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At least 685 records · Page 38Linked to original sources

Post-fire temporal trends in soil-physical and -hydraulic properties and simulated runoff generation: Insights from different burn severities in the 2013 Black Forest Fire, CO, USA

Burn severity influences on post-fire recovery of soil-hydraulic properties controlling runoff generation are poorly understood despite the importance for parameterizing infiltration models. We measured soil-hydraulic properties of field-saturated hydraulic conductivity (Kfs), sorptivity (S), and wetting front potential (ψf) for four years after the 2013 Black Forest Fire, Colorado, USA at six sites across a gradient of initial remotely sensed burn severity using the change in the normalized burn ratio (dNBR). These measurements were correlated with soil-physical property measurements of bulk density (ρb), loss on ignition (LOI, a measure of soil organic matter), and ground cover composition to provide insight into causal factors for temporal changes in Kfs, S, and ψf . Modeled infiltration using the Smith-Parlange approach parameterized with measured Kfs, S, and ψf further discerned the role of precipitation intensity on runoff generation. Temporal trends of soil-physical properties and ground cover showed influences from initial burn severity. Trends in soil-hydraulic properties , surprisingly, were not strongly influenced by initial burn severity despite inferred effects of ρb, LOI, and ground cover on trends in Kfs and S. Calculations of dNBR at the time of sampling showed strong correlations with Kfs and S, demonstrating a new approach for estimating long-unburned Kfs and S values, infiltration model parameters after fire, and assessing the time of return to pre-fire values. Simulated infiltration-excess runoff, in contrast, did depend on initial burn severity. Time series of the ratio S2/Kfs ≈ ψf tended to converge between 1 to 10 mm four years after wildfire, potentially (i) defining a long-unburned forest domain of S2/Kfs and ψf from 1 to 10 mm with relatively high Kfs values, and (ii) providing a new post-fire soil-hydraulic property recovery metric (i.e. S2/Kfs ≈ ψf in the range of 1 to 10 mm) for sites in the Rocky Mountains of the USA.

Colorado↗

Synergistic soil, land use, and climate influences on wind erosion on the Colorado Plateau: Implications for management

Two decades of drought in the southwestern USA are spurring concerns about increases in wind erosion, dust emissions, and associated impacts on ecosystems, agriculture, human health, and water supply. Different avenues of investigation into primary drivers of wind erosion and dust have yielded mixed results depending on the spatial and temporal sensitivity of the evidence. We monitored passive aeolian sediment traps from 2017 to 2020 across eighty-one sites near Moab UT to understand patterns of sediment flux. At measurement sites we collated climate, soil, topography and vegetation spatial layers to better understand the context of wind erosion and then combine these data with field observations of land use in models to characterize the influence of cattle grazing, oil and gas well pads, and vehicle/heavy equipment disturbance that potentially drive both exposure of bare soil and increases in erodible sediment supply that increase vulnerability to erosion. Disturbed areas with low soil calcium carbonate content yielded high sediment transport in dry years, but notably areas with little disturbance and low bare soil exposure had much less activity. Cattle grazing had the largest land use association with erosional activity with analyses suggesting that both herbivory and trampling from cattle could be drivers. The amount and distribution of bare soil exposure from new sub-annual fractional cover remote sensing products proved very helpful in mapping erosion, and new predictive maps informed by field data are presented to help depict spatial patterns of wind erosion activity. Our results suggest that despite the magnitude of current droughts, minimizing surface disturbance in vulnerable soils can mitigate a large portion of dust emissions. Results can help managers identify eroding areas where disturbance reduction and soil surface protection measures can be prioritized.

Science of the Total Environment↗

Weathering rates as a function of flow through an alpine soil

The effect of flow on release rates of solutes from soil in a 39-m2 alpine catchment in the Colorado Rockies was measured during the summers of 1990-1994. Flow rates through the soil were varied by augmenting natural rainfall with deionized irrigation water. Daily water inputs averaged between 96 and 216 1 day-1 during the five field seasons, and mean discharge (inputs minus evapotranspiration) varied from 35 to 175 1 day-1. Volume-weighted mean concentrations of base cations and silica decreased only moderately in response to the increased water inputs. Input fluxes of solutes in precipitation were similar in each of the study seasons, but output fluxes of base cations and silica in surface outflow increased substantially in conjunction with the average water input rate for the season. Weathering rates calculated from the chemical fluxes increased substantially in response to increases in water input rates. The increases appear to be largely attributable to enhanced transport of solutes from the soil matrix under high flow conditions. At high flow, physical flushing of micropores presumably occurs to a greater extent than during low-flow periods because of greater soil wetness and higher hydrologic head. Increased flushing would also cause an increased rate of diffusion of solutes from microcracks in mineral surfaces and constricted pore spaces in response to an increased concentration gradient between those regions and adjacent areas in the soil matrix. Another consequence of the increased flushing that occurs during periods of high flow is that concentrations throughout the soil matrix tend to be lower, which might increase chemical weathering rates of some silicate minerals such as microcline, which are relatively close to saturation. Decreased Si concentrations under high-flow conditions appear to promote dissolution of amorphous aluminosilicates or desorption of Si from mineral surfaces, buffering Si concentrations in the soil solutions. Thus, both physical transport of solutes and subsequent chemical effects appear to be responsible for the positive relation observed between fluxes of weathering products and water input rates.

Chemical Geology↗

Soil quality assessment using weighted fuzzy association rules

Fuzzy association rules (FARs) can be powerful in assessing regional soil quality, a critical step prior to land planning and utilization; however, traditional FARs mined from soil quality database, ignoring the importance variability of the rules, can be redundant and far from optimal. In this study, we developed a method applying different weights to traditional FARs to improve accuracy of soil quality assessment. After the FARs for soil quality assessment were mined, redundant rules were eliminated according to whether the rules were significant or not in reducing the complexity of the soil quality assessment models and in improving the comprehensibility of FARs. The global weights, each representing the importance of a FAR in soil quality assessment, were then introduced and refined using a gradient descent optimization method. This method was applied to the assessment of soil resources conditions in Guangdong Province, China. The new approach had an accuracy of 87%, when 15 rules were mined, as compared with 76% from the traditional approach. The accuracy increased to 96% when 32 rules were mined, in contrast to 88% from the traditional approach. These results demonstrated an improved comprehensibility of FARs and a high accuracy of the proposed method.

Pedosphere↗

Migration through soil of organic solutes in an oil-shale process water

The migration through soil of organic solutes in an oil-shale process water (retort water) was studied by using soil columns and analyzing leachates for various organic constituents. Retort water extracted significant quantities of organic anions leached from ammonium-saturated-soil organic matter, and a distilled-water rinse, which followed retort-water leaching, released additional organic acids from the soil. After being corrected for organic constitutents extracted from soil by retort water, dissolved-organic-carbon fractionation analyses of effluent fractions showed that the order of increasing affinity of six organic compound classes for the soil was as follows: hydrophilic neutrals nearly equal to hydrophilic acids, followed by the sequence of hydrophobic acids, hydrophilic bases, hydrophobic bases, and hydrophobic neutrals. Liquid-chromatographic analysis of the aromatic amines in the hydrophobic- and hydrophilic-base fractions showed that the relative order of the rates of migration through the soil column was the same as the order of migration on a reversed-phase, octadecylsilica liquid-chromatographic column.

Environmental Science & Technology↗

Environmental fate of roxarsone in poultry litter. Part II. Mobility of arsenic in soils amended with poultry litter

Poultry litter often contains arsenic as a result of organo-arsenical feed additives. When the poultry litter is applied to agricultural fields, the arsenic is released to the environment and may result in increased arsenic in surface and groundwater and increased uptake by plants. The release of arsenic from poultry litter, litter-amended soils, and soils without litter amendment was examined by extraction with water and strong acids (HCI and HN03). The extracts were analyzed for As, C, P, Cu, Zn, and Fe. Copper, zinc, and iron are also poultry feed additives. Soils with a known history of litter application and controlled application rate of arsenic-containing poultry litter were obtained from the University of Maryland Agricultural Experiment Station. Soils from fields with long-term application of poultry litter were obtained from a tilled field on the Delmarva Peninsula (MD) and an untilled Oklahoma pasture. Samples from an adjacent forest or nearby pasture that had no history of litter application were used as controls. Depth profiles were sampled for the Oklahoma pasture soils. Analysis of the poultry litter showed that 75% of the arsenic was readily soluble in water. Extraction of soils shows that weakly bound arsenic mobilized by water correlates positively with C, P, Cu, and Zn in amended fields and appears to come primarily from the litter. Strongly bound arsenic correlates positively with Fe in amended fields and suggests sorption or coprecipitation of As and Fe in the soil column.

Environmental Science & Technology↗

Dynamic replacement and loss of soil carbon on eroding cropland

Links between erosion/sedimentation history and soil carbon cycling were examined in a highly erosive setting in Mississippi loess soils. We sampled soils on (relatively) undisturbed and cropped hillslopes and measured C, N, 14 C, and CO 2 flux to characterize carbon storage and dynamics and to parameterize Century and spreadsheet 14 C models for different erosion and tillage histories. For this site, where 100 years of intensive cotton cropping were followed by fertilization and contour plowing, there was an initial and dramatic decline in soil carbon content from 1870 to 1950, followed by a dramatic increase in soil carbon. Soil erosion amplifies C loss and recovery: About 100% of the original, prehistoric soil carbon was likely lost over 127 years of intensive land use, but about 30% of that carbon was replaced after 1950. The eroded cropland was therefore a local sink for CO 2 since the 1950s. However, a net CO 2 sink requires a full accounting of eroded carbon, which in turn requires that decomposition rates in lower slopes or wetlands be reduced to about 20% of the upland value. As a result, erosion may induce unaccounted sinks or sources of CO 2 , depending on the fate of eroded carbon and its protection from decomposition. For erosion rates typical of the United States, the sink terms may be large enough (1 Gt yr −1 , back‐of‐the‐envelope) to warrant a careful accounting of site management, cropping, and fertilization histories, as well as burial rates, for a more meaningful global assessment.

Global Biogeochemical Cycles↗

Weathering controls on mechanisms of carbon storage in grassland soils

On a sequence of soils developed under similar vegetation, temperature, and precipitation conditions, but with variations in mineralogical properties, we use organic carbon and 14C inventories to examine mineral protection of soil organic carbon. In these soils, 14C data indicate that the creation of slow-cycling carbon can be modeled as occurring through reaction of organic ligands with Al3+ and Fe3+ cations in the upper horizons, followed by sorption to amorphous inorganic Al compounds at depth. Only one of these processes, the chelation Al3+ and Fe3+ by organic ligands, is linked to large carbon stocks. Organic ligands stabilized by this process traverse the soil column as dissolved organic carbon (both from surface horizons and root exudates). At our moist grassland site, this chelation and transport process is very strongly correlated with the storage and long-term stabilization of soil organic carbon. Our 14C results show that the mechanisms of organic carbon transport and storage at this site follow a classic model previously believed to only be significant in a single soil order (Spodosols), and closely related to the presence of forests. The presence of this process in the grassland Alfisol, Inceptisol, and Mollisol soils of this chronosequence suggests that this process is a more significant control on organic carbon storage than previously thought. Copyright 2004 by the American Geophysical Union.

Global Biogeochemical Cycles↗

Effects of native forest restoration on soil hydraulic properties, Auwahi, Maui, Hawaiian Islands

Over historic time Hawai'i's dryland forests have been largely replaced by grasslands for grazing livestock. On-going efforts have been undertaken to restore dryland forests to bring back native species and reduce erosion. The reestablishment of native ecosystems on land severely degraded by long-term alternative use requires reversal of the impacts of erosion, organic-matter loss, and soil structural damage on soil hydraulic properties. This issue is perhaps especially critical in dryland forests where the soil must facilitate native plants' optimal use of limited water. These reforestation efforts depend on restoring soil ecological function, including soil hydraulic properties. We hypothesized that reforestation can measurably change soil hydraulic properties over restoration timescales. At a site on the island of Maui (Hawai'i, USA), we measured infiltration capacity, hydrophobicity, and abundance of preferential flow channels in a deforested grassland and in an adjacent area where active reforestation has been going on for fourteen years. Compared to the nearby deforested rangeland, mean field-saturated hydraulic conductivity in the newly restored forest measured by 55 infiltrometer tests was greater by a factor of 2.0. Hydrophobicity on an 8-point scale increased from average category 6.0 to 6.9. A 4-point empirical categorization of preferentiality in subsurface wetting patterns increased from an average 1.3 in grasslands to 2.6 in the restored forest. All of these changes act to distribute infiltrated water faster and deeper, as appropriate for native plant needs. This study indicates that vegetation restoration can lead to ecohydrologically important changes in soil hydraulic properties over decadal time scales.

Hawai'i↗

Modeling thermal dynamics of active layer soils and near-surface permafrost using a fully coupled water and heat transport model

Thawing and freezing processes are key components in permafrost dynamics, and these processes play an important role in regulating the hydrological and carbon cycles in the northern high latitudes. In the present study, we apply a well-developed soil thermal model that fully couples heat and water transport, to simulate the thawing and freezing processes at daily time steps across multiple sites that vary with vegetation cover, disturbance history, and climate. The model performance was evaluated by comparing modeled and measured soil temperatures at different depths. We use the model to explore the influence of climate, fire disturbance, and topography (north- and south-facing slopes) on soil thermal dynamics. Modeled soil temperatures agree well with measured values for both boreal forest and tundra ecosystems at the site level. Combustion of organic-soil horizons during wildfire alters the surface energy balance and increases the downward heat flux through the soil profile, resulting in the warming and thawing of near-surface permafrost. A projection of 21st century permafrost dynamics indicates that as the climate warms, active layer thickness will likely increase to more than 3 meters in the boreal forest site and deeper than one meter in the tundra site. Results from this coupled heat-water modeling approach represent faster thaw rates than previously simulated in other studies. We conclude that the discussed soil thermal model is able to well simulate the permafrost dynamics and could be used as a tool to analyze the influence of climate change and wildfire disturbance on permafrost thawing.

Journal of Geophysical Research D: Atmospheres↗

Soil genesis on the island of Bermuda in the Quaternary: the importance of African dust transport and deposition

The origin of terra rossa, red or reddish-brown, clay-rich soils overlying high-purity carbonate substrates, has intrigued geologists and pedologists for decades. Terra rossa soils can form from accumulation of insoluble residues during dissolution of the host limestones, addition of volcanic ash, or addition of externally derived, long-range-transported (LRT) aeolian particles. We studied soils and paleosols on high-purity, carbonate aeolianites of Quaternary age on Bermuda, where terra rossa origins have been debated for more than a century. Potential soil parent materials on this island include sand-sized fragments of local volcanic bedrock, the LRT, fine-grained (<20 &mu;m) component of distal loess from the lower Mississippi River Valley, and LRT dust from Africa. These parent materials can be characterized geochemically using trace elements that are immobile in the soil-forming environment. Results indicate that local volcanic bedrock on Bermuda has Sc-Th-La, Cr-Ta-Nd, and Eu/Eu*, La N /Yb N , Gd N /Yb N that can be distinguished from African dust and lower Mississippi River valley loess. Bermuda soils have Sc-Th-La, Cr-Ta-Nd, and Eu/Eu*, La N /Yb N , Gd N /Yb N that indicate derivation from a combination of LRT dust from Africa and local volcanic bedrock. Our results indicate that soils on islands in a very broad latitudinal belt of the western Atlantic margin have been influenced by African LRT dust inputs over much of the past &ndash;500 ka.

Journal of Geophysical Research↗

Quantifying soil surface change in degraded drylands: shrub encroachment and effects of fire and vegetation removal in a desert grassland

Woody plant encroachment, a worldwide phenomenon, is a major driver of land degradation in desert grasslands. Woody plant encroachment by shrub functional types ultimately leads to the formation of a patchy landscape with fertile shrub patches interspaced with nutrient-depleted bare soil patches. This is considered to be an irreversible process of land and soil degradation. Recent studies have indicated that in the early stages of shrub encroachment, when there is sufficient herbaceous connectivity, fires (prescribed or natural) might provide some reversibility to the shrub encroachment process by negatively affecting shrub demography and homogenizing soil resources across patches within weeks to months after burning. A comprehensive understanding of longer term changes in microtopography and spatial patterning of soil properties following fire in shrub-encroached grasslands is desirable. Here, we investigate the changes in microtopography with LiDAR (light detection and ranging), vegetation recovery, and spatial pattering of soil properties in replicated burned, clipped, and control areas in a shrub-grass transition zone in the northern Chihuahuan Desert four years after prescribed fire or clipping. Results indicate a greater homogeneity in soil, microtopography, and vegetation patterning on burned relative to clipped and control treatments. Findings provide further evidence that disturbance by prescribed fire may allow for reversal of the shrub encroachment process, if the event occurs in the early stages of the vegetation shift. Improved understanding of longer-term effects of fire and associated changes in soil patterning can inform the use and role of fire in the context of changing disturbance regimes and climate.

Chihuahuan Desert↗

Modeling transient soil moisture limitations on microbial carbon respiration: A cost-performance comparison

Soil microorganisms are known to survive periods of aridity and to recover rapidly after wetting events, with the ability to transition between a dormant state in dry conditions and an active state in wet conditions. Though this dynamic behavior has been previously incorporated into soil carbon respiration modeling frameworks, a direct comparison between this active-dormant transition mechanism and a more simplified first-order model has yet to be made. Here, we demonstrate the necessary extent of model complexity needed to reproduce transient carbon respiration rates obtained from a set of soil incubation experiments implemented over a range of soil depths and time intervals. Two approaches are tested, one uses simplified first-order kinetics whereas the other employs a transition between active and dormant biomass. The performance of each model is evaluated using an Akaike Information Criterion (AIC) based on the accuracy with which they reproduce an experimental dataset consisting of two sets of time series soil incubations collected across a range of time and depth resolutions. Based on the AIC evaluation and model-data comparison, we conclude that a dormancy-enabled model featuring two distinct microbial strategists performs best for the majority of the soil profile (above 108 cm) for both high- and low- depth resolution and sampling frequency, despite the added parameters required. In contrast, the first-order model achieves better AIC scores when simulating our deepest soils (112-165 cm), where moisture fluctuations are expected to be less prevalent. These results guide how and where we choose to apply more cost intensive models.

Biogeosciences↗

Clumped isotopes record a glacial-interglacial shift in seasonality of soil carbonate accumulation in the San Luis Valley, southern Rocky Mountains, USA

Clumped isotope paleothermometry using pedogenic carbonates is a powerful tool for investigating past climate changes. However, location-specific seasonal patterns of precipitation and soil moisture cause systematic biases in the temperatures they record, hampering comparison of data across large areas or differing climate states. To account for biases, more systematic studies of carbonate forming processes are needed. We measured modern soil temperatures within the San Luis Valley of the Rocky Mountains and compared them to paleotemperatures determined using clumped isotopes. For Holocene-age samples, clumped isotope results indicate carbonate accumulated at a range of temperatures with site averages similar to the annual mean. Paleotemperatures for late Pleistocene-age samples (ranging 19–72 ka in age) yielded site averages only 2°C lower, despite evidence that annual temperatures during glacial periods were 5–9°C colder than modern. We use a 1D numerical model of soil physics to support the idea that differences in hydrologic conditions in interglacial versus glacial periods promote differences in the seasonal distribution of soil carbonate accumulation. Model simulations of modern (Holocene) conditions suggest that soil drying under low soil p CO 2 favors year-round carbonate accumulation in this region but peaking during post-monsoon soil drying. During a “glacial” simulation with lowered temperatures and added snowpack, more carbonate accumulation shifted to the summer season. These experiments show that changing hydrologic regimes could change the seasonality of carbonate accumulation, which in this study blunts the use of clumped isotopes to quantify glacial-interglacial temperature changes. This highlights the importance of understanding seasonal biases of climate proxies for accurate paleoenvironmental reconstruction.

Colorado, new Mexico↗

The effect of drying boreal lakes on plants, soils, and microbial communities in lake margin habitats

Decadal scale lake drying in interior Alaska results in lake margin colonization by willow shrub and graminoid vegetation, but the effects of these changes on plant production, biodiversity, soil properties, and soil microbial communities are not well known. We studied changes in soil organic carbon (SOC) and nitrogen (N) storage, plant and microbial community composition, and soil microbial activities in drying and non-drying lakes in the Yukon Flats National Wildlife Refuge. Historic changes in lake area were determined using Landsat imagery. Results showed that SOC storage in drying lake margins declined by 0.13 kg C m −2 yr −1 over 30 years of exposure of lake sediments, with no significant change in soil N. Lake drying resulted in an increase in graminoid and shrub aboveground net primary production (ANPP, +3% yr −1 ) with little change in plant functional composition. Increases in ANPP were similar in magnitude (but opposite in sign) to losses in SOC over a 30-year drying trend. Potential decomposition rates and soil enzyme activities were lower in drying lake margins compared to stable lake margins, possibly due to high salinities in drying lake margin soils. Microbial communities shifted in response to changing plant communities, although they still retained a legacy of the previous plant community. Understanding how changing lake hydrology impacts the ecology and biogeochemistry of lake margin terrestrial ecosystems is an underexamined phenomenon with large impacts to landscape processes.

Alaska↗

Discrete element investigation of the influence of shallow soil density on the manifestations of strike-slip surface fault rupture

This study investigates the influence of soil relative density on strike-slip surface fault rupture manifestations using three-dimensional numerical simulations performed with the discrete element method (DEM). The simulations capture the formation of distinctive fault strands within complex flower structures using tens of millions of grains. The tendency for dense soils to localize shear manifests as multiple localized shear bands within wide zones of deformation in strike-slip fault rupture, whereas diffuse shear deformation develops within narrow zones in loose soils. The spatial extents of soil deformation are consistent between simulations having similar relative densities but with different quantities of grains. However, individual shears are more distinguishable in assemblages of finer grains than in assemblages of coarser grains. The simulations show the progressive development of new shears within the bounds of previously developed shears. Shear activity transitions inward as fault activity diminishes along the outermost shears and continues along newly developed shears until a vertically dipping throughgoing shear structure develops that accommodates most of the fault displacement thereon. The throughgoing fault develops at smaller fault displacements in looser soils because the first shear rupture propagates closer to the vertical direction and does not undergo as much inward translation of shear activity, as is observed in denser soils. In all simulations, ground surface uplift develops between nonintersecting active shears, and ground surface subsidence tends to develop where new shears intersect previous shears. The surface traces in these simulations are shown to be consistent with analog models and case histories of surface fault rupture occurring in different shallow subsurface materials. Although computationally costly, these modeling results are valuable for providing a strong numerical supplement to traditional analog models used to represent the mechanics of strike-slip zones in soil, and they provide quantifiable stresses and large-strain deformations throughout the model domain.

Journal of Geotechnical and Geoenvironmental Engin↗

Relations between soil hydraulic properties and burn severity

Wildfire can affect soil hydraulic properties, often resulting in reduced infiltration. The magnitude of change in infiltration varies depending on the burn severity. Quantitative approaches to link burn severity with changes in infiltration are lacking. This study uses controlled laboratory measurements to determine relations between a remotely sensed burn severity metric ( dNBR , change in normalised burn ratio) and soil hydraulic properties (SHPs). SHPs were measured on soil cores collected from an area burned by the 2013 Black Forest fire in Colorado, USA. Six sites with the same soil type were selected across a range of burn severities, and 10 random soil cores were collected from each site within a 30-m diameter circle. Cumulative infiltration measurements were made in the laboratory using a tension infiltrometer to determine field-saturated hydraulic conductivity, K fs , and sorptivity, S . These measurements were correlated with dNBR for values ranging from 124 (low severity) to 886 (high severity). SHPs were related to dNBR by inverse functions for specific conditions of water repellency (at the time of sampling) and soil texture. Both functions had a threshold value for dNBR between 124 and 420, where K fs and S were unchanged and equal to values for soil unaffected by fire. For dNBR s >~420, the K fs was an exponentially decreasing function of dNBR and S was a linearly decreasing function of dNBR . These initial quantitative empirical relations provide a first step to link SHPs to burn severity, and can be used in quantitative infiltration models to predict post-wildfire infiltration and resulting runoff.

International Journal of Wildland Fire↗

Alder ( Alnus crispa ) effects on soils in ecosystems of the Agashashok River valley, northwest Alaska

At the northern limit of the boreal forest biome, alder ( Alnus crispa [Ait.] Pursh) shrubs occur in a variety of ecosystems. We assessed the effects of individual alder shrubs on soil properties and understory plant tissue nitrogen in floodplain terraces, valley slopes and tussock tundra ridges. The three ecosystems differed with respect to soil properties and abiotic conditions and supported distinct plant communities. Alder increased resin-exchangeable soil N and NO 3 production significantly in each ecosystem. The greatest difference between alder canopy and surrounding soil NO 3 measured both under field and laboratory conditions occurred in floodplain sites. The shrub effect on soil pH and soil organic matter was greatest on tundra ridges. Alder shrubs also influenced the nitrogen nutrition of plants growing beneath their canopies. Plants growing below alder canopies had higher foliar nitrogen concentration and natural abundance 15 N composition and lower carbon to nitrogen ratio than open-grown plants. Similar to soil N availability, understory plant leaf chemistry responded more to alder on floodplains than on slope or tundra ecosystems. This pattern suggests that understory plants rely more heavily on alder-fixed-N in this resource-poor ecosystem.

Alaska↗