economy dependent on tourism, health care, and services. Counties that had population declines, such as Deer Lodge, Silver Bow, and Meagher Counties, also had declines in agriculture and mining activity, and they had railroad closures as well. Climate varies from north to south and from the east side of the Continental Divide to the west side. However, all areas are semiarid with long cold winters and short growing seasons. In the western part of the ecoregion, Beaverhead, Bitterroot, Flathead, and Lolo National Forests provide the natural resources, particularly timber, that form the economic base for towns within nearby valleys. Mineral resources from mines in and around Anaconda, Deer Lodge, and Butte have long provided an economic base for these towns (fig. 3).

Idaho;Montana;Wyoming↗

The effects of management practices on grassland birds—Greater Prairie-Chicken ( Tympanuchus cupido pinnatus )

The keys to Greater Prairie-Chicken ( Tympanuchus cupido pinnatus ) management are maintaining expansive grasslands; preventing populations of Greater Prairie-Chickens from becoming small and isolated; managing grasslands to maintain proper grassland height, density, and vigor; and reducing woody plant invasion and excessive litter buildup. Within these grasslands, areas should contain short herbaceous cover for lek sites; tall residual grasses for nesting; and disturbed habitats for broods with adequate vegetation regrowth that provides insects for food and cover from predators and weather. This account does not address population or harvest management but rather focuses on habitat management. Greater Prairie-Chickens have been reported to use habitats with 5–113 centimeter (cm) average vegetation height, 5–40 cm visual obstruction reading, 18–95 percent grass cover, 1–35 percent forb cover, <45 percent litter cover, <5 percent shrub cover, 3–25 percent bare ground, and <12 cm litter depth.

Professional Paper↗

Conifer health classification for Colorado, 2008

Colorado has undergone substantial changes in forests due to urbanization, wildfires, insect-caused tree mortality, and other human and environmental factors. The U.S. Geological Survey Rocky Mountain Geographic Science Center evaluated and developed a methodology for applying remotely-sensed imagery for assessing conifer health in Colorado. Two classes were identified for the purposes of this study: healthy and unhealthy (for example, an area the size of a 30- x 30-m pixel with 20 percent or greater visibly dead trees was defined as ?unhealthy?). Medium-resolution Landsat 5 Thematic Mapper imagery were collected. The normalized, reflectance-converted, cloud-filled Landsat scenes were merged to form a statewide image mosaic, and a Normalized Difference Vegetation Index (NDVI) and Renormalized Difference Infrared Index (RDII) were derived. A supervised maximum likelihood classification was done using the Landsat multispectral bands, the NDVI, the RDII, and 30-m U.S. Geological Survey National Elevation Dataset (NED). The classification was constrained to pixels identified in the updated landcover dataset as coniferous or mixed coniferous/deciduous vegetation. The statewide results were merged with a separate health assessment of Grand County, Colo., produced in late 2008. Sampling and validation was done by collecting field data and high-resolution imagery. The 86 percent overall classification accuracy attained in this study suggests that the data and methods used successfully characterized conifer conditions within Colorado. Although forest conditions for Lodgepole Pine (Pinus contorta) are easily characterized, classification uncertainty exists between healthy/unhealthy Ponderosa Pine (Pinus ponderosa), Pi?on (Pinus edulis), and Juniper (Juniperus sp.) vegetation. Some underestimation of conifer mortality in Summit County is likely, where recent (2008) cloud-free imagery was unavailable. These classification uncertainties are primarily due to the spatial and temporal resolution of Landsat, and of the NLCD derived from this sensor. It is believed that high- to moderate-resolution multispectral imagery, coupled with field data, could significantly reduce the uncertainty rates. The USGS produced a four-county follow-up conifer health assessment using high-resolution RapidEye remotely sensed imagery and field data collected in 2009.

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