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Climate variation at Flagstaff, Arizona - 1950 to 2007

Much scientific research demonstrates the existence of recent climate variation, particularly global warming. Climate prediction models forecast that climate will change; it will become warmer, droughts will increase in number and severity, and extreme climate events will recur often—desiccating aridity, extremely wet, unusually warm, or even frigid at times. However, the global models apply to average conditions in large grids approximately 150 miles on an edge (Thorpe, 2005), and how or whether specific areas within a grid are affected is unclear. Flagstaff's climate is mentioned in the context of global change, but information is lacking on the amount and trend of changes in precipitation, snowfall, and temperature. The purpose of this report is to understand what may be happening to Flagstaff's climate by reviewing local climate history. Flagstaff is in north-central Arizona south of San Francisco Mountain, which reaches 12,633 feet, the highest in Arizona (fig. 1). At 6,900 feet, surrounded by ponderosa pine forest, Flagstaff enjoys a four-season climate; winter-daytime temperatures are cool, averaging 45 degrees (Fahrenheit). Summer-daytime temperatures are comfortable, averaging 80 degrees, which is pleasant compared with nearby low-elevation deserts. Flagstaff's precipitation averages 22-inches per year with a range of 9 to 39 inches. Snowfall occurs each season, averaging 97 inches annually. This report, written for the non-technical reader, interprets climate variation at Flagstaff as observed at the National Weather Service (NWS) station at Pulliam Field (or Airport), a first-order weather station staffed by meteorologists (Staudenmaier and others, 2007). The station is on a flat-topped ridge surrounded by forest 5-miles south of Flagstaff at an elevation of 7,003 feet. Data used in this analysis are daily measurements of precipitation (including snowfall) and temperature (maximum and minimum) covering the period from 1950, when the station began operation, through spring 2007. Conversations with Byron Peterson and Michael Staudenmaier of the NWS helped us understand the difficulties of collecting consistent weather data, operation of the station, and Flagstaff's climate. Weather is the daily or even instantaneous state of temperature and precipitation. Climate is the average or accumulation of these parameters over longer time scales such as a week, month, or year. Seasonal (winter, spring, summer, and fall) and annual averages of temperature and accumulated precipitation describe the temporal variation of Flagstaff's climate, which is shown graphically with time series (figs. 2, 4, 6, 8-15). These plots show precipitation or temperature on the ordinate plotted against time on the abscissa, which is a year for annually repeating data or the year of a particular season. The plots reveal changing patterns of precipitation and temperature related to droughts, wet episodes, and rising temperatures.

Arizona↗

Evaluation of streamflow traveltime and streamflow gains and losses along the lower Purgatoire River, southeastern Colorado, 1984-92

Traveltime and gains and losses within a stream are important basic characteristics of streamflow. The lower Purgatoire River flows more than 160 river miles from Trinidad to the Arkansas River near Las Animas. A better knowledge of streamflow traveltime and streamflow gains and losses along the lower Purgatoire River would enable more informed management decisions about the availability of water supplies for irrigation use in southeastern Colorado. In 1994-95, the U.S.\x11Geological Survey, in cooperation with the Purgatoire River Water Conservancy District and the Arkansas River Compact Administration, evaluated streamflow traveltime and estimated streamflow gains and losses using historical surface-water records. Traveltime analyses were used along the lower Purgatoire River to determine when streamflows would arrive at selected downstream sites. The substantial effects of diversions for irrigation and unmeasured return flows in the most upstream reach of the river prevented the tracking of streamflow through reach\x111. Therefore, the estimation of streamflow traveltime for the 60.6 miles of river downstream from Trinidad could not be made.Hourly streamflow data from 1990 through 1994 were used to estimate traveltimes of more than 30 streamflow events for about 100 miles of the lower Purgatoire River. In the middle reach of the river, the traveltime of streamflow for the 40.1\x11miles ranged from about 11 to about 47\x11hours, and in the lower reach of the river, traveltime for the 58.5 miles ranged from about 6 to about 61 hours.Traveltime in the river reaches generally increased as streamflow decreased, but also varied for a specific streamflow in both reaches. Streamflow gains and losses were estimated using daily streamflow data at the upstream and downstream sites, available tributary inflow data, and daily diversion data. Differences between surface-water inflows and surface-water outflows in a reach determined the quantity of water gained or lost. In the most upstream reach of the river near Trinidad, difficulties in establishing streamflow traveltimes prevented the estimation of streamflow gains or losses. From 1984 through 1992, more than 2,900 daily estimates of streamflow gains or losses were made for the last 100\x11miles of the lower Purgatoire River that indicated daily gains and losses in streamflow were common during all four seasons of the year. Although some large daily streamflow gains and losses were computed, most daily estimates indicated small gains and losses in streamflow. The daily median streamflow gain or loss for the middle reach of the river was close to zero during every season, whereas median values for the lower most reach of the river indicated a daily gain in streamflow during every season.

Water-Resources Investigations Report↗

Water-quality assessment of south-central Texas — Descriptions and comparisons of nutrients, pesticides, and volatile organic compounds at three intensive fixed sites, 1996-98

Water-quality samples were collected during April 1996-April 1998 at three intensive fixed sites in the San Antonio region of the South-Central Texas study unit as part of the U.S. Geological Survey National Water-Quality Assessment Program. The sampling strategy for the intensive fixed-site assessment is centered on obtaining information about the occurrence and seasonal patterns of selected constituents including nutrients, pesticides, and volatile organic compounds. The three sites selected to determine the effects of agriculture and urbanization on surface-water quality in the study unit are Medina River at LaCoste (agriculture indicator site), Salado Creek (lower station) at San Antonio (urban indicator site), and San Antonio River near Elmendorf (integrator site). Concentrations of two nutrients, dissolved nitrite plus nitrate nitrogen and total phosphorus, were largest at the integrator site, which is downstream of municipal wastewater treatment plants. Nitrite plus nitrate nitrogen concentrations at this site often exceeded the U.S. Environmental Protection Agency (EPA) maximum contaminant level (MCL) for drinking water. All total phosphorus concentrations at the site exceeded the EPA recommended maximum concentration for streams not discharging directly into reservoirs. Nitrite plus nitrate nitrogen concentrations at the integrator site tended to be smaller, and total phosphorus concentrations at the urban site tended to be larger in samples collected during stormflow than during base flow. The most detections and largest concentrations of three pesticides (atrazine, diazinon, and prometon) were in samples collected at the urban site. Some pesticide concentrations at the agriculture site showed a seasonal pattern of increasing concentrations during spring, the peak application season. Four pesticides (atrazine, deethylatrazine, diazinon, and prometon) were detected in at least 38 percent of samples collected at all three sites. The concentrations of all detected pesticides that have an MCL were less than the MCL at the three sites. More volatile organic compounds (VOC) were detected at the urban indicator site than at the agriculture indicator site, mostly likely because more sources are located in urbanized areas. The most VOCs detected and the largest concentrations of two VOCs (chloroform and tetrahydrofuran) were in samples from the integrator site. More VOCs were detected in samples collected at the integrator site during stormflow than during base flow. The concentrations of all detected VOCs that have an MCL were less than the MCL at the three sites.

Texas↗

The use of sea ice habitat by female polar bears in the Beaufort Sea

Polar bears ( Ursus maritimus ) depend on ice-covered seas to satisfy life history requirements. Modern threats to polar bears include oil spills in the marine environment and changes in ice composition resulting from climate change. Managers need practical models that explain the distribution of bears in order to assess the impacts of these threats. We used stepwise procedures to create resource selection models of habitat use for radio-collared female polar bears in the Beaufort Sea. Sea ice characteristics and ocean depths at known polar bear locations were compared to the same features at randomly selected locations. Models generated for each of four seasons confirmed complexities of habitat use by polar bears and their response to numerous factors. Bears preferred shallow water areas where ice concentrations were > 80 % and different ice types intersected. Variation among seasons was reflected mainly in differential selection of ice stages, floe sizes, and their interactions. Water depth, total ice concentration and distance to the nearest interface between different ice types were significant terms in models for most seasons. Variation in ice stage and form also appeared in three models, and several interaction effects were identified. Habitat selection by polar bears is likely related to prey abundance and availability. Use of habitats in shallow water possibly reflects higher productivity in those areas. Habitat use in close proximity to ice edges is probably related to greater access of prey in those habitats.

Conference Paper↗

Using discrete choice modeling to generate resource selection functions for female polar bears in the Beaufort Sea

Polar bears ( Ursus maritimus ) depend on ice-covered seas to satisfy life history requirements. Modern threats to polar bears include oil spills in the marine environment and changes in ice composition resulting from climate change. Managers need practical models that explain the distribution of bears in order to assess the impacts of these threats. We explored the use of discrete choice models to describe habitat selection by female polar bears in the Beaufort Sea. Using stepwise procedures we generated resource selection models of habitat use. Sea ice characteristics and ocean depths at known polar bear locations were compared to the same features at randomly selected locations. Models generated for each of four seasons confirmed complexities of habitat use by polar bears and their response to numerous factors. Bears preferred shallow water areas where different ice types intersected. Variation among seasons was reflected mainly in differential selection of total ice concentration, ice stages, floe sizes, and their interactions. Distance to the nearest ice interface was a significant term in models for three seasons. Water depth was selected as a significant term in all seasons, possibly reflecting higher productivity in shallow water areas. Preliminary tests indicate seasonal models can predict polar bear distribution based on prior sea ice data.

Alaska↗

Coefficients of productivity for Yellowstone's grizzly bear habitat

This report describes methods for calculating coefficients used to depict habitat productivity for grizzly bears in the Yellowstone ecosystem. Calculations based on these coefficients are used in the Yellowstone Grizzly Bear Cumulative Effects Model to map the distribution of habitat productivity and account for the impacts of human facilities. The coefficients of habitat productivity incorporate detailed information that was collected over a 20-year period (1977-96) on the foraging behavior of Yellowstone's bears and include records of what bears were feeding on, when and where they fed, the extent of that feeding activity, and relative measures of the quantity consumed. The coefficients also incorporate information, collected primarily from 1986 to 1992, on the nutrient content of foods that were consumed, their digestibility, characteristic bite sizes, and the energy required to extract and handle each food. Coefficients were calculated for different time periods and different habitat types, specific to different parts of the Yellowstone ecosystem. Stratifications included four seasons of bear activity (spring, estrus, early hyperphagia, late hyperphagia), years when ungulate carrion and whitebark pine seed crops were abundant versus not, areas adjacent to (< 100 m) or far away from forest/nonforest edges, and areas inside or outside of ungulate winter ranges. Densities of bear activity in each region, habitat type, and time period were incorporated into calculations, controlling for the effects of proximity to human facilities. The coefficients described in this report and associated estimates of grizzly bear habitat productivity are unique among many efforts to model the conditions of bear habitat because calculations include information on energetics derived from the observed behavior of radio-marked bears.

Biological Science Report↗

Delayed conifer mortality after fuel reduction treatments: Interactive effects of fuel, fire intensity, and bark beetles

Many low‐elevation dry forests of the western United States contain more small trees and fewer large trees, more down woody debris, and less diverse and vigorous understory plant communities compared to conditions under historical fire regimes. These altered structural conditions may contribute to increased probability of unnaturally severe wildfires, susceptibility to uncharacteristic insect outbreaks, and drought‐related mortality. Broad‐scale fuel reduction and restoration treatments are proposed to promote stand development on trajectories toward more sustainable structures. Little research to date, however, has quantified the effects of these treatments on the ecosystem, especially delayed and latent tree mortality resulting directly or indirectly from treatments. In this paper, we explore complex hypotheses relating to the cascade of effects that influence ponderosa pine (Pinus ponderosa) and Douglas‐fir (Pseudotsuga menziesii) mortality using structural equation modeling (SEM). We used annual census and plot data through six growing seasons after thinning and four growing seasons after burning from a replicated, operational‐scale, completely randomized experiment conducted in northeastern Oregon, USA, as part of the national Fire and Fire Surrogate study. Treatments included thin, burn, thin followed by burn (thin + burn), and control. Burn and thin + burn treatments increased the proportion of dead trees while the proportion of dead trees declined or remained constant in thin and control units, although the density of dead trees was essentially unchanged with treatment. Most of the new mortality (96%) occurred within two years of treatment and was attributed to bark beetles. Bark beetle‐caused tree mortality, while low overall, was greatest in thin + burn treatments. SEM results indicate that the probability of mortality of large‐diameter ponderosa pine from bark beetles and wood borers was directly related to surface fire severity and bole charring, which in turn depended on fire intensity, which was greater in units where thinning increased large woody fuels. These results have implications when deciding among management options for restoring ecosystem health in similar ponderosa pine and Douglas‐fir forests.

Ecological Applications↗

Determination of the sources of nitrate contamination in karst springs using isotopic and chemical indicators

The sources of nitrate (NO-3) in groundwater of the shallow karst aquifer in southwestern Illinois' sinkhole plain were investigated using chemical and isotopic techniques. The groundwater in this aquifer is an important source of potable water for about half of the residents of the sinkhole plain area. Previous work has shown that groundwater from approximately 18% of the wells in the sinkhole plain has NO-3 concentrations in excess of the USEPA's drinking water standard of 10 mg N/1. Relative to background levels, the NO-3 concentrations in water from 52% of the wells, and probably all of the springs in the study area, are anomalously high, suggesting that sources other than naturally occurring soil organic matter have contributed additional NO-3 to groundwater in the shallow karst aquifer. This information, and the dominance of agriculture in the study area, suggest that agrichemical contributions may be significant. To test this hypothesis, water samples from 10 relatively large karst springs were collected during four different seasons and analyzed for inorganic constituents, dissolved organic carbon, atrazine, and ??15N and ??18O of the NO-3 ions. The isotopic data were most definitive and suggested that the sources of NO-3 in spring water are dominated by N-fertilizer with some possible influence of atmospheric NO-3 and, to a much lesser extent, human and/or animal waste. Differences in the isotopic composition of NO-3 and some of the chemical characteristics were observed during the four consecutive seasons in which spring water samples were collected. Isotopic values for ??15N and ??18O of the NO-3 ranged from 3.2??? to 19.1??? and from 7.2??? to 18.7???, respectively. The trend of ??15N and ??18O data for NO-3 also indicated that a significant degree of denitrification is occurring in the shallow karst hydrologic system (within the soil zone, the epikarst and the shallow karst aquifer) prior to discharging to springs. ?? 2001 Elsevier Science B.V. All rights reserved.

Chemical Geology↗

Intersexual and temporal variation in foraging ecology of prothonotary warblers during the breeding season

We studied foraging ecology of Prothonotary Warblers ( Protonotaria citrea ) over four breeding seasons to determine if this species exhibited sex-specific or temporal variation in foraging behavior. Significant differences between sexes during the prenestling period were found for foraging height and substrate height (foraging method, plant species/substrate, perch diameter, horizontal location from trunk, and prey location were not significantly different). During the nestling period, this divergence between sexes was evident for foraging height, substrate height, substrate / tree species, and prey location. Additionally, male warblers significantly altered their behavior for all seven foraging variables between the two periods, whereas females exhibited changes similar to those of males for five of the foraging variables. This parallel shift suggests a strong behavioral response by both sexes to proximate factors (such as vegetation structure, and prey abundance and distribution) that varied throughout the breeding season. Sex-specific foraging behavior during the prenestling period was best explained by differences in reproductive responsibilities rather than by the theory of intersexual competition for limited resources. During the nestling period, neither hypothesis by itself explained foraging divergences adequately. However, when integrated with the temporal responses of the warblers to changes in prey availability, reproductive responsibilities seemed to be of primary importance in explaining intersexual niche partitioning during the nestling period. We emphasize the importance of considering both intersexual and intraseasonal variation when quantifying a species' foraging ecology.

The Auk↗

Water-quality, bed-sediment, and biological data (October 2013 through September 2014) and statistical summaries of data for streams in the Clark Fork Basin, Montana

Water, bed sediment, and biota were sampled in streams from Butte to near Missoula, Montana, as part of a monitoring program in the upper Clark Fork Basin of western Montana. The sampling program was led by the U.S. Geological Survey, in cooperation with the U.S. Environmental Protection Agency, to characterize aquatic resources in the Clark Fork Basin, with emphasis on trace elements associated with historic mining and smelting activities. Sampling sites were located on the Clark Fork and selected tributaries. Water samples were collected periodically at 20 sites from October 2013 through September 2014. Bed-sediment and biota samples were collected once at 14 sites during August 2014. This report presents the analytical results and qualityassurance data for water-quality, bed-sediment, and biota samples collected at sites from October 2013 through September 2014. Water-quality data include concentrations of selected major ions, trace elements, and suspended sediment. At 12 sites, dissolved organic carbon and turbidity samples were collected. In addition, nitrogen (nitrate plus nitrite) samples were collected at two sites. Daily values of mean suspended-sediment concentration and suspended-sediment discharge were determined for four sites. Seasonal daily values of turbidity were determined for four sites. Bed-sediment data include trace-ele&shy;ment concentrations in the fine-grained fraction. Biological data include trace-element concentrations in wholebody tissue of aquatic benthic insects. Statistical summaries of water-quality, bed-sediment, and biological data for sites in the upper Clark Fork Basin are provided for the period of record.

Montana↗

Pesticides in rain in four agricultural watersheds in the United States

Rainfall samples were collected during the 2003 and 2004 growing seasons at four agricultural locales across the USA in Maryland, Indiana, Nebraska, and California. The samples were analyzed for 21 insecticides, 18 herbicides, three fungicides, and 40 pesticide degradates. Data from all sites combined show that 7 of the 10 most frequently detected pesticides were herbicides, with atrazine (70%) and metolachlor (83%) detected at every site. Dacthal, acetochlor, simazine, alachlor, and pendimethalin were detected in more than 50% of the samples. Chlorpyrifos, carbaryl, and diazinon were the only insecticides among the 10 most frequently detected compounds. Of the remaining pesticide parent compounds, 18 were detected in fewer than 30% of the samples, and 13 were not detected. The most frequently detected degradates were deethylatrazine; the oxygen analogs (OAs) of the organophosphorus insecticides chlorpyrifos, diazinon, and malathion; and 1-napthol (degradate of carbaryl). Deethylatrazine was detected in nearly 70% of the samples collected in Maryland, Indiana, and Nebraska but was detected only once in California. The OAs of chlorpyrifos and diazinon were detected primarily in California. Degradates of the acetanilide herbicides were rarely detected in rain, indicating that they are not formed in the atmosphere or readily volatilized from soils. Herbicides accounted for 91 to 98% of the total pesticide mass deposited by rain except in California, where insecticides accounted for 61% in 2004. The mass of pesticides deposited by rainfall was estimated to be less than 2% of the total applied in these agricultural areas.

Journal of Environmental Quality↗

Modelling effects of flow withdrawal scenarios on riverine and riparian features of the Yampa River in Dinosaur National Monument

The National Park Service (NPS) is charged with maintaining natural riverine resources and processes in its parks along the Yampa River and downstream along the Green River. This mission requires information on how proposed water withdrawals would affect resources. We present a methodology that quantifies the impact on natural riverine and riparian features of Dinosaur National Monument based on alternative withdrawals that vary in volume and timing. This methodology uses a reverse quantification and develops tools to enable the NPS to ensure that if withdrawals must occur, the adverse impacts would be minimized by prescribing or constraining the timing, magnitude, and duration of withdrawal. The reverse quantification, well-suited for unregulated rivers such as the Yampa, strives to protect all flows minus extractions from daily flows based on three parameters: 1) a minimum flow, below which water diversion does not occur; 2) the percentage of the flow above the minimum that is diverted; 3) the maximum daily flow that is diverted. We apply 350 flow extraction scenarios, each defined by a unique set of parameters, to the 99 historic annual hydrographs of daily flows (water year (WY) 1922–2020), and to the more recent 20 years (WY 2001–2020). We also consider how hydrologic year type (wet to dry) influences the flow volume extracted and impact to the resource. Recognizing the seasonal differences in flow and ecological and geomorphic response, we divide each year into four distinct seasonal periods and use relations from the literature between flow, channel change, riparian vegetation and fish behavior, physiology, and habitat to define hydrograph and resource metrics used to evaluate impacts to the resource. While our analysis demonstrates that all withdrawals will damage the resource, extractions during the Early Runoff Period (March 15 – April 30) are least detrimental and extractions during the Summer Baseflow Period (July 16 – October 31) are most detrimental. We find that most aspects of the resource are more sensitive to increasing extractions during drier years than during wetter years. Recent decades have seen a shift towards more frequent drier years, resulting in less water in most periods. As a result, our analysis suggests that extractions in recent decades would have had a greater impact on the resource when compared to similar extractions during the full historical record. Finally, we demonstrate how the NPS may use these results to develop limits on extractions for resource protection.

Colorado, Utah↗

Hydrology and chemistry of selected prairie wetlands in the Cottonwood Lake area, Stutsman County, North Dakota, 1979-82

The relation of hydrologic setting and temporal variability in hydrology to nutrient content and geochemical characteristics of a group of prairie wetlands and adjacent ground water was studied during the period 1979-82. Although data were collected from many wetlands and wells at the study site, emphasis in this report primarily is on four wetlands two seasonal and two semipermanent and four wells contiguous to them along a hydrologic section. The seasonal wetlands, T8 and T3, contained water only for a few weeks to months after filling in spring and early summer; both were completely dry by August. The semipermanent wetlands, PI and P8, contained water throughout each year and were ice covered in winter. One wetland, T8, recharges ground water. Wetlands PI and P8 are in areas of ground-water discharge. None of the wetlands received water by channelized surface-water inlets. Only wetland P8 had a channelized surface-water outlet. Ground-water-level data showed that high points of the water table did not always occur beneath land-surface highs. Reversals of ground-water flow occurred occasionally between two of the wetlands, T3 and PI. Significant differences existed in the chemical composition of the wetlands based on their hydrologic setting. In general, the dominant cation and anion in the wetlands were potassium and bicarbonate in wetland T8, calcium and sulfate in wetland T3, magnesium and sulfate in wetland PI, and magnesium and bicarbonate in wetland P8. Significant seasonal differences existed in the water chemistry of the wetlands in ground-water discharge areas. Water in three of the wetlands, T3, Pi, and P8, was most dilute while they filled in spring after icemelt. Concentration increased during the open-water period, and two of the wetlands, PI and P8, became most concentrated under ice cover. Concentrations of total phosphorus and total nitrogen were greatest in wetlands in areas of ground-water recharge and least in wetlands in areas of ground-water discharge. Differences in the chemistry of water from wells in the adjacent ground water resulted primarily from the positions of the wells in the ground-water flow system. The chemical type of water from well 12, which was located in a ground-water recharge area, was calcium sodium bicarbonate. Water from well 4, located downgradient from wetland T8, and from well 16, located downgradient from wetland PI, typically was a calcium sulfate type. Water from well 13, located between wetlands T3 and PI in an area of changing ground-water flow directions, was a magnesium sulfate type. Data from this study show that an understanding of hydrologic conditions is important in the interpretation of the water chemistry of wetlands in the study area.

Professional Paper↗

Detecting annual and seasonal changes in a sagebrush ecosystem with remote sensing-derived continuous fields

Climate change may represent the greatest future risk to the sagebrush ecosystem. Improved ways to quantify and monitor gradual change resulting from climate influences in this ecosystem are vital to its future management. For this research, the change over time of five continuous field cover components including bare ground, herbaceous, litter, sagebrush, and shrub were measured on the ground and by satellite across six seasons and four years. Ground-measured litter and herbaceous cover exhibited the highest variation annually and herbaceous cover the highest variation seasonally. Correlation of ground measurements to corresponding remote-sensing predictions indicated that annual predictions tracked ground measurements more closely than seasonal ones, and QuickBird predictions tracked ground measurements more closely than Landsat predictions. When annual linear slope values from ground plots and sensor predictions were correlated by component, the direction of ground-measured change was tracked better with QuickBird components than with Landsat components. Component predictions were correlated to annual and seasonal DAYMET precipitation. QuickBird components on average had the best response to precipitation patterns, followed by Landsat components. Overall, these results demonstrate the ability of sagebrush ecosystem components as predicted by regression trees to incrementally measure changing components of a sagebrush ecosystem.

Wyoming↗

Water velocity and suspended solids measurements by in-situ instruments in Upper Klamath Lake, Oregon

The U. S. Geological Survey conducted hydrodynamic measurements in Upper Klamath Lake during four summer seasons (approximately mid-June to mid-September) during 2003 to 2006. Measurements included water current profiles made by acoustic Doppler current profilers at a number of fixed locations in the lake during all four years as well as from a moving boat during 2005 and 2006. Measurements of size distribution of suspended material were made at four locations in the lake during 2004-2006. Raw (unfiltered) data are presented as time series of measurements. In addition, water-velocity data have been filtered to remove wind-induced variations with periods less than thirty hours from the measurements. Bar graphs of horizontal and vertical water speed and acoustic backscatter have been generated to discern diurnal variations, especially as they relate to wind patterns over the lake. Mean speeds of the horizontal currents in the lake range between about 3.5 to 15 cm/s with the higher speeds at the deep locations in the trench on the west side of the lake. Current directions generally conform to the lake’s bathymetry contours and the water circulation pattern is usually in a clockwise direction around the lake as established by the prevailing north to northwesterly surface winds in the region. Diurnal patterns in horizontal currents probably relate to diurnal wind patterns with minimum wind speeds near noon and maximum wind speeds near 2100. Diurnal variations in vertical velocities do not appear to be related to wind patterns; they do appear to be related to expected patterns of vertical migration of Aphanizomenon flos aquae , (AFA) the predominant species of blue-green algae in the lake. Similarly, diurnal variations in acoustic backscatter, especially near the lake’s surface, are probably related to the vertical migration of AFA.

Oregon↗

Grasslands maintain stability in productivity through compensatory effects and dominant species stability under extreme precipitation patterns

Extreme climatic events are likely to intensify under climate change and can have different effects on ecosystems depending on their timing and magnitude. Understanding how productivity responds to extreme precipitation patterns requires assessing stability and vulnerability during critical growing periods at the plant community level. In this study, we experimentally imposed two contrasting types of extreme precipitation patterns, including extreme drought (excluding all rainfall for 30 consecutive days) during early-, mid-, and late-stages of the growing season, and heavy rainfall (adding 14.1 mm of rainfall every day for 20 consecutive days) during mid- and late-stages of the growing season over four years (2013–2016) in a steppe community in Inner Mongolia, China. We found that extreme drought and heavy rainfall had no effect on community aboveground net primary productivity (ANPP), species richness, and dominance at any stage of the growing season. Community stability in response to extreme drought was mainly driven by compensation among species and the stability of dominant species, while the compensatory effect among species and functional groups, and the stability of dominant species contributed to the community stability in response to heavy rainfall. Overall, our findings indicate that the responses of the ecosystem to intra-seasonal contrasting extreme precipitation patterns can be driven by similar stability mechanisms and suggest that semiarid temperate steppe communities may have strong initial resistance to more frequent extreme climatic events in the future.

Ecosystems↗

High diet overlap between native small-bodied fishes and nonnative fathead minnow in the Colorado River, Grand Canyon, Arizona

River regulation may mediate the interactions among native and nonnative species, potentially favoring nonnative species and contributing to the decline of native populations. We examined food resource use and diet overlap among small-bodied fishes in the Grand Canyon section of the Colorado River as a first step in evaluating potential resource competition. We compared the diets of the predominant small-bodied fishes (native Speckled Dace Rhinichthys osculus , juvenile Flannelmouth Sucker Catostomus latipinnis , and juvenile Bluehead Sucker C. discobolus , and nonnative Fathead Minnow Pimephales promelas ) across seasons at four sites downstream of Glen Canyon Dam using nonmetric multidimensional scaling and Schoener's similarity index. The diets of these fishes included diatoms, amorphous detritus, aquatic invertebrates (especially simuliid and chironomid larvae), terrestrial invertebrates, and terrestrial vegetation. Diets varied with season and were affected by high turbidity. Fish consumed more amorphous detritus and terrestrial vegetation during the summer monsoon season (July–September), when turbidity was higher. The diets of all species overlapped, but there was large variation in the degree of overlap. The diets of juvenile suckers and Fathead Minnows were most similar, while Speckled Dace had relatively distinct diets. The differences took the form of higher proportions of diatoms and amorphous detritus in the diets of Bluehead Suckers and Fathead Minnows and higher proportions of simuliids and chironomids in those of Speckled Dace. If food resources are or become limiting, diet overlap suggests that competition may occur among native and nonnative species, which could have implications for the population dynamics of these fishes and for the management of the Colorado River ecosystem in Grand Canyon.

Arizona↗

Are there optimal densities for prairie birds?

The major forces of food and predation shape fitness-enhancing decisions of birds at all stages of their life cycles. During the breeding season, birds can minimize nest loss due to predation by selecting sites with a lower probability of predation. To understand the environmental and social aspects and consequences of breedingsite selection in prairie birds, we explored variation in nest-survival patterns of the Lark Bunting (Calamospiza melanocorys) in the shortgrass prairie region of North America. Over four breeding seasons, we documented the survival of 405 nests, conducted 60 surveys to estimate bird densities, and measured several vegetative features to describe habitat structure in 24 randomly selected study plots. Nest survival varied with the buntings' density as described by a quadratic polynomial, increasing with density below 1.5 birds ha -1 and decreasing with density between 1.5 and 3 birds ha -1 , suggesting that an optimal range of densities favors reproductive success of the Lark Bunting, which nests semi-colonially. Nest survival also increased with increasing vegetation structure of study plots and varied with age of the nest, increasing during early incubation and late in the nestling stage and declining slightly from mid-incubation to the middle of the nestling period. The existence of an optimal range of densities in this semi-colonial species can be elucidated by the "commodity-selection hypothesis" at low densities and density dependence at high densities. ?? The Cooper Ornithological Society 2010.

Condor↗