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U.S. Geological Survey Karst Interest Group Proceedings, Carlsbad, New Mexico, April 29-May 2, 2014

Karst aquifer systems are present throughout parts of the United States and some of its territories, and have developed in carbonate rocks (primarily limestone and dolomite) that span an interval of time encompassing more than 550 million years. The depositional environments, diagenetic processes, post-depositional tectonic events, and geochemical weathering processes that form karst aquifers are varied and complex, and involve biological, chemical, and physical changes. These factors, combined with the diverse climatic regimes under which karst development in these rocks has taken place, result in the unique dual- or triple-porosity nature of karst aquifers. These complex hydrogeologic systems typically represent challenging and unique conditions to scientists attempting to study groundwater flow and contaminant transport in these terrains. The dissolution of carbonate rocks and the subsequent development of distinct and beautiful landscapes, caverns, and springs has resulted in the most exceptional karst areas of the United States being designated as national or state parks; commercial caverns and known privately owned caves number in the tens of thousands. Both public and private properties provide access for scientists to study the flow of groundwater in situ. Likewise, the range and complexity of landforms and groundwater flow systems associated with karst terrains are enormous, perhaps more than for any other aquifer type. Karst aquifers and landscapes that form in tropical areas, such as the cockpit karst along the north coast of Puerto Rico, differ greatly from karst landforms in more arid climates, such as the Edwards Plateau in west-central Texas or the Guadalupe Mountains near Carlsbad, New Mexico, where hypogenic processes have played a major role in speleogenesis. Many of these public and private lands also contain unique flora and fauna associated with these karst hydrogeologic systems. As a result, numerous federal, state, and local agencies have a strong interest in the study of karst terrains. Many of the major springs and aquifers in the United States have developed in carbonate rocks, such as the Floridan aquifer system in Florida and parts of Alabama, Georgia, and South Carolina; the Ozark Plateaus aquifer system in parts of Arkansas, Kansas, Missouri, and Oklahoma; and the Edwards-Trinity aquifer system in west-central Texas. These aquifers, and the springs that discharge from them, serve as major water-supply sources and as unique ecological habitats. Competition for the water resources of karst aquifers is common, and urban development and the lack of attenuation of contaminants in karst areas can impact the ecosystem and water quality of these aquifers. The concept for developing a platform for interaction among scientists within the U.S. Geological Survey (USGS) working on karst-related studies evolved from the November 1999 National Ground-Water Meeting of the USGS. As a result, the Karst Interest Group (KIG) was formed in 2000. The KIG is a loose-knit, grass-roots organization of USGS and non-USGS scientists and researchers devoted to fostering better communication among scientists working on, or interested in, karst science. The primary mission of the KIG is to encourage and support interdisciplinary collaboration and technology transfer among scientists working in karst areas. Additionally, the KIG encourages collaborative studies between the different mission areas of the USGS as well as other federal and state agencies, and with researchers from academia and institutes. The KIG also encourages younger scientists by participation of students in the poster and oral sessions. To accomplish its mission, the KIG has organized a series of workshops that are held near nationally important karst areas. To date (2014) six KIG workshops, including the workshop documented in this report, have been held. The workshops typically include oral and poster sessions on selected karst-related topics and research, as well as field trips to local karst features. Proceedings of the workshops are published by the USGS and are available online at http://water.usgs.gov/ogw/karst/kig . The first KIG workshop was held in St. Petersburg, Florida, February 13–16, 2001, in the vicinity of the large springs and other karst features of the Floridan aquifer system. The second KIG workshop was held August 20–22, 2002, in Shepherdstown, West Virginia, in proximity to the carbonate aquifers of the northern Shenandoah Valley and highlighted an invited presentation on karst literature by the late Barry F. Beck of P.E. LaMoreaux and Associates. The third KIG workshop was held September 12–15, 2005, in Rapid City, South Dakota, nearby to karst features in evaporites and limestones of the Madison Group in the Black Hills of South Dakota, including Wind Cave National Park and Jewel Cave National Monument. The workshop also included a featured presentation by Thomas Casadevall, Central Region Director, USGS, on the status of earth science at the USGS and evening trips to Jewel Cave led by Mike Wiles, National Park Service (NPS) and Wind Cave led by Rod Horrocks, NPS. The fourth KIG workshop was held May 27–29, 2008, and hosted by the Hoffman Environmental Research Institute and Center for Cave and Karst Studies at Western Kentucky University in Bowling Green, Kentucky, near Mammoth Cave National Park and karst features of the Chester Upland and Pennyroyal Plateau. The workshop featured a late-night field trip into Mammoth Cave with Rickard Toomey and Rick Olsen, NPS. The fifth workshop was held April 26–29, 2011, and was a joint meeting of the USGS KIG and University of Arkansas HydroDays, hosted by the Department of Geosciences at the University of Arkansas in Fayetteville. The workshop featured an outstanding field trip to the unique karst terrain along the Buffalo National River of the southern Ozarks and a keynote presentation on paleokarst in the United States by Art and Peggy Palmer. This sixth and current 2014 KIG workshop is hosted by the National Cave and Karst Research Institute (NCKRI) in Carlsbad, New Mexico, with Director of NCKRI, George Veni, serving as co-chair of the workshop with Eve Kuniansky, USGS. The session planning committee for this sixth workshop includes Van Brahana, USGS retired and University of Arkansas Professor Emeritus; Tom Byl, USGS and Tennessee State University; Zelda Bailey, former Director of NCKRI and retired Director, National Institute of Standards and Technology, Boulder Laboratory, Colorado; Patrick Tucci, USGS retired; and Mike Bradley, Allan Clark, Geoff Delin, Daniel Doctor, James Kaufmann, Eve Kuniansky, Randy Orndorff, Larry Spangler, and Dave Weary of the USGS. The karst hydrology field trip on Thursday will be led by Lewis Land (NCKRI karst hydrologist) and the optional Friday field trip on the geology of Carlsbad Caverns National Park will be led by George Veni. The keynote speaker is Dr. Penelope Boston, Director of Cave and Karst Studies at New Mexico Tech, Socorro, and Academic Director at NCKRI, who will address the future of karst research. Additionally, there is a featured presentation “Irish karst and its management,” by Caoimhe Hickey, The Geological Survey of Ireland, preceding a panel discussion on “Collaboration During Times of Limited Resources.” The extended abstracts of USGS authors were peer reviewed and approved for publication by the U.S. Geological Survey. Articles submitted by university researchers and other federal and state agencies did not go through the formal USGS peer review and approval process, and therefore may not adhere to our editorial standards or stratigraphic nomenclature and is not research conducted or data collected by the USGS. However, all articles had at a minimum of two peer reviews, and all articles were edited for consistency of appearance in the published Proceedings. The use of trade, firm or product names in any article is for descriptive purposes only and does not imply endorsement by the U.S. Government. The USGS, Office of Groundwater, provides technical support for the Karst Interest Group website and public availability of the Proceedings from these workshops, and the USGS Groundwater Resources Program funds the publication costs. Finally, the cover illustration is the work of Ann Tihansky, USGS, used since the first KIG workshop in 2000.

Scientific Investigations Report↗

The effects of bird use on nutrient removal in a constructed wastewater-treatment wetland

A 9.9-ha constructed wetland designed to reduce nitrogen in municipal wastewater following conventional secondary treatment began operating in southern California's San Jacinto Valley in September 1994. The wetland incorporated zones of bulrush ( Schoenoplectus acutus and S. californicus ) for effluent treatment, plus areas of 1.8-m deep open water and other features to benefit wintering waterfowl. A one-year long program to monitor bird use and evaluate their contribution to loadings of nitrogen and phosphorus was initiated seven months later and a second, four-month long period of monitoring was initiated after a 20-month hiatus. Daily bird use peaked at nearly 12,000 individuals during the second period. Estimates of maximum daily nitrogen and phosphorus input by birds were 139 g N ha −1 day −1 and 56 g P ha −1 day −1 . Following a reconfiguration of the wetland that increased the area of open water, a third year-long period of monitoring was initiated in September 2000. Estimated maximum daily loading attributable to birds during this period reached 312 g N ha −1 day −1 and 124 g P ha −1 day −1 . These levels represent only 2.6% and 7.0%, respectively, of the mean daily loads of N and P in inflow water from the wastewater-treatment plant. Wintering waterfowl contributed the most to nutrient loading, but the numerically dominant species was the colonial Red-winged Blackbird ( Agelaius phoeniceus ). The wetland's nutrient-removal efficiency was negatively correlated to bird loading. However, the greatest bird loading occurred during November to March, when winter conditions would reduce microbial nutrient-removal processes and plant uptake in the wetland. Multiple regression analysis indicated that variation in nutrient removal efficiency over a one-year period was best explained by wetland water temperature (R 2 = 0.21) and that little additional insight was gained by adding bird loading and inflow nutrient load data (R 2 = 0.22). This case study supports the concept that a constructed wetland can be designed both to reduce nutrients in municipal wastewater and to provide habitat for wetland birds.

Wetlands↗

Bedrock depth influences spatial patterns of summer baseflow, temperature and flow disconnection for mountainous headwater streams

In mountain headwater streams, the quality and resilience of summer cold-water habitat is generally regulated by stream discharge, longitudinal stream channel connectivity and groundwater exchange. These critical hydrologic processes are thought to be influenced by the stream corridor bedrock contact depth (sediment thickness), a parameter often inferred from sparse hillslope borehole information, piezometer refusal and remotely sensed data. To investigate how local bedrock depth might control summer stream temperature and channel disconnection (dewatering) patterns, we measured stream corridor bedrock depth by collecting and interpreting 191 passive seismic datasets along eight headwater streams in Shenandoah National Park (Virginia, USA). In addition, we used multi-year stream temperature and streamflow records to calculate several baseflow-related metrics along and among the study streams. Finally, comprehensive visual surveys of stream channel dewatering were conducted in 2016, 2019 and 2021 during summer low flow conditions (124 total km of stream length). We found that measured bedrock depths along the study streams were not well-characterized by soils maps or an existing global-scale geologic dataset where the latter overpredicted measured depths by 12.2 m (mean) or approximately four times the average bedrock depth of 2.9 m. Half of the eight study stream corridors had an average bedrock depth of less than 2 m. Of the eight study streams, Staunton River had the deepest average bedrock depth (3.4 m), the coldest summer temperature profiles and substantially higher summer baseflow indices compared to the other study steams. Staunton River also exhibited paired air and water annual temperature signals suggesting deeper groundwater influence, and the stream channel did not dewater in lower sections during any baseflow survey. In contrast, Paine Run and Piney River did show pronounced, patchy channel dewatering, with Paine Run having dozens of discrete dry channel sections ranging from 1 to greater than 300 m in length. Stream dewatering patterns were apparently influenced by a combination of discrete deep bedrock (20+ m) features and more subtle sediment thickness variation (1–4 m) depending on local stream valley hydrogeology. In combination, these unique datasets show the first large-scale empirical support for existing conceptual models of headwater stream disconnection based on spatially variable underflow capacity and shallow groundwater supply.

Virginia↗

Water-quality assessment of the Rio Grande Valley, Colorado, New Mexico and Texas: Fish communities at selected sites, 1993-95

Fish communities at 10 sites in the Rio Grande Basin were sampled during low-flow periods between 1993 and 1995 as part of the U.S. Geological Survey National Water-Quality Assessment Program. The ecology of fish communities is one of several lines of evidence used to characterize water-quality conditions. This report describes the fish communities at selected sites in the Rio Grande Basin and relates the structure of these fish communities to the physical and chemical characteristics of the streams. Twenty-nine species of fish representing 10 families were identified in 25 samples collected during this study. Species richness ranged from 1 to 13. Cluster analysis of the 25 samples collected during this study delineated four groups of sites that were based on the similarity of the fish communities. The first two groups were individual sites with low species richness. The third group contained the most samples, and the fourth group consisted of samples from the Rio Grande at Isleta, New Mexico, and the Rio Grande at El Paso, Texas. The shift in community structure of samples from group 3 to group 4 reflects changes from predominantly coldwater fishes to warmwater fishes. Four metrics of biotic integrity (percentages of introduced individuals, omnivores, tolerant individuals, and anomalies) were used in this study to provide a broad overview of the community structure. The relative percentages of introduced species at the Rio Grande near Del Norte, Colorado; Saguache Creek near Saguache, Colorado; Rio Grande below Taos Junction Bridge, near Taos, New Mexico; and Rio Grande at Isleta are indicative of biological stress on the communities at these sites. The dominance of omnivores in samples from the Rio Grande below Taos Junction Bridge, near Taos; Rio Chama near Chamita, New Mexico; Rio Grande at Isleta; and Rio Grande at El Paso is an indication of environmental stress at these sites. In 1995, tolerant species accounted for the entire fish community at the Rio Grande at Isleta. In all samples the occurrence of anomalies was less than 2 percent of the individuals, with the exception of the sample from the Rio Grande at Isleta. On the basis of the relative percentages of introduced individuals, omnivores, tolerant individuals, and anomalies, the biotic integrity at the Rio Grande at Isleta appears to be the most impaired of all sites and shows indications of potential chemical and physical perturbations. Fish communities from three reaches at the Santa Fe River above Cochiti Lake, New Mexico, and the Rio Grande at Isleta were sampled in 1995 to assess small-scale spatial patterns in the structure of fish communities. The spatial pattern at these sites might be associated with natural variability of the fish communities or with the presence of habitat features such as pools. The total number of individuals and relative abundance in a sample varied at sites sampled yearly during this study. All sites, with the exception of the Rio Grande near Del Norte, had a decline in total number of individuals in a sample. The temporal decline in the total number of individuals at these site might be associated with the natural variability within the fish communities.

Colorado, New Mexico, Texas↗

Distribution and movements of female northern pintails radiotagged in San Joaquin Valley, California

To improve understanding of northern pintail (Anas acuta) distribution in central California (CCA), we radiotagged 191 Hatch-Year (HY) and 228 After-Hatch-Year (AHY) female northern pintails during late August-early October, 1991-1993, in the San Joaquin Valley (SJV) and studied their movements through March each year. Nearly all (94.3%) wintered in CCA, but 5.7% went to southern California, Mexico, or unknown areas; all that went south left before hunting season. Of the 395 radiotagged pintails that wintered in CCA, 83% flew from the SJV north to other CCA areas (i.e., Sacramento Valley [SACV], Sacramento-San Joaquin River Delta [Delta], Suisun Marsh, San Francisco Bay) during September-January; most went during December. Movements coincid- ed with start of hunting seasons and were related to pintail age, mass, capture location, study year, and weather. Among pintails with less than average mass, AHY individuals tended to leave the SJV earlier than HY individuals. Weekly distribution was similar among capture locations and years but a greater percentage of pintails radiotagged in Tulare Basin (south part of SJV) were known to have (10.3% vs. 0.9%) or probably (13.8% vs. 4.6%) wintered south of CCA than pintails radiotagged in northern SJV areas (i.e., Grassland Ecological Area [EA] and Mendota Wildlife Area [WA]). Also, a greater percentage of SJV pintails went to other CCA areas before hunting season in the drought year of 1991-1992 than later years (10% vs. 3-5%). The percent of radiotagged pintails from Grass- land EA known to have gone south of CCA also was greater during 1991-1992 than later years (2% vs. 0%), but both the known (19% vs. 4%) and probable (23% vs. 12%) percent from Tulare Basin that went south was greatest during 1993-1994, when availability of flooded fields there was lowest. The probability of pintails leaving the SJV was 57% (95% CI = 8-127%) greater on days with than without rain, and more movements per bird out of SJV occurred in years with more rain and fog but fewer days with southerly winds. Movements by pintails and changes in pintail distributions, direct recovery distributions, and harvest rates suggest the disproportionate decline of pin- tails in Tulare Basin was due to a lower percentage of pintails moving there in fall and a greater percentage or ear- lier movements north and south out of Tulare Basin. With fewer in Tulare Basin to replace Grasslands EA pintails going north in December, pintail abundance in the northern SJV declined during late winter. Changes in move- ment patterns correspond to habitat loss in Tulare Basin and increased habitats in SACV and western mainland Mexico. Habitat improvements, especially in Tulare Basin, that increase food, sanctuary, and winter survival would probably help restore pintails throughout the SJV.

Journal of Wildlife Management↗

U.S. Geological Survey Karst Interest Group Proceedings, San Antonio, Texas, May 16–18, 2017

Introduction and Acknowledgments Karst aquifer systems are present throughout parts of the United States and some of its territories, and have developed in carbonate rocks (primarily limestone and dolomite) and evaporites (gypsum, anhydrite, and halite) that span an interval of time encompassing more than 550 million years. The depositional environments, diagenetic processes, post-depositional tectonic events, and geochemical weathering processes that form karst aquifers are varied and complex. These factors involve biological, chemical, and physical changes that when combined with the diverse climatic regimes in which karst development has taken place, result in the unique dual- or triple-porosity nature of karst aquifers. These complex hydrogeologic systems typically represent challenging and unique conditions to scientists attempting to study groundwater flow and contaminant transport in these terrains. The dissolution of carbonate rocks and the subsequent development of distinct and beautiful landscapes, caverns, and springs have resulted in the most exceptional karst areas being designated as national or state parks. Tens of thousands of similar areas in the United States have been developed into commercial caverns and known privately owned caves. Both public and private properties provide access for scientists to study the flow of groundwater in situ . Likewise, the range and complexity of landforms and groundwater flow systems associated with karst terrains are enormous, perhaps more than for any other aquifer type. Karst aquifers and landscapes that form in tropical areas, such as the cockpit karst along the north coast of Puerto Rico, differ greatly from karst landforms in more arid climates, such as the Edwards Plateau in west-central Texas or the Guadalupe Mountains near Carlsbad, New Mexico, where hypogenic processes have played a major role in speleogenesis. Many of these public and private lands also contain unique flora and fauna associated with these karst hydrogeologic systems. As a result, numerous federal, state, and local agencies have a strong interest in the study of karst terrains. Many of the major springs and aquifers in the United States have developed in carbonate rocks, such as the Floridan aquifer system in Florida and parts of Alabama, Georgia, and South Carolina; the Ozark Plateaus aquifer system in parts of Arkansas, Kansas, Missouri, and Oklahoma; and the Edwards-Trinity aquifer system in west-central Texas. These aquifers, and the springs that discharge from them, serve as major water-supply sources and form unique ecological habitats. Competition for the water resources of karst aquifers is common, and urban development and the lack of attenuation of contaminants in karst areas due to dissolution features that form direct pathways into karst aquifers can impact the ecosystem and water quality associated with these aquifers. The concept for developing a platform for interaction among scientists within the U.S. Geological Survey (USGS) working on karst-related studies evolved from the November 1999 National Groundwater Meeting of the USGS. As a result, the Karst Interest Group (KIG) was formed in 2000. The KIG is a loose-knit, grass-roots organization of USGS and non-USGS scientists and researchers devoted to fostering better communication among scientists working on, or interested in, karst science. The primary mission of the KIG is to encourage and support interdisciplinary collaboration and technology transfer among scientists working in karst areas. Additionally, the KIG encourages collaborative studies between the different mission areas of the USGS as well as with other federal and state agencies, and with researchers from academia and institutes. To accomplish its mission, the KIG has organized a series of workshops that have been held near nationally important karst areas. To date (2017) seven KIG workshops, including the workshop documented in this report, have been held. The workshops typically include oral and poster sessions on selected karst-related topics and research, as well as field trips to local karst areas. To increase non-USGS participation an effort was made for the workshops to be held at a university or institute beginning with the fourth workshop. Proceedings of the workshops are published by the USGS and are available online at the USGS publications warehouse https://pubs.er.usgs.gov/ by using the search term “karst interest group.” The first KIG workshop was held in St. Petersburg, Florida, in 2001, in the vicinity of the large springs and other karst features of the Floridan aquifer system. The second KIG workshop was held in 2002, in Shepherdstown, West Virginia, in proximity to the carbonate aquifers of the northern Shenandoah Valley, and highlighted an invited presentation on karst literature by the late Barry F. Beck of P.E. LaMoreaux and Associates. The third KIG workshop was held in 2005, in Rapid City, South Dakota, near evaporite karst features in limestones of the Madison Group in the Black Hills of South Dakota. The Rapid City KIG workshop included field trips to Wind Cave National Park and Jewel Cave National Monument, and featured a presentation by Thomas Casadevall, then USGS Central Region Director, on the status of Earth science at the USGS. The fourth KIG workshop in 2008 was hosted by the Hoffman Environmental Research Institute and Center for Cave and Karst Studies at Western Kentucky University in Bowling Green, Kentucky, near Mammoth Cave National Park and karst features of the Chester Upland and Pennyroyal Plateau. The workshop featured a late-night field trip into Mammoth Cave led by Rickard Toomey and Rick Olsen, National Park Service. The fifth KIG workshop in 2011 was a joint meeting of the USGS KIG and University of Arkansas HydroDays, hosted by the Department of Geosciences at the University of Arkansas in Fayetteville. The workshop featured an outstanding field trip to the unique karst terrain along the Buffalo National River in the southern Ozarks, and a keynote presentation on paleokarst in the United States was delivered by Art and Peggy Palmer. The sixth KIG workshop was hosted by the National Cave and Karst Research Institute (NCKRI) in 2014, in Carlsbad, New Mexico. George Veni, Director of the NCKRI, served as a co-chair of the workshop with Eve Kuniansky of the USGS. The workshop featured speaker Dr. Penelope Boston, Director of Cave and Karst Studies at New Mexico Tech, Socorro, and Academic Director at the NCKRI, who addressed the future of karst research. The field trip on evaporite karst of the lower Pecos Valley was led by Lewis Land (NCKRI karst hydrologist), and the field trip on the geology of Carlsbad Caverns National Park was led by George Veni. This current seventh KIG workshop is being held in San Antonio at the University of Texas at San Antonio (UTSA). This 2017 workshop is being hosted by the Department of Geological Sciences’ Student Geological Society (SGS), and student chapters of the American Association of Petroleum Geologists (AAPG) and Association of Engineering Geologists (AEG), with support by the UTSA Department of Geological Sciences and Center for Water Research. The UTSA student chapter presidents, Jose Silvestre (SGS), John Cooper (AAPG), and Tyler Mead (AEG) serve as co-chairs of the 2017 workshop with Eve Kuniansky of the USGS. The technical session committee is chaired by Eve Kuniansky, USGS, and includes Michael Bradley, Tom Byl, Rebecca Lambert, John Lane, and James Kaufmann, all USGS, and Patrick Tucci, retired USGS. The logistics committee includes Amy Clark, Yongli Gao, and Lance Lambert (Department Chair), UTSA Department of Geological Sciences; and Ryan Banta and Allan Clark, USGS, San Antonio, Texas. The field trip committee is chaired by Allan Clark and includes Amy Clark, Yongli Gao, and Keith Muehlestein, UTSA; Marcus Gary, Edwards Aquifer Authority and University of Texas at Austin; Ron Green, Southwest Research Institute; Geary Schindel, Edwards Aquifer Authority; and George Veni, NCKRI. Additionally, two organizations have assisted the UTSA student chapters in hosting the meeting by donating funds to the chapters: the Edwards Aquifer Authority, San Antonio, Texas, and the Barton Springs Edwards Aquifer Authority, Austin, Texas. Additionally, Yongli Gao, Center for Water Research and Department of Geological Sciences, UTSA, helped develop sessions on cave and karst research in China for this workshop. These proceedings could not have been accomplished without the assistance of Lawrence E. Spangler as co-editor who not only has subject matter expertise, but also serves as an editor with the USGS Science Publishing Network. We sincerely hope that this workshop continues to promote future collaboration among scientists of varied and diverse backgrounds, and improves our understanding of karst aquifer systems in the United States and its territories. The extended abstracts of USGS authors were peer reviewed and approved for publication by the USGS. Articles submitted by university researchers and other federal and state agencies did not go through the formal USGS peer review and approval process, and therefore may not adhere to USGS editorial standards or stratigraphic nomenclature. However, all articles had a minimum of two peer reviews and were edited for consistency of appearance in the proceedings. The use of trade, firm or product names is for descriptive purposes only and does not imply endorsement by the U.S. Government. The USGS Water Availability and Use Science Program funded the publication costs of the proceedings.

Scientific Investigations Report↗

Effects of Climate and land use on diversity, prevalence, and seasonal transmission of avian hematozoa in American Samoa

The indigenous forest birds of American Samoa are increasingly threatened by changing patterns of rainfall and temperature that are associated with climate change as well as environmental stressors associated with agricultural and urban development, invasive species, and new introductions of avian diseases and disease vectors. Long term changes in their distribution, diversity, and population sizes could have significant impacts on the ecological integrity of the islands because of their critical role as pollinators and seed dispersers. We documented diversity of vector borne parasites on Tutuila and Ta‘u Islands over a 10-year period to expand earlier observations of Plasmodium, Trypanosoma , and filarial parasites, to provide better parasite identifications, and to create a better baseline for detecting new parasite introductions. We also identified potential mosquito vectors of avian Plasmodium and Trypanosoma , determined whether land clearing and habitat alterations associated with subsistence farming within the National Park of American Samoa can influence parasite prevalence, and determined whether parasite prevalence is correlated with seasonal changes in rainfall, temperature and wind speed. Three taxonomically distinct lineages of Plasmodium were identified from mosquito vectors and forest birds based on partial sequence data from parasite mitochondrial genes. All three have been described from passerine and galliform birds in Australasia. Two lineages, SCEDEN01 and ORW1, had elongate gametocytes and large schizonts that were consistent with species of Plasmodium in the subgenus Giavannolaia, but were taxonomically distinct from known morphological species of Plasmodium based on a Bayesian phylogenetic analysis of a 478 bp region of the parasite cytochrome b gene. Both are candidates for description as new species. The third lineage (GALLUS02) was detected only in mosquito vectors on Tutuila and was similar in cytochrome b sequence to P. juxtanucleare , a pathogenic species of Plasmodium from chickens and other galliform birds from Australasia, Africa, and South America. Plasmodium relictum , the malarial parasite that has had such a devastating impact on Hawaiian forest birds, was not detected. We observed large, striated trypanosomes in avian hosts from both Tutuila and Ta‘u Islands that fell within the same taxonomic clade as T. corvi and T. culicavium based on 18S ribosomal DNA sequence. We also observed sheathed microfilariae with pointed tails that had some morphological similarities to microfilaria from species of Pelecitus, Struthiofilaria and Eulimdana , but identification will require recovery and examination of adult filarial worms from the connective tissue or body cavities of infected birds. We also observed one or more species of haemococcidians (Isospora, synonym = Atoxoplasma ) within circulating lymphocytes from multiple avian host species. Overall prevalence of Plasmodium was higher on Ta‘u (22%, 75/341) than Tutuila (9.2%, 27/294), with most infections occurring in Polynesian starlings, Samoan starlings, Wattled honeyeaters, and Cardinal honeyeaters. Prevalence was relatively constant from year to year and between seasons at individual study sites, but varied among study sites, with highest rates of infection in areas with agricultural activity at Faleasao (37.4%, 73/195, Ta‘u Island) and Amalau Valley (9.7%, 21/216, Tutuila Island). Prevalence in more remote areas of the National Park of American Samoa was lower, ranging from 1.4% (2/146) at Laufuti and Luatele on Ta‘u to 7.7% (6/78) at Olo Ridge on Tutuila. Similar trends were evident for infections with Trypanosoma and filarial worms. Overall prevalence was not influenced significantly by warmer, wet (summer) or cooler, dry (winter) season. We detected Plasmodium infections in Culex sitiens and C. quinquefasciatus through either salivary gland and midgut dissections or PCR amplification of parasite cytochrome b genes in pooled or individual samples of mosquitoes that were collected on Tutuila. Pooled or individual Aedes oceanicus, A. polynesiensis, A. tutuilae, A. upolensis, A. nocturnus, Aedes (Finlaya) (mixed pools of A. samoanus, A. oceanicus, A. tutuilae), Aedes (Stegomyia) (mixed pools of A. aegypti, A. upolensis, A. polynesiensis ), and C. annulirostris were negative for Plasmodium , but we detected infections with Trypanosoma through midgut and salivary gland dissections in a single C. sitiens from Amalau Valley, Tutuila and three A. oceanicus from Faleasao, Ta‘u. Two of the A. oceanicus from Faleasao amplified successfully with Trypanosoma primers, but sequences were distinctly different from those obtained from avian hosts. We found a strong association between land use and prevalence of mosquito-transmitted parasites on Ta‘u Island with odds of being infected more than 20 times greater in agricultural plots than more remote native forest. This relationship was evident on Tutuila Island but not statistically significant because of the close proximity of study sites and observed movement of birds between native forest and agricultural land. Our data support previous studies that have suggested that Plasmodium and other vector-borne parasites are part of the indigenous parasite fauna in American Samoa. Transmission dynamics appear to be affected by environmental changes associated with land use practices.

HI↗

Investigation of land cover within wetland complexes at Dixie Meadows, Churchill County, Nevada, from October 2015 to January 2022

The U.S. Geological Survey investigated land cover at subannual time steps within six wetland areas in Dixie Valley, Churchill County, Nevada, from October 2015 to January 2022. As requested by the U.S. Fish and Wildlife Service, we used aerial photography and satellite remote sensing data to map surface water and other land cover types within the wetland complexes. We identified five land cover classes using the green normalized difference vegetation index (gNDVI) and its inverse relationship to the normalized difference water index (NDWI) within three U.S. Department of Agriculture National Agriculture Imagery Program aerial images (acquired in 2015, 2017, and 2019) and 110 European Space Agency Sentinel-2 satellite images (acquired 2015–2022). The relative wetness of soil conditions within each land cover class is estimated by comparison to previously published observations of relative conductivity measured by 79 field-based sensors within the wetlands from 2019 to 2021. We mapped the areal coverage of the five land cover classes for approximately 385 acres (1,559,000 square meters [m²]) comprising six individual wetland complexes as well as a larger 1,298- acre (5,254,000-m 2 ) area of interest inclusive of the wetland complexes and adjacent landscape. Land cover of open water (Class 5) primarily within ponds at one of the wetland complexes comprised 8,333 m 2 , on average, of the wetland complexes. Land cover of mixed shallow surface water, saturated soil, and vegetation (Class 4) comprised 111,723 m 2 on average of the wetland complexes. Land cover of dense green vegetation canopy cover (Class 3) that often (46 percent of observations) had underlying surface water or saturated soil conditions comprised 592,522 m 2 on average of the wetland complexes. The remaining areas of the wetland complexes not mapped as these three land cover types (Classes 2 and 1) had sparse vegetation or bare soil cover and commonly (greater than or equal to 67 percent of observations) had dry soil conditions. The investigation of land cover detailed in this report could inform future efforts to map land cover more precisely via higher resolution remote sensing or ground-based surveying or could be incorporated with other environmental monitoring data to characterize habitat and hydrology of the wetland complexes at Dixie Meadows.

Nevada↗

The shallow stratigraphy and sand resources offshore of the Mississippi Barrier Islands

Coastal Mississippi is protected by a series of barrier islands ranging in length from 10-25 kilometers that are less than 2 kilometers wide. The majority of these islands comprise the Gulf Islands National Seashore (GUIS), an ecologically diverse shoreline that provides habitat for wildlife including migratory birds and endangered animals. The majority of GUIS is submerged, and aquatic environments include dynamic tidal inlets, ebb-tide deltas, and seagrass beds. The islands are in a state of decline, with land areas severely reduced during the past century by storms, sea-level rise, and human alteration. Morton (2008) estimates that since the mid-1800s up to 64 percent of island surface area has been lost. Heavy damage was inflicted in 2005 by Hurricane Katrina, which passed by as a Category 3 storm and battered the islands with winds of more than 160 kilometers per hour and a storm surge up to 9 meters. Since 2007, the U.S. Geological Survey (USGS), in collaboration with the National Park Service, has been mapping the seafloor and substrate around the islands as part of the USGS Northern Gulf of Mexico Ecosystem Change and Hazard Susceptibility project. The purpose of these investigations is to characterize the near-surface stratigraphy and identify the influence it may have on island evolution and fate. In 2009, this effort provided the basis for a collaborative effort with the U.S. Army Corps of Engineers (USACE) to expand the investigation outside of GUIS boundaries as part of the Mississippi Coastal Improvement Project (MsCIP). The MsCIP program consists of structural, nonstructural, and environmental project elements to restore portions of coastal Mississippi and GUIS affected by storm impact. The project includes the placement of sand along the islands, both on the present beaches and within the littoral zone, to mitigate shoreline erosion and breaching. This action requires the location and assessment of offshore sand or sediment deposits that can provide suitable material for shoreline renourishment. The geophysical and sample information collected by the USGS during geologic investigations provides this information. As part of the MsCIP program, in March 2010 the USGS mapped approximately 300 square kilometers of seafloor around GUIS. Interferometric swath bathymetry, sidescan sonar, and Chirp sub-bottom profiling were used to characterize seafloor elevations, texture, and the underlying stratigraphy. On the basis of this information, potential sediment resources were identified. The most promising offshore deposits for beach restoration include shoals, lowstand valley fill, tidal delta deposits, abandoned barrier deposits, and dredge spoil. Of these, lowstand valley fill deposits and dredge spoil are less desirable; lowstand deposits are buried under a 2- to 4-meter blanket of mud, and dredge spoil volume is small. A relict tidal delta and submerged shoals are the most desirable deposits; the tidal delta contains a large volume of material still exposed on the seafloor, and parts of submerged shoals have modest volume and thin mud cover.

Mississippi↗

2005 annual progress report: Elk and bison grazing ecology in the Great Sand Dunes complex of lands

In 2000 the U.S. Congress authorized the expansion of the former Great Sand Dunes National Monument by establishing a new Great Sand Dunes National Park and Preserve in its place, and establishing the Baca National Wildlife Refuge. The establishment of Great Sand Dunes National Park and Preserve and the new Baca National Wildlife Refuge in the San Luis Valley (SLV), Colorado was one of the most significant land conservation actions in the western U.S. in recent years. The action was a result of cooperation between the National Park Service (NPS), U.S. Fish and Wildlife Service (USFWS), Bureau of Land Management (BLM), U.S. Forest Service (USDA-FS), and The Nature Conservancy (TNC). The new national park, when fully implemented, will consist of 107,265 acres, the new national preserve 41,872 acres, and the new national wildlife refuge (USFWS lands) 92,180 acres (fig. 1). The area encompassed by this designation protects a number of natural wonders and features including a unique ecosystem of natural sand dunes, the entire watershed of surface and groundwaters that are necessary to preserve and recharge the dunes and adjacent wetlands, a unique stunted forest, and other valuable riparian vegetation communities that support a host of associated wildlife and bird species. When the National Park was initially established, there were concerns about overconcentrations and impacts on native plant communities of the unhunted segments of a large and possibly growing elk (Cervus elaphus) population. This led to the designation of the Preserve as a compromise solution, where the elk could be harvested. The Preserve Unit, however, will not address all the ungulate management challenges. In order to reduce the current elk population, harvests of elk may need to be aggressive. But aggressive special hunts of elk to achieve population reductions can result in elk avoidance of certain areas or elk seeking refuge in areas where they cannot be hunted, while removals of whole herd segments and abandonment or alterations of migration routes can occur (Smith and Robbins, 1994; Boyce and others, 1991). Elk may seek refuge from hunting in the newly expanded Park Unit and TNC lands where they might overconcentrate and impact unique vegetation communities. In these sites of refugia, or preferred loafing sites, elk and bison could accelerate a decline in woody riparian shrubs and trees. This decline may also be due to changes in hydrology, climatic, or dunal processes, but ungulate herbivory might exacerbate the effects of those processes. To address the questions and needs of local resource managers, a multi-agency research project was initiated in 2005 to study the ecology, forage relations, and habitat relations of elk and bison in the Great Sand Dunes–Sangre de Cristo–Baca complex of lands. Meetings and discussions of what this research should include were started in 2001 with representatives from NPS, USFWS, TNC, the Colorado Division of Wildlife (CDOW), and USDA-FS/BLM. The final study plan was successfully funded in 2004 with research scheduled to start in 2005. The research was designed to encompass three major study elements: (1) animal movements and population dynamics, (2) vegetation and nutrient effects from ungulate herbivory, and (3) development of ecological models, using empirical data collected from the first two components, that will include estimates of elk carrying capacity and management scenarios for resource managers.

Colorado↗

Species richness and relative abundance of breeding birds in forests of the Mississippi Alluvial Valley

In 1992, the Vicksburg Field Research Station of the National Wetlands Research Center initiated research on the ecology of migratory birds within forests of the Mississippi Alluvial Valley (MAV). The MAV was historically a nearly contiguous bottomland hardwood forest, however, only remnants remain. These remnants are fragmented and often influenced by drainage projects, silviculture, agriculture, and urban development. Our objectives are to assess species richness and relative abundance, and to relate these to the size, quality, and composition of forest stands. Species richness and relative abundance were estimated for 53 randomly selected forest sites using 1 to 8 point counts per site, depending on the size of the forest fragment. However, statistical comparisons among sites will be restricted to an equal number ofpoint counts within the sites being compared. Point counts, lasting five minutes, were conducted from 11 May to 29 June 1992, foltowing Ralph, Sauer, and Droege (Point Count Standards; memo dated 9 March 1992). Vegetation was measured at the first three points on each site using a modification of the methods employed by Martin and Roper (Condor 90: 5 1-57; 1988). During 252 counts, 7 1 species were encountered, but only 62 species were encountered within a 50-m radius of point center. The mean number of species encountered within 50 m of a point, was 7.3 (s.d. = 2.7) and the mean number of individuals was 11.2 (s.d. = 4.2). The mean number of species detected at any distance was 9.6 (s.d, = 2.8) and the mean number of individuals was 15.6 (s.d. = 7.9). The most frequently encountered warblers in the MAV were Prothonotary Warbler and Northern Parula. Rarely encountered warblers were American Redstart and Worm-eating Warbler. The genera, Quercus, Ulmus, Carya, and Celtis were each encountered at 80 or more of the 152 points at which vegetation was sampled. Species most frequentlyencountered were: sugarberry (Celtis laevagata), water hickory (Caqa aquatica), American elm (Ulmus arnericana), sweetgum (Liquidambar styraciflua) and willow oak (Quercus phellos) The mean basal area of all trees 10 cm diameter-at-breast height (dbh) was 28 m2 /ha (range 7-70). The mean canopy cover was 87 percent, mean canopy height was 20 m, ground cover was 60 percent, and vegetation density (2-7 m) was 47 percent. The most frequently encountered understory species were sugarberry, ash (Fraxinus spp.), maple (Acer spp.), and elm (Ulnrus spp.). A cooperative GIs effort among the U.S. Fish and Wildlife Service, the Nature Conservancy, and the University of Arkansas is currently classifying forested habitats within the MAV. This effort will provide information on stand size and topology which will be used in concert with our current data, and data from visits to additional forest stands in 1993, to assess the relationship between the size, quatity, and composition of forests within the MAV and their breeding bird community.

Book chapter↗

Hydrogeology and sources of water to select springs in Black Canyon, south of Hoover Dam, Lake Mead National Recreation Area, Nevada and Arizona

Springs in Black Canyon of the Colorado River, directly south of Hoover Dam in the Lake Mead National Recreation Area, Nevada and Arizona, are important hydrologic features that support a unique riparian ecosystem including habitat for endangered species. Rapid population growth in areas near and surrounding Black Canyon has caused concern among resource managers that such growth could affect the discharge from these springs. The U.S. Geological Survey studied the springs in Black Canyon between January 2008, and May 2014. The purposes of this study were to provide a baseline of discharge and hydrochemical data from selected springs in Black Canyon and to better understand the sources of water to the springs. Various hydrologic, hydrochemical, geochemical, and geologic data were collected and analyzed during this study. More than 100 hydrologic sites consisting of springs, seeps, pools, rivers, reservoirs, and wells were investigated, and measurements were taken at 75 of these sites. Water levels were measured or compiled for 42 wells and samples of water were collected from 36 unique sites and submitted for laboratory analyses of hydrochemical constituents. Measurements of discrete discharge were made from nine unique spring areas and four sites in Black Canyon were selected for continuous monitoring of discharge. Additionally, samples of rock near Hoover Dam were collected and analyzed to determine the age of spring deposits. Results of hydrochemical analyses indicate that discharge from springs in Black Canyon is from two sources: (1) Lake Mead, and (2) a local and (or) regional source. Discharge from springs closest to Hoover Dam contains a substantial percentage (>50 percent) of water from Lake Mead. This includes springs that are between Hoover Dam and Palm Tree Spring. Discharge from springs south of Palm Tree Spring contains a substantial percentage (>50 percent) of the water that is believed to come from a combination of other local and regional sources, although the exact location and nature of these sources is not clear. The unique hydrochemistry of some springs, such as Bighorn Sheep Spring and Latos Pool, suggests that little if any water discharging from these springs comes from Lake Mead. Geochronological results of spring deposits at several sites near Hoover Dam indicate that most deposits are young and likely formed after the construction of Hoover Dam. Several major faults, including the Salt Cedar Fault and the Palm Tree Fault, play an important role in the movement of groundwater. Groundwater may move along these faults and discharge where faults intersect volcanic breccias or fractured rock. Vertical movement of groundwater along faults is suggested as a mechanism for the introduction of heat energy present in groundwater from many of the springs. Groundwater altitudes in the study area indicate a potential for flow from Eldorado Valley to Black Canyon although current interpretations of the geology of this area do not favor such flow. If groundwater from Eldorado Valley discharges at springs in Black Canyon then the development of groundwater resources in Eldorado Valley could result in a decrease in discharge from the springs. Geology and structure indicate that it is not likely that groundwater can move between Detrital Valley and Black Canyon. Thus, the development of groundwater resources in Detrital Valley may not result in a decrease in discharge from springs in Black Canyon.

Arizona, Nevada↗

Of travertine and time: otolith chemistry and microstructure detect provenance and demography of endangered humpback chub in Grand Canyon, USA

We developed a geochemical atlas of the Colorado River in Grand Canyon and in its tributary, the Little Colorado River, and used it to identify provenance and habitat use by Federally Endangered humpback chub, Gila cypha. Carbon stable isotope ratios (δ 13 C) discriminate best between the two rivers, but fine scale analysis in otoliths requires rare, expensive instrumentation. We therefore correlated other tracers (SrSr, Ba, and Se in ratio to Ca) to δ 13 C that are easier to quantify in otoliths with other microchemical techniques. Although the Little Colorado River’s water chemistry varies with major storm events, at base flow or near base flow (conditions occurring 84% of the time in our study) its chemistry differs sufficiently from the mainstem to discriminate one from the other. Additionally, when fish egress from the natal Little Colorado River to the mainstem, they encounter cold water which causes the otolith daily growth increments to decrease in size markedly. Combining otolith growth increment analysis and microchemistry permitted estimation of size and age at first egress; size at first birthday was also estimated. Emigrants < 1 year old averaged 51.2 ± 4.4 (SE) days and 35.5 ± 3.6 mm at egress; older fish that had recruited to the population averaged 100 ± 7.8 days old and 51.0 ± 2.2 mm at egress, suggesting that larger, older emigrants recruit better. Back-calculated size at age 1 was unimodal and large (78.2 ± 3.3 mm) in Little Colorado caught fish but was bimodally distributed in Colorado mainstem caught fish (49.9 ± 3.6 and 79 ± 4.9 mm) suggesting that humpback chub can also rear in the mainstem. The study demonstrates the coupled usage of the two rivers by this fish and highlights the need to consider both rivers when making management decisions for humpback chub recovery.

Arizona↗

Hydrogeologic framework, ground-water geochemistry, and assessment of nitrogen yield from base flow in two agricultural watersheds, Kent County, Maryland

Hydrostratigraphic and geochemical data collected in two adjacent watersheds on the Delmarva Peninsula, in Kent County, Maryland, indicate that shallow subsurface stratigraphy is an important factor that affects the concentrations of nitrogen in ground water discharging as stream base flow. The flux of nitrogen from shallow aquifers can contribute substantially to the eutrophication of streams and estuaries, degrading water quality and aquatic habitats. The information presented in this report includes a hydrostratigraphic framework for the Locust Grove study area, analyses and interpretation of ground-water chemistry, and an analysis of nutrient yields from stream base flow. An understanding of the processes by which ground-water nitrogen discharges to streams is important for optimal management of nutrients in watersheds in which ground-water discharge is an appreciable percentage of total streamflow. The U.S. Geological Survey, in cooperation with the U.S. Environmental Protection Agency (USEPA), collected and analyzed hydrostratigraphic and geochemical data in support of ground-water flow modeling by the USEPA. The adjacent watersheds of Morgan Creek and Chesterville Branch have similar topography and land use; however, reported nitrogen concentrations are generally 6 to 10 milligrams per liter in Chesterville Branch but only 2 to 4 milligrams per liter in Morgan Creek. Ground water in the surficial aquifer in the recharge areas of both streams has high concentrations of nitrate (greater than 10 milligrams per liter as N) and dissolved oxygen. One component of the ground water discharging to Morgan Creek typically is anoxic and contains virtually no dissolved nitrate; most of the ground water discharging to Chesterville Branch is oxygenated and contains moderately high concentrations of nitrate. The surficial aquifer in the study area is composed of the deeply weathered sands and gravels of the Pensauken Formation (the Columbia aquifer) and the underlying glauconitic sands of the upper Aquia Formation (the Aquia aquifer). The lower 6 to 9 meters of the Aquia Formation is a low-permeability silt-clay with abundant glauconite. The Aquia confining layer underlies the Columbia-Aquia surficial aquifer throughout the study area. The sediment redox transition, identified in cores, that occurs in the upper 0.5 to 1 meter of the Aquia confining layer is thought to be a site for subsurface denitrification of ground water. The first confined aquifer is composed of the glauconitic sands in the upper 9 to 11 meters of the Hornerstown Formation. The Hornerstown aquifer is underlain by 10 to 15 meters of glauconitic silt-clay at the base of the Hornerstown Formation (the Hornerstown confining layer), and 5 meters of low-permeability clay in the underlying Severn Formation. The Aquia and Hornerstown Formations dip and thicken to the southeast, and the Aquia confining layer subcrops shallowly (within 5 meters of the land surface) in a band that strikes southwest to northeast across the northern edge of the study area. The surficial aquifer is very thin (generally less than 5 meters) north of Morgan Creek, and the alluvial valley of Morgan Creek has incised into the top of the Aquia confining layer. In contrast, the Aquia confining layer lies 22 meters below Chesterville Branch, and the surficial aquifer approaches 30 meters in thickness (away from the creek). Chemically reduced iron sulfides and glauconite in the Aquia confining layer are likely substrates for denitrification of nitrate in ground water. Evidence from the dissolved concentrations of nitrate, sulfate, iron, argon, and nitrogen gas, and stable nitrogen isotopes support the interpretation that ground water flowing near the top of the Aquia confining layer, or through the confined Hornerstown aquifer, has undergone denitrification. This process appears to have the greatest effect on ground-water chemistry north of Morgan Creek, where the surficial aquifer is thin and a greater percentage of the ground water contacts the Aquia confining layer. The base-flow discharges of total nitrogen from the two watersheds are of similar magnitude, although Chesterville Branch has somewhat higher loads (29,000 kilograms of nitrogen per year) than Morgan Creek (20,000 kilograms of nitrogen per year), although Morgan Creek has a larger drainage area and a greater discharge of water. The base-flow yield of nitrogen (load per unit area) in Chesterville Branch (median of 0.058 grams per second per square kilometer at the outlet) is more than twice that of Morgan Creek (median of 0.022 grams per second per square kilometer at the outlet), reflecting the higher concentration of nitrate in ground water discharging to Chesterville Branch. Total nitrogen concentrations tend to decrease downstream in Chesterville Branch and increase downstream in Morgan Creek. The downstream trend in Chesterville Branch may be affected by instream nitrogen uptake and denitrification, and an increasing proportion of older, denitrified ground water in downstream discharge. The downstream trends in Morgan Creek may be affected by inflow from tributaries, downstream changes in the source of discharge water, and downstream changes in the riparian zone, which could affect the processes and degree of denitrification. Although these two watersheds appear to have landscape features (such as topography, land use, and soils) that would produce similar nitrogen discharges, a more detailed examination of landscape features indicates that Chesterville Branch has soils that are slightly better drained, tributary stream outlets at higher altitudes, and a slightly higher percentage of agricultural land. All of these factors have been related to higher nitrogen yields. Nonetheless, most of the data support the interpretation that hydrostratigraphy has the greatest effect in producing the difference in nitrogen yields between the two watersheds.

Maryland↗

Preliminary volcano-hazard assessment for the Katmai volcanic cluster, Alaska

The world’s largest volcanic eruption of the 20th century broke out at Novarupta (fig. 1) in June 1912, filling with hot ash what came to be called the Valley of Ten Thousand Smokes and spreading downwind more fallout than all other historical Alaskan eruptions combined. Although almost all the magma vented at Novarupta, most of it had been stored beneath Mount Katmai 10 km away, which collapsed during the eruption. Airborne ash from the 3-day event blanketed all of southern Alaska, and its gritty fallout was reported as far away as Dawson, Ketchikan, and Puget Sound (fig. 21). Volcanic dust and sulfurous aerosol were detected within days over Wisconsin and Virginia; within 2 weeks over California, Europe, and North Africa; and in latter-day ice cores recently drilled on the Greenland ice cap. There were no aircraft in Alaska in 1912—fortunately! Corrosive acid aerosols damage aircraft, and ingestion of volcanic ash can cause abrupt jet-engine failure. Today, more than 200 flights a day transport 20,000 people and a fortune in cargo within range of dozens of restless volcanoes in the North Pacific. Air routes from the Far East to Europe and North America pass over and near Alaska, many flights refueling in Anchorage. Had this been so in 1912, every airport from Dillingham to Dawson and from Fairbanks to Seattle would have been enveloped in ash, leaving pilots no safe option but to turn back or find refuge at an Aleutian airstrip west of the ash cloud. Downwind dust and aerosol could have disrupted air traffic anywhere within a broad swath across Canada and the Midwest, perhaps even to the Atlantic coast. The great eruption of 1912 focused scientific attention on Novarupta, and subsequent research there has taught us much about the processes and hazards associated with such large explosive events (Fierstein and Hildreth, 1992). Moreover, work in the last decade has identified no fewer than 20 discrete volcanic vents within 15 km of Novarupta (Hildreth and others, 1999, 2000, 2001; Hildreth and Fierstein, 2000), only half of which had been named previously—the four stratovolcanoes Mounts Katmai, Mageik, Martin, and Griggs; the cone cluster called Trident Volcano; Snowy Mountain; and the three lava domes Novarupta, Mount Cerberus, and Falling Mountain. The most recent eruptions were from Trident Volcano (1953–74), but there have been at least eight other, probably larger, explosive events from the volcanoes of this area in the past 10,000 years. This report summarizes what has been learned about the volcanic histories and styles of eruption of all these volcanoes. Many large earthquakes occurred before and during the 1912 eruption, and the cluster of Katmai volcanoes remains seismically active. Because we expect an increase in seismicity before eruptions, seismic monitoring efforts to detect volcanic unrest and procedures for eruption notification and dissemination of information are included in this report. Most at risk from future eruptions of the Katmai volcanic cluster are (1) air-traffic corridors of the North Pacific, including those approaching Anchorage, one of the Pacific’s busiest international airports, (2) several regional airports and military air bases, (3) fisheries and navigation on the Naknek Lake system and Shelikof Strait, (4) pristine wildlife habitat, particularly that of the Alaskan brown bear, and (5) tourist facilities in and near Katmai National Park.

Alaska↗

Quantifying Ground-Water and Surface-Water Discharge from Evapotranspiration Processes in 12 Hydrographic Areas of the Colorado Regional Ground-Water Flow System, Nevada, Utah, and Arizona

Rapid population growth in southern Nevada has increased the demand for additional water supplies from rural areas of northern Clark and southern Lincoln counties to meet projected water-supply needs. Springs and rivers in these undeveloped areas sustain fragile riparian habitat and may be susceptible to ground-water withdrawals. Most natural ground-water and surface-water discharge from these basins occurs by evapotranspiration (ET) along narrow riparian corridors that encompassed about 45,000 acres or about 1 percent of the study area. This report presents estimates of ground- and surface-water discharge from ET across 3.5 million acres in 12 hydrographic areas of the Colorado Regional Ground-Water Flow System. Ground-and surface-water discharge from ET were determined by identifying areas of ground- and surface-water ET, delineating areas of similar vegetation and soil conditions (ET units), and computing ET rates for each of these ET units. Eight ET units were identified using spectral-reflectance characteristics determined from 2003 satellite imagery, high-resolution aerial photography, and land classification cover. These ET units are dense meadowland vegetation (200 acres), dense woodland vegetation (7,200 acres), moderate woodland vegetation (6,100 acres), dense shrubland vegetation (5,800 acres), moderate shrubland vegetation (22,600 acres), agricultural fields (3,100 acres), non-phreatophytic areas (3,400,000 acres), and open water (300 acres). ET from diffuse ground-water and channelized surface-water is expressed as ETgs and is equal to the difference between total annual ET and precipitation. Total annual ET rates were calculated by the Bowen ratio and eddy covariance methods using micrometeorological data collected from four sites and estimated at 3.9 ft at a dense woodland site (February 2003 to March 2005), 3.6 ft at a moderate woodland site (July 2003 to October 2006), 2.8 ft at a dense shrubland site (June 2005 to October 2006), and 1.5 ft at a moderate shrubland site (April 2006 to October 2006). Annual ETgs rates were 3.4 ft for dense woodland vegetation, 3.2 ft for moderate woodland vegetation, 2.2 ft for dense shrubland vegetation, and 1.0 ft for moderate shrubland vegetation. Published annual rates of ETgs were used for the other ET units found in the study area. These rates were 3.4 ft for dense meadowland vegetation, 5.2 ft for agricultural fields, and 4.9 ft for open water. For the non-phreatophytic ET unit, ETgs was assumed to be zero. Estimated ground- and surface-water discharge from ET was calculated by multiplying the ETgs by the ET-unit acreage and equaled 24,480 acre-ft for dense woodland vegetation, 19,520 acre-ft for moderate woodland vegetation, 12,760 acre-ft for dense shrubland vegetation, 22,600 acre-ft for moderate shrubland vegetation, 680 acre-ft for dense meadowland vegetation, 16,120 acre-ft for agricultural fields, 1,440 acre-ft for open water, and 0 acre-ft for the non-phreatophytic ET unit. Estimated ground-water and surface-water discharge from ET from each hydrographic area was calculated by summing the total annual ETgs rate for ET units found within each hydrographic area and equaled 1,952 acre-ft for the Black Mountains Area, 6,080 acre-ft for California Wash, 4,090 acre-ft for the Muddy River Springs Area, 11,510 acre-ft for Lower Moapa Valley, 51,960 acre-ft for the Virgin River Valley, 16,168 acre-ft for Lower Meadow Valley Wash, 5,840 acre-ft for Clover Valley, and 0 acre-ft for Coyote Spring Valley, Kane Springs Valley, Tule Desert, Hidden Valley (North), and Garnet Valley. The annual discharge from ETgs for the study area totals about 98,000 acre-ft.

Scientific Investigations Report↗

Reservoir evolution, downstream sediment transport, downstream channel change, and synthesis of geomorphic responses of Fall Creek and Middle Fork Willamette River to water years 2012–18 streambed drawdowns at Fall Creek Lake, Oregon

Executive Summary Chapter A. Introduction Fall Creek Dam impounds Fall Creek Lake, a 10-kilometer-long reservoir in western Oregon and is operated by the U.S. Army Corps of Engineers (USACE) primarily for flood-risk management (or flood control) in late autumn through early spring months, as well as for water quality, irrigation, recreation, and habitat in late spring through early autumn. Since 2011 (water year [WY] 2012), Fall Creek Lake has been temporarily drawn down each year to facilitate downstream passage of juvenile spring Chinook salmon ( Oncorhynchus tshawytscha ) through the 55-meter (m) high dam. This annual dam operation is temporary, typically lasting about 1–2 weeks from WY 2012 through 2020 (drawdown operations in WY 2022–24 have increased to more than 6 weeks). Drawdown of the reservoir results in lake levels being lowered to the elevation near the historical, pre-dam streambed. The annual streambed drawdowns of WY 2012–18 have improved fish passage and led the USACE to formally adopt streambed drawdowns as part of annual operations at Fall Creek Dam. However, temporarily lowering the lake to streambed creates free-flowing conditions in the reservoir that result in the erosion and episodic export of predominantly sand and finer-grained sediments (less than 2 millimeters [mm]) to the lower gravel-bed reaches of Fall Creek and the Middle Fork Willamette River. The introduction of large volumes of sand and finer-grain sediment into the dam-regulated reaches downstream from Fall Creek Dam prompted questions about the geomorphic responses to annual streambed drawdowns within Fall Creek Lake and downstream reaches along Fall Creek and the Middle Fork Willamette River. The U.S. Geological Survey (USGS) in partnership with USACE initiated a comprehensive geomorphic and sediment transport investigation to assess the coupled processes of reservoir erosion, sediment evacuation from Fall Creek Lake, and patterns of sediment transport and deposition in reaches downstream from the Fall Creek Dam that have resulted from annual streambed drawdowns. The purpose of this report is to systematically describe the processes of sediment erosion, transport, and deposition at Fall Creek Lake and geomorphic interactions between reaches upstream and downstream from Fall Creek Dam that relate to dam operations. Specifically, this report focuses on evaluating geomorphic responses to streambed drawdowns from WY 2012 through 2018 and placing drawdown-induced geomorphic responses within the broader context of physiographic and historical conditions and dam operations of Fall Creek and Middle Fork Willamette Rivers. Key objectives for this study were to characterize changes in reservoir morphology and substrate at Fall Creek Lake, describe the character and temporal pattern of sediment transport downstream from Fall Creek Dam, characterize geomorphic changes in channel reaches downstream from the Fall Creek Dam, and relate these data to the annual streambed drawdowns of WY 2012–18. This study uses multiple independent monitoring and measurement approaches to assess site, reach, and river-scale geomorphic responses to drawdowns to inform dam and reservoir management. Patterns and processes of reservoir evolution were assessed with geomorphic mapping and volumetric analyses of topography through comparison of multiple digital surface models (DSMs). Just downstream from Fall Creek Dam, analyses of sediment export from the reservoir focused on suspended sediment but also incorporated bedload analyses to assess sediment sizes. Geomorphic assessments downstream from the dam used reach-scale and site-scale approaches to document changes in channel morphology and substrate, including site measurements of sand and finer-grained sediment deposition and in-channel bed-material, volumetric change analyses from comparison of digital elevation models (DEMs), and repeat geomorphic mapping. Findings from this study inform river management and dam operations by providing an understanding of (1) coupled upstream-downstream geomorphic responses to the Fall Creek Lake streambed drawdowns, (2) geomorphic responses of Fall Creek Lake streambed drawdowns in comparison to drawdowns at other large dams, (3) controls on reservoir erosion and downstream geomorphic responses, and (4) implications for future hydrogeomorphic changes that may result from continued drawdowns and monitoring activities to assess those changes. Chapter B. Reservoir Morphology and Evolution Related to Dam Operations at Fall Creek Lake To understand the volume and distribution of sediment accumulation in Fall Creek Lake since dam closure in 1965, decadal-scale sedimentation patterns (spanning approximately 1965–2016) are evaluated using a combination of storage curve analyses and geomorphic mapping. Short-term (drawdown event-scale) patterns of erosion, sedimentation, and sediment export downstream are evaluated using a combination of geomorphic mapping and change detection analyses that quantify the distribution and total volume of sediment erosion and deposition within Fall Creek Lake. Geomorphic mapping of reservoir topography and analyses of historical datasets reveals four categories of landforms and sediment processes within Fall Creek Lake related to lake level operations: lacustrine sedimentation expressed in the reservoir floor, fluvial erosion and deposition within historical stream channels during streambed drawdowns, channel-like features created by erosion within the reservoir floor during streambed drawdowns, and erosion on reservoir hillslopes. Where the reservoir floor is mapped for this study as pelagic (deep water), deposition up to 3 meters (m) thick by lacustrine processes and burial of pre-dam topography with deposits thinning toward the edges of the valley floor and upstream areas of reservoir are observed. Despite over 50 years of sediment accumulation since dam construction, the main stream channels of Fall and Winberry Creeks (or reservoir thalwegs) through the reservoir are well defined, though their distinct morphology is likely influenced by a long history of recurring historical drawdowns to or near streambed since dam construction. Unregulated streamflow and sediment transport through the reservoir primarily are confined to these channels during the streambed drawdown periods. Erosional channel-like features created by drawdowns are carved through underlying, unconsolidated reservoir floor sediments and are most prominent in the lower reservoir below minimum conservation pool (the low pool elevation during winter flood season); sediment generated from the formation of these drawdown channels is more likely to be transported through and out of the reservoir than sediment deposits along the reservoir hillslopes at the valley margins that are separated from main channels by areas of low-gradient reservoir floor. Morphologic changes in the lower reservoir topography between January 2012 and November 2016 indicate overall net erosion of about 129,500 cubic meters (m 3 ). The most prominent geomorphic changes occurred along the main channels of Fall and Winberry Creeks near the Fall Creek Dam where incision, lateral migration, and slumping banks resulted in vertical and lateral adjustments to channel position, whereas most changes fell below the detectable limit on higher-elevation reservoir floor surfaces except where erosion occurred along features mapped as drawdown channels. Chapter C. Sediment Delivery from Fall Creek Lake and Transport through Downstream Reaches USGS implemented a sediment monitoring program in WY 2013–18 to evaluate the quantity and character of reservoir sediment exported from Fall Creek Lake during streambed drawdowns. Turbidity and suspended sediments were monitored annually autumn through spring to span the WY 2013–18 streambed drawdowns; however, unequal monitoring timeframes each year reduced the ability to compare results and factors affecting sediment export from the reservoir difficult between years. These data were originally measured to develop regressions and compute suspended-sediment loads (SSL). Bedload sediment monitoring from a cableway at the Fall Creek streamgage was completed in the autumn-winter of WY 2013 and 2017. The limited number of samples and presumed variability in sediment supply from the reservoir precluded construction of streamflow and bedload discharge relations to compute more than instantaneous bedload. Sand and finer-grained silts and clays were transported from the reservoir in suspension, though some coarser grains (up to 32 mm) were also mobilized and transported downstream from the dam as bedload. Observations of increased sediment transport downstream from Fall Creek Dam coincided with lake levels approaching about 3 m (10 feet [ft] or elevation 690 ft) above the streambed regulating outlets. Suspended-sediment loads computed for the full monitoring periods WY 2013–18 at the Fall Creek streamgage, located 1.4 kilometers (km) downstream from Fall Creek Dam, range from 54,700 metric tons (t) in WY 2013 to 13,900 t in WY 2018. Although the total annual SSL varied from year to year, the overall seasonal patterns of suspended sediment transport throughout each year were similar during monitoring in WY 2013-18. Suspended-sediment loads were low prior to the drawdown, then increased rapidly as lake levels lowered and approached the streambed. In the weeks following the drawdown period, as pool levels were increased, SSL remained slightly elevated above pre-drawdown levels but generally declined through the following winter and spring except during streamflow-driven pulses of suspended-sediment transport. WY 2013 had the greatest total computed SSL for each streambed drawdown and partial-year monitoring period. SSL computed for the partial-year period have generally decreased since WY 2013 and have varied by about 6,800 t with the exception of WY 2014. WY 2014 SSL reflects anomalously low sediment export due to low streamflows and freezing conditions that stabilized reservoir floor deposits. Bedload measurements in the short 1.4-km reach between Fall Creek Dam and the Fall Creek streamgage showed an inverse correlation between bedload transport rates and discharge, which probably reflects diminishing supply of coarse-sized sediment. Sand was more abundant (60–100 percent) than gravel in bedload samples confirming sand and finer-grained sediment dominated sediment evacuated from the reservoir during streambed drawdowns at Fall Creek Lake. Chapter D. Geomorphic Responses to Fall Creek Lake Streambed Drawdowns Downstream from Fall Creek Dam In the days, weeks, and months following streambed drawdown operations at Fall Creek Dam through WY 2018, sites downstream from the dam displayed a variety of geomorphic responses to reservoir sediment delivery within the main channel and overbank areas. Evaluation of streambed elevations at two streamgages located 1.4 km downstream from the dam on Fall Creek and 16.3 km downstream from the dam on the Middle Fork Willamette River indicated the effects of drawdown sediment on bed elevations were modest and transient. Repeat particle size measurements (October 2015 and September 2016) at five sites along Fall Creek and the Middle Fork Willamette River showed similar grain-sized distributions that do not reveal substantial deposition of fine-grained sediment related to the WY 2016 streambed drawdown. Altogether, these findings indicate that transport capacity in the main, low-flow channels of Fall Creek and Middle Fork Willamette River during WY 2012–18 was sufficient to mobilize and evacuate reservoir sediments from streambed drawdowns or other bank material and tributary sources. However, other monitoring for this study indicate low-velocity zones in off-channel areas are prime locations for sand and finer-grain sediment deposition. Patterns of overbank sediment accumulation indicate that the magnitude and timing of overbank deposition on bars and low-elevation floodplain varies with proximity to the dam, geomorphic setting, streamflows, and other factors. Sand and finer-grained reservoir sediments carried as suspended-sediment load in the reaches downstream from Fall Creek Dam were deposited in overbank areas as observed with clay-horizon markers during WY 2016–17. Overbank deposition quantified with Geomorphic Change Detection (GCD) software evaluated landform-scale patterns of erosion and deposition using repeat light detection and ranging (lidar) surveys at two sites in the Upper Fall Creek reach and one site in the Jasper reach for 3 years (2012–15) and one site in the Clearwater reach for 6 years (2009–15). Deposition thickness and spatial patterns from the GCD analysis were variable; some sites had dispersed but measurable deposition while at others, deposition was highly localized and exceeded 1 m in depth. Patterns of overbank deposition illustrate interactions among bar morphology, local hydraulics, and suspended-sediment transport dynamics that can create patches of highly localized deposition. The measured deposition at the two Fall Creek GCD sites likely resulted from reservoir sediments released from Fall Creek Lake during streambed drawdowns in WY 2016 and 2017 because the limited sediment inputs from bank material (geomorphically laterally stable reach) or tributaries (no significant tributaries) provided few other sediment sources. On the Middle Fork Willamette River, observed patterns of overbank deposition could reflect sediment sourced from upstream tributaries, bank erosion, or Fall Creek Lake streambed drawdown operations. Despite the introduction of several thousand tons of reservoir sediment delivered from the Fall Creek Lake streambed drawdowns to below-dam river corridors, reach-scale mapping of channel features downstream from Fall Creek Dam shows minimal evidence of changes in channel planform or landforms that can be attributed to a drawdowns in WY 2012–16. On Upper Fall Creek reach, widespread increases in gravel bars or other in-channel sediment did not result from the five streambed drawdowns. The main changes attributable to sediment releases from Fall Creek Lake were localized increases in vegetated bar area, particularly on channel margin areas where sand and finer-grain sediment was deposited and rapidly colonized by vegetation. The area of mapped secondary water features decreased between 2005 and 2016, but that may be due to lower discharges depicted in the 2016 aerial photographs and less mapped area of inundation. Primary changes along the Lower Fall Creek reach include a 6.4 percent decrease in area of secondary water features between 2011 and 2016, and a nearly twofold increase in the area of unvegetated bars. Immediately downstream from the Fall Creek confluence, there were negligible changes in the location and areas of vegetated bars and the main wetted channel between 2005 and 2016, and local increases in bar area cannot be attributed solely to deposition of reservoir sediments from Fall Creek Lake because (1) areas along the Middle Fork Willamette River just upstream from the Fall Creek confluence display similar type and magnitude of changes and (2) some of the increases at the confluence area pre-date the drawdowns. The cumulative effect of sediment releases from Fall Creek Lake streambed drawdowns from WY 2012 to 2016 on downstream channel planform and landforms are modest compared to the river-scale transformations and planform changes that occurred in the decades following dam construction. Chapter E. Discussion of Geomorphic Responses of Fall Creek and Middle Fork Willamette River to Streambed Drawdowns at Fall Creek Lake Multiple aspects of Fall Creek Dam infrastructure and operations exert first-order controls on the magnitudes of reservoir erosion that occur during the streambed drawdowns and ultimately determine the sediment delivery to downstream reaches. Key aspects of the dam and its operations that are most relevant to assessing geomorphic responses to streambed drawdowns include the (1) dam infrastructure, including configuration and size of regulating outlets and their proximity to the streambed which dictates the capacity and competence of the river to deliver sediment to downstream reaches and mode of sediment transport as suspended-sediment load or bedload; (2) frequency of historical drawdowns and long-term, year-round dam operations and lake level management, which partly dictate reservoir morphology and locations and magnitudes of readily erodible materials; (3) dam operations and hydroclimatic conditions during the streambed drawdown (including length of the drawdown and streamflows entering the reservoir), which directly control the timing, duration and magnitude of reservoir erosion and sediment evacuation; and (4) dam operations following the streambed drawdown operation that regulate streamflows (and thereby sediment transport conditions) downstream of Fall Creek Dam which primarily reflect interactions between hydroclimatic conditions and flood control operations. Patterns of sediment erosion and evacuation observed in this study at Fall Creek Lake from WY 2012–18 suggest that reservoir erosion during annual streambed drawdowns may remain similar or decrease in future years assuming (1) annual streambed drawdown operations are implemented in similar manner as the WY 2012–18 drawdowns (in terms of duration, late autumn or early winter implementation, rate of pool-level lowering to reach streambed, and other factors), (2) streambed drawdowns coincide with similar conditions as were observed WY 2012–18 (similar sediment yield into reservoir, low reservoir inflows, limited precipitation, moderate air temperature), and (3) no major geomorphic changes in the main reservoir channels of Fall and Winberry Creeks occur (for example, channel avulsion). Under such conditions, it is hypothesized that the stream channel within the reservoir would achieve a quasi-equilibrium state with respect to annual influx and export of sediment and aided by the substantial amount of in-channel bedrock, will remain laterally stable without erosion across reservoir deposits. Patterns of sediment transport measured at the Fall Creek streamgage downstream from Fall Creek Dam provide insight into the potential effects of future streambed drawdowns at Fall Creek Lake. Analyses of suspended sediment measured in WY 2013–18 show a major reduction in suspended-sediment loads between WY 2013 and later years, indicating streamflows transporting sediment through the reservoir to downstream reaches during streambed drawdowns have become supply limited. The 6-year suspended-sediment monitoring and sampling program is insufficient to make predictions about future sediment transport conditions because of uneven monitoring periods and varying controls on reservoir sediment erosion. It is likely that future suspended-sediment loads will be variable but similar to those observed in WY 2015–18 if operational, climatic, and geomorphological factors remain similar to those monitored WY 2015–18. Suspended-sediment loads downstream from Fall Creek Lake will likely remain highest when regulating outlets are fully open and Fall Creek is free flowing with the reservoir fully drained with little to no residual pool. Over time, it is possible that the suspended-sediment loads would reflect mobilization of reservoir sediment deposited in the previous year rather than erosion of sediment deposited years or decades earlier. Bedload is likely to remain a small fraction of the total sediment load evacuated from the reservoir and is relatively modest compared with pre-dam bedload transport rates because most coarse sediment remains trapped by the dam. If sediment releases from Fall Creek Lake and ensuing streamflow conditions follow a similar pattern in the future as was assessed in this study spanning WY 2012–18, near-term geomorphic adjustments downstream of the dam are expected to be modest. Barring major operational, climatic, and geomorphological changes, local site-scale deposition on bars, overbank areas, or off-channel features that persists several months after the streambed drawdown will likely continue to be highly variable, ranging from negligible to several centimeters of deposition. At the landform-scale, low velocity areas nearest to Fall Creek Dam will likely continue to undergo rapid deposition immediately during and after a streambed drawdown event, similar to patterns observed for WY 2012–18. Some of the sediment entering these off-channel features and margin areas may be temporarily stored, then later remobilized and dispersed farther downstream. But if newly deposited sediment persists through the following spring, there is a greater likelihood that local vegetation will establish, reinforce deposited material, and trap sediment during later drawdowns. The reach-scale geomorphic changes may become more apparent if (1) streambed drawdowns continued for several decades, and geomorphic changes were measured at decadal scales or (2) the amount of sediment introduced to downstream reaches substantially increased and (or) sediment transport capacity decreased. The continued streamflow regulation of Fall Creek Dam after sediment releases provides an opportunity to strategically manage streamflows during and after the streambed drawdowns to minimize downstream sediment impacts and ensure other operational thresholds are satisfied. This study provides a comprehensive foundation of datasets and geomorphic analyses to inform dam operations at Fall Creek Lake, monitor sediment transport downstream, and consider operational schemes for future drawdowns. The datasets from this study also provide baselines of sediment transport and geomorphic conditions to assess future changes in reservoir and downstream environments. Future monitoring could be tailored to address specific questions regarding the long-term geomorphic effects of streambed drawdowns on fluvial habitats, flood hazards, cultural resources, or downstream water quality. Future monitoring activities could focus on the relevant geomorphic processes and spatial domains within the three categories used for this study: (1) reservoir erosion and net sediment evacuation, (2) sediment delivery to downstream reaches, including magnitude and temporal pattern of sediment transport, and (3) geomorphic responses of downstream reaches to sediment delivery. Specifically, high priority future monitoring activities could include: Repeat topographic or photographic surveys in the reservoir to characterize changes occurring within individual drawdowns, to quantify sediment export, to determine temporal changes in reservoir storage, and to identify locations of erosion and deposition. Continuous, year-round turbidity monitoring supplemented with suspended-sediment measurements at a streamflow-gaging station immediately downstream from the dam to quantify sediment export. Repeat geomorphic monitoring, mapping, or modeling in downstream reaches to track changes in channel and over bank features using a combination of site- and reach-scale monitoring approaches. This could support assessments of sediment deposition and ensuing vegetation encroachment on flood hazards and habitats and examine how sediment transport and depositional processes may be affected by different sediment supply, streamflow, or dam management scenarios.

Oregon↗

Distribution and demography of Coastal Cactus Wrens in Southern California, 2015–19

Surveys and monitoring for the coastal Cactus Wren ( Campylorhynchus brunneicapillus ) were completed in San Diego County between March 2015 and July 2019. A total of 383 plots were surveyed across 3 genetic clusters (Otay, Lake Jennings, and Sweetwater/Encanto). From 2015 to 2019, 317 plots were surveyed 8 times (twice per year in 2015, 2017–19). Additional plots were added in later years as wrens were discovered in new locations. We found differences in the proportion of plots occupied in the genetic clusters, with a lower proportion of plots occupied in the Otay cluster than in the Lake Jennings and Sweetwater/Encanto clusters in all years. Plot occupancy increased each year in the Otay and Sweetwater/Encanto clusters but not in the Lake Jennings cluster. The number of Cactus Wren territories increased from 2015 through 2018, and then decreased in 2019 in all three genetic clusters. We monitored nesting activities for two populations of Cactus Wrens in southern San Diego County. The Otay population consisted of two sites within the Otay genetic cluster, and the San Diego population consisted of two sites within the Sweetwater/Encanto and Lake Jennings genetic clusters. Nest monitoring occurred at 10–13 territories per year in the Otay population and 14–18 territories in the San Diego population from 2015 through 2019. All territories were occupied by pairs except two territories in 2015, five in 2016, and two in 2019. Between 46 and 74 Cactus Wren nests were monitored each year, which totaled 295 monitored nests from 2015 to 2019. To evaluate the direct influence of precipitation on breeding success, bio-year precipitation (“precipitation”) was calculated from July 1 of the prior year through June 30 of the breeding season year. Overall apparent nest success was positively influenced by precipitation with the lowest apparent nest success of 50 percent in 2015 and the highest apparent nest success of 72 percent in 2017, corresponding to the second lowest and the highest precipitation years, respectively. Apparent nest success also was higher in the Otay population than in the San Diego population. The number of brood nests initiated per pair and the number of renesting attempts per pair also were higher in years with more precipitation. Other metrics of Cactus Wren nesting success and productivity were positively influenced by the amount of precipitation, including clutch size and egg hatching success. The percent of hatchlings that fledged was greater in the Otay population than in the San Diego population but was not influenced by precipitation. The number of fledglings per pair was higher in years with more precipitation and was greater in the Otay population than in the San Diego population. Predation was the predominant cause of nest failure in both populations. Analysis of Cactus Wren daily nest survival rate indicated that there was a population, and possibly a precipitation effect on nest survival, with the daily survival rate for the Otay population significantly higher than for the San Diego population and weak increase in the daily survival rate with more precipitation. A total of 629 Cactus Wrens were banded during the course of the study, 360 in the San Diego population and 269 in the Otay population. Between 2015 and 2019, we resighted 301 color-banded adult birds that ranged between 1 and 8 years old. One additional color-banded bird was resighted in San Pasqual Valley (as part of a separate study); this bird originated in the San Diego population and was excluded from our analyses. Annual survival was higher for adult Cactus Wrens (ranging from 60 to 70 percent) than for first-year wrens (ranging from 20 to 28 percent) and varied by year. Annual survival was also weakly but positively correlated with precipitation. Annual survival was higher for first year and adult Cactus Wrens following years with increased precipitation. We found no evidence that survival differed by population. Banding also allowed us to examine whether there were differences in movement of adult and first-year Cactus Wrens by year or by population. We found that average dispersal distance for first-year Cactus Wrens was 1.9 kilometers in the Otay population and 1.6 kilometers in the San Diego population and did not differ by population or year. Dispersal between populations was not common. We detected five instances of movement of first-year wrens between the San Diego and Otay populations. All movements into and out of the San Diego population were from or into territories in the Sweetwater area. We detected no movement between the Lake Jennings site and either of the Sweetwater or Otay sites; however, we did detect one wren that dispersed from Lake Jennings to the San Pasqual Valley population in 2019, which was a distance of 26.4 kilometers. Adult Cactus Wrens were site-faithful, with 87 percent of adults remaining on the same territory between breeding seasons. Precipitation may be a weak driver of movement for adult Cactus Wrens, with adults more likely to remain on the same territory following years of increased precipitation. There was no difference in adult movement between populations. Arthropods were collected in pitfall traps and by vacuum in 23 Cactus Wren territories during 3 sampling periods in 2016 (early nesting, peak nesting, and late nesting). Arthropods of 19 orders and at least 128 families were collected. Analysis of 43 Cactus Wren fecal samples identified 10 arthropod orders that were present in more than 10 percent of fecal samples. The most abundant arthropod order collected was Hymenoptera; however, Cactus Wrens consumed arthropods in the order Hymenoptera significantly less than their availability, suggesting that this order was avoided. No other orders were significantly selected or avoided; however, selection indices of arthropod families identified that two families of arthropods (Isopoda Porcellionidae [woodlice] and Hymenoptera Formicidae [ants]) were avoided. After excluding the taxa that were avoided or not represented in fecal samples, 95 percent of Cactus Wren prey items were collected in pitfall traps and 5 percent were collected by vacuum. The most abundant prey orders captured were Diptera, Coleoptera, Hemiptera, Hymenoptera, and Aranea. Analysis of the abundance of Cactus Wren prey items by vegetation type and sampling period indicated that vegetation type by itself was not a significant predictor of arthropod abundance but interacted with sampling period. Seasonal availability of arthropods was highest in the peak nesting period, followed by early and late nesting periods for California sagebrush ( Artemisia californica ), lemonadeberry ( Rhus integrifolia ), non-native grass, and bare ground, whereas availability increased from early to late nesting periods for blue elderberry ( Sambucus mexicana spp. caerulea ), cactus ( Opuntia spp. and Cylindropuntia spp.), California buckwheat ( Eriogonum fasciculatum ), native bunch grasses, and black mustard ( Brassica nigra ). During the early nesting period, arthropods were most abundant in native bunch grasses and least abundant in lemonadeberry. During the peak nesting period, arthropods were most abundant in native bunch grasses and in areas of bare ground and were least abundant in cactus and blue elderberry. During late nesting, arthropods were most abundant in blue elderberry and non-native grass and least abundant in lemonadeberry and mustard. Each year from 2015 to 2019, vegetation data were collected at the same 23 territories where arthropods were sampled: 9 territories in the Otay population and 14 territories in the San Diego population. Cactus, California buckwheat, and non-native grasses were detected within at least 60 percent of sampling points in the Otay population. Cactus, California sagebrush, California buckwheat, non-native grass, and black mustard each were detected within an average of 40 percent of sampling points in the San Diego population. No native bunch grass or lemonadeberry were recorded at the Lake Jennings site within the San Diego population. The cover of shrub species was relatively stable throughout the 5 years. Cover of herbaceous species and bare ground had greater annual variation than shrub species. We found that vegetation cover varied widely among territories, with territory accounting for 69 percent of the variation in vegetation cover. Redundancy analysis allowed us to identify the vegetation types that accounted for the most variation. We used the top scores from the redundancy analysis to identify six vegetation types to be used in generalized linear mixed models analyzing the relationships between vegetation type, precipitation, and Cactus Wren breeding productivity. Three vegetation variables influenced the number of fledglings produced per pair. California sagebrush had a positive effect on the number of fledglings per pair whereas non-native grass and black mustard had a negative effect. Breeding productivity, survival, and movements of adult and first-year Cactus Wrens indicated that the Otay population behaved similarly to, if not out-performed, the San Diego population during the span of our project, suggesting that the driving forces behind low numbers of Cactus Wrens in the Otay population before 2015 were no longer in effect. The Cactus Wren populations in Otay and San Diego reached a peak in 2018, which followed a year of high productivity and survivorship, both of which were correlated with high precipitation. This peak in population size was consistent with reproductive timing and productivity in other bird populations in semi-arid ecosystems that were linked to precipitation and arthropod abundance. We did not find a strong link among arthropod abundance, vegetation composition, and Cactus Wren breeding productivity, likely in part because arthropod abundance varied by vegetation type and sampling period, suggesting that different vegetation types provided important sources of prey at different periods of the breeding season. Arthropod abundance also may not represent arthropod availability when vegetation structure discourages the ground foraging behavior of species such as Cactus Wrens. Cover of non-native grass negatively influenced breeding productivity, although arthropods were abundant in non-native grass. Other factors that could have influenced differential breeding productivity between the Otay and San Diego populations were habitat restoration, control of annual herbaceous vegetation, human disturbance, lingering effects of wildfire, and nest predation. Overall, precipitation appeared to be a driver of Cactus Wren breeding productivity and possibly survival, potentially obscuring proximate effects of arthropod or vegetation composition.

California↗