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Assessing biodiversity in Nuevo Leon, Mexico: Are nature reserves the answer?

The Mexican state of Nuevo Leon, located in the northeastern portion of the country, currently has 26 state and three federal nature reserves covering approximately 4.5% of its land area. These reserves were established for a variety of reasons not necessarily related to conservation purposes. In 2000 in response to a growing concern about the lack of organized conservation reserve planning to protect the important biological and physical features of Mexico, the Mexican Commission for Knowledge and Use of Biodiversity proposed 12 new terrestrial reserves for Nuevo Leon. The new reserves, if established, would increase the proportion of protected lands in the state to almost 24% of the state's land area. We compiled a Geographic Information System (GIS) analysis using digital thematic maps of physical and ecological features to examine how well the existing and proposed reserves incorporated the major biological and physical features of the state. The existing reserves are located primarily in regions with elevations > 1,000-1,500 m, on less productive soils, and are dominated by pine and oak forest cover types. As a result, the state's dominant biotic region - low elevation coastal plain with xeric scrub vegetation - is disproportionately under represented in the current reserve system. The new reserves would expand the protection of biophysical resources throughout the state. However, the inclusion of important resources in the low elevation coastal lands would still be limited.

Nuevo Leon↗

Level II scour analysis for Bridge 6 (BRISVT01160006) on State Highway 116, crossing Little Notch Brook, Bristol, Vermont

This report provides the results of a detailed Level II analysis of scour potential at structure BRISVT01160006 on State Route 116 crossing the Little Notch Brook, Bristol, Vermont (figures 1–8). A Level II study is a basic engineering analysis of the site, including a quantitative analysis of stream stability and scour (U.S. Department of Transportation, 1993). Results of a Level I scour investigation also are included in Appendix E of this report. A Level I investigation provides a qualitative geomorphic characterization of the study site. Information on the bridge, gleaned from Vermont Agency of Transportation (VTAOT) files, was compiled prior to conducting Level I and Level II analyses and is found in Appendix D. The site is in the Green Mountain section of the New England physiographic province of West-central Vermont in the town of Bristol. The 8.59-mi 2 drainage area is in a predominantly rural and forested basin. In the vicinity of the study site, the surface cover is dense forest except for the downstream left side, which is row crops. In the study area, Little Notch Brook has a sinuous channel with a slope of approximately 0.005 ft/ft, an average channel top width of 32 ft and an average channel depth of 4 ft. The predominant channel bed material is sand and gravel with a median grain size (D 50 ) of 17.4 mm (0.0570 ft). The geomorphic assessment at the time of the Level I and Level II site visit on June 13, 1996, indicated that the reach was laterally unstable. The sinuous configuration of the channel with fine bed and bank material, a sharp channel bend upstream, and point bars and cut-banks upstream and downstream of this site are among the primary characteristics, which suggest lateral instability. In addition, there is evidence of streambed degradation at this site. A large eddy was noted at the location where Little Notch Brook enters the New Haven River about 100 feet downstream. There was a large scour hole noted at the location of the eddy, which is likely to remove streambed material at least as quickly as supplied from upstream on Little Notch Brook. Hence, channel degradation may be significant during a flood event. The state route 116 crossing of Little Notch Brook is a 24-ft-long, two-lane bridge consisting of one 21-foot concrete span (Vermont Agency of Transportation, written communication, December 14, 1995). The bridge is supported by vertical, concrete abutments with wingwalls. The channel is skewed approximately 15 degrees to the opening while the opening-skew-to-roadway is 25 degrees. There was one foot of scour evident along the downstream half of the left abutment footing and some separation of the left abutment wall from the deck above due to settling. The left abutment footing was undermined up to a foot at the downstream end. The scour protection measures at the site were type-1 stone fill (less than 12 inches diameter) on the upstream left bank and type-2 stone fill (less than 36 inches diameter) on the right banks and right wingwalls upstream and downstream of the structure. Additional details describing conditions at the site are included in the Level II Summary and Appendices D and E. Scour depths and rock rip-rap sizes were computed using the general guidelines described in Hydraulic Engineering Circular 18 (Richardson and others, 1995). Total scour at a highway crossing is comprised of three components: 1) long-term streambed degradation; 2) contraction scour (due to accelerated flow caused by a reduction in flow area at a bridge) and; 3) local scour (caused by accelerated flow around piers and abutments). Total scour is the sum of the three components. Equations are available to compute depths for contraction and local scour and a summary of the results of these computations follows. Contraction scour for all modelled flows ranged from 3.2 to 4.3 ft. The worst-case contraction scour occurred at the 500-year discharge. Abutment scour ranged from 6.0 to 10.0 ft. The worst-case abutment scour occurred at the 500-year discharge. Additional information on scour depths and depths to armoring are included in the section titled “Scour Results”. Scoured-streambed elevations, based on the calculated scour depths, are presented in tables 1 and 2. A cross-section of the scour computed at the bridge is presented in figure 8. Scour depths were calculated assuming an infinite depth of erosive material and a homogeneous particle-size distribution. It is generally accepted that the Froehlich equation (abutment scour) gives “excessively conservative estimates of scour depths” (Richardson and others, 1995, p. 47). Usually, computed scour depths are evaluated in combination with other information including (but not limited to) historical performance during flood events, the geomorphic stability assessment, existing scour protection measures, and the results of the hydraulic analyses. Therefore, scour depths adopted by VTAOT may differ from the computed values documented herein.

Vermont↗

Effect of timber harvest on soil carbon storage at Blodgett Experimental Forest, California

Four plots from a mixed conifer forest were similarly cleared, burned, and replanted at various times over 17 years; a plot logged 79 years before sampling was used as a control. The plots had similar slope (2 to 15%, midslope position), aspect (south to southeast), and soil type (Holland series: mesic Haploxeralf; a Gray Brown Luvisol in the Canadian classification system). Twenty sites at each plot were sampled volumetrically by horizon to 20 cm below the organic–mineral soil boundary. Samples were analyzed for bulk density, organic C, and total N. There was an initial loss (15%) of organic C from the soil within 1 to 7 years, likely the result of oxidation (burning and decomposition) and erosion. For 17 years of forest regrowth, the soil continued to lose C (another 15%), probably owing to decomposition of slash material and possibly erosion, despite the slight accumulation of new litter and roots. After 80 years of regrowth, rates of carbon accumulation exceeded rates of loss, but carbon storage had declined and was not likely to recover to preharvest levels. Timber harvest and site preparation dramatically altered soil C and N distribution, in which C/N ratios after site preparation were initially high throughout the upper 20 cm. Subsequently, C/N ratios became lower with depth and with recovery age. Although stocks of C and N varied considerably among the plots and did not change consistently as a function of recovery age, the C/N ratios did vary systematically with recovery age. We hypothesize that the amount of C ultimately stored in the soil at steady state depends largely on N reserves and potentials, which appear to vary with erosion, intensity of burning, and site treatment.

California↗

Evaluating management alternatives for Wyoming elk feedgrounds in consideration of chronic wasting disease

Executive Summary The authors used decision and modeling analyses to evaluate management alternatives for a decision on whether to permit Cervus canadensis (elk) feeding on two sites on Bridger-Teton National Forest, Dell Creek and Forest Park. Supplemental feeding of elk could increase the transmission of chronic wasting disease (CWD) locally and disease spread regionally, potentially impacting elk populations over time with wider implications for Odocoileus hemionus (mule deer) and Odocoileus virginianus (white-tailed deer) populations and hunting, tourism, and regional revenue. Supplemental feeding is thought to improve overwinter elk survival and reduce the commingling of elk with cattle during months when brucellosis transmission risk is highest. We worked with the U.S. Department of Agriculture Forest Service to identify their fundamental objectives and associated performance metrics related to this feedground decision. We then developed disease and habitat selection models to quantify the effect of four management alternatives on select performance metrics. The four alternatives were to continue to permit feeding, phaseout permits to feed in three years, permit feeding on an emergency basis, or stop permitting feeding. In this report, we present methods and summarized results on disease and habitat selection models and summaries of other performance metrics analyzed by BIO-WEST, Inc. and Cirrus Ecological Solutions as part of an Environmental Impact Statement. Data from Wyoming Game and Fish Department (WGFD) supported the assumption that supplemental elk feeding allows for larger elk populations in a region. We documented that herd units (HU) without feedgrounds had 23 percent lower densities of elk per area of winter range when compared against HUs with feedgrounds, after accounting for differences in sightability of elk during counts on and off feedgrounds. Thus, throughout our analyses, we assumed feedground closures would reduce elk carrying capacity resulting in an average decline of previously fed elk population segments by 23 percent (5th and 95th percentiles = [11 percent, 35 percent]) by year 20. Most of that decline occurred within the first few years after a feedground ceases to operate. We used a panel of CWD experts to help estimate CWD trans-mission in fed and unfed elk population segments. In aggregate, the expert panel estimated that median values of direct and indirect transmission of CWD are expected to be 1.9 and 4 times higher, respectively, in fed elk populations compared to unfed elk. We used these disease transmission estimates in combination with local elk demographic rates and carrying capacity estimates to project disease and population dynamics. In year 20, we predicted CWD prevalence would increase to 42 percent (5th and 95th percentiles = [29 percent, 55 percent]), and 13 percent (5th and 95th percentiles = [4 percent, 26 percent]) on average for fed and unfed elk population segments, respectively, given a starting prevalence of 1.6 percent. The prevalence estimates for the unfed elk population segments are in the range of previous observations of CWD in elk in the western United States. The average CWD prevalence from 2016 to 2018 in the unfed elk population of Wind Cave National Park in South Dakota was 18 percent overall but up to 30 percent in some regions (Sargeant and others, 2021). Meanwhile, CWD prevalence in the Iron Mountain and Laramie Peak elk herds in Wyoming from 2016 to 2018 was 14 percent and 7 percent, respectively, despite being present since at least 2002 (Wyoming Game and Fish Department, 2020b). From 2016 to 2020, elk that were fed at Dell Creek and Forest Park constituted on average 12–20 percent of the total elk on their respective HUs. As a result, the differences between management alternatives are modest when considering the closure of only one feedground on a HU. The no feeding alternative for Forest Park resulted in a CWD prevalence of 17 percent (SD = 7 percent) in the Afton HU compared to 20 percent (SD = 7 percent) with continued feeding by year 20. In the Upper Green River HU, no feeding on Dell Creek resulted in a CWD prevalence of 27 percent (SD = 6 percent) compared to 30 percent (SD = 5 percent) with continued feeding. In terms of disease-associated mortality, we predicted the closure of Forest Park and Dell Creek feedgrounds would reduce the total number of CWD mortalities by 9 percent in the Upper Green River HU and 26 percent in the Afton HU during the 20-year timespan. Our spatial analyses predicted that management alternative effects vary by HU as a function of private property and other wildlife winter ranges proximity relative to feedground location. The predicted number of elk abortions on private land, as a proxy for brucellosis risk to cattle, may increase by 8–21 percent in the absence of feeding at Dell Creek and Forest Park. Eight feedgrounds are located on Bridger-Teton National Forest, all of which have permits that have expired or will expire prior to 2028. In addition, WGFD could change their management of feedgrounds given new information; therefore, we also assessed the cumulative effects of continued feeding, phaseout, and no feeding management alternatives across five HUs south of Jackson, Wyoming (Afton HU, Fall Creek HU, Piney HU, Pinedale HU, and Upper Green River HU). These five HUs ranged from about 41 to 85 percent of the elk herd using feedgrounds, which corresponded to a CWD prevalence at year 20 of 23–34 percent if all feedgrounds in those five HUs remained open relative to 12 to 14 percent if all feedgrounds were closed. We predicted feedground closures may result in immediate reductions in population size relative to alternatives that continue feeding (for example, continued feeding and emergency feeding alternatives); however, over longer periods of time, CWD-associated mortality leads to larger population reductions. The no feeding alternative resulted in higher elk population sizes compared to the continued feeding alternative after about 10 years of implementation. Delayed action under a phaseout alternative resulted in increasing the CWD prevalence to 20 percent relative to 12 to 14 percent, on average, without feeding on HUs with a large population of fed elk such as the Upper Green River HU. Summarizing our cumulative results across all five of the analyzed HUs, we predicted continued feeding will lead to fewer elk by year 20 (mean = 8,300, standard deviation [SD] = 740) compared to no feeding at U.S. Department of Agri-culture Forest Service sites (10,700, SD = 890). The closure of all feedgrounds was projected to result in the largest elk populations at year 20 (12,500, SD = 980). No feeding at all sites also resulted in the largest cumulative harvest of 57,700 (SD = 2,600) compared to 51,100 (SD = 3,800) for continued feeding at all current feedground sites on the five HUs. Continued feeding also resulted in the lowest brucellosis costs to producers ($194,600, SD = $11,500) compared to no feeding on all feedgrounds ($243,000, SD = $13,700). Assuming moderate reductions in hunter interest because of increasing CWD prevalence in elk, we predicted that no feeding resulted in regional revenues generated by hunting activities of $190 million (SD = $10 million) compared to $173 million (SD = $10 million) for continued feeding over the 20-year timeframe. Recent CWD detections in mule deer and elk in Grand Teton National Park has elevated the importance of the cur-rent decision on whether, and how, to permit elk feeding on Dell Creek and Forest Park and the management of the other feedgrounds. Aggressive male harvest has slowed, but not stopped, the increasing prevalence of CWD in mule deer (Conner and others, 2021). It is unclear whether harvest management can be an effective tool to slow the spread of CWD in elk. There are also no effective treatments or vaccines for CWD, and it is unlikely that any will be developed that can be easily deployed in the near future. Thus, reducing artificial aggregations is one of the few management approaches suggested by the Western Association of Fish and Wildlife Agencies (Almberg and others, 2017). Future surveillance and monitoring can be designed to resolve uncertainties that can improve future decision-making. If feedgrounds close, research could quantify elk population reductions in the absence of feeding, the redistribution of fed elk to other places, or the consequences of elk movement on private property. If feedgrounds remain open, research could assess how rapidly CWD spreads in artificial aggregations of elk; however, surveillance programs would need to be designed with sufficient power to detect initial changes of CWD prevalence. Delaying action on feedground management was projected to be costly. Results of the phaseout alternative relative to the no feeding alternative suggested a 3-year delay was enough for substantial long-term changes in CWD prevalence. The long-term persistence of infectious CWD prions in the environment suggests that feedground management decisions may have long-lasting consequences. Our results indicated tradeoffs in the ability of a management agency to achieve all their objectives, and all management alternatives resulted in significant reductions in elk population size. This report contains the foundational elements for formal decision analysis methods, which can be implemented to help decision makers transparently evaluate the consequences of decision alternatives and identify the set of actions that best achieve agency and stakeholder priorities.

Wyoming↗

2018 Kaua'i forest bird population estimates and trends

Kaua‘i's native forest birds have experienced steep declines since the beginning of systematic surveys in 1981, and declines have accelerated in recent decades. This report details the analysis of the most recent surveys conducted in 2018. Incorporating the new survey results, long-term trends continue to show sharp declines for all native honeycreeper species with the exception of ‘apapane (Himatione sanguinea), which has stable numbers in core areas of its range. Kaua‘i ‘elepaio (Chasiempis sclateri) continued to decline in the outer portions of its range but increased in the core areas of its range. Abundance estimates of forest birds ranged from slightly higher to slightly lower in most species, indicating a relatively stable period from 2012–2018, and a pause from the rapid declines seen in earlier periods. Many native species now exist in very low numbers, and variation in estimates from survey to survey will increase.

Hawaii↗

Millennial-scale climatic and cultural impacts on vegetation and fire at the southern edge of the Rocky Mountains, USA

Mountain forests and woodlands in semiarid regions of the world are threatened by climatic change and other human impacts. In the southwestern USA, climate and culturally driven changes to the structure and fire regimes of dry coniferous forests over recent centuries are well documented by tree-ring archives. However, the roles of climate and people as drivers of millennial-scale changes are less understood. We present a new record of vegetation dynamics and regional fire activity inferred from pollen, microscopic charcoal, and sediment geochemistry from Santa Fe Lake, NM (3532 m a.s.l.), the southernmost natural lake in the Rocky Mountains. Declining elemental intensities (e.g., Ti, Fe, Si) record local deglaciation after 14,000 cal yr BP followed by upslope expansion of trees indicated by increasing Pinus and Picea pollen percentages as the climate warmed during the Late Glacial. Cool, dry growing seasons probably limited the establishment of dense forests until after 12,200 cal yr BP when we hypothesize that Pinus ponderosa (ponderosa pine) expanded regionally (i.e., within 10–100 km) due to continued warming and a possible increase in monsoonal precipitation. Subalpine forests established near Santa Fe Lake by 10,300 cal yr BP as indicated by abundant conifer stomates and increasing Picea and Pinus aristata pollen percentages, which are highest after 5100 cal yr BP. Increasing Juniperus -type, Quercus , and Pinus edulis -type pollen record changes in vegetation belts at lower elevations after 10,300 cal yr BP. Maize pollen, an unambiguous indicator of Puebloan agriculture, first appears at 800 cal yr BP (1150 cal yr CE) and is present through 1600 cal yr CE when Spanish colonization brought cultural upheaval and population loss to local indigenous communities. Regional fire activity, inferred from microscopic charcoal influx, remained relatively constant for most of our record. However, average microscopic charcoal influx more than doubled relative to the rest of the Holocene during the last 2000 years, with the highest influx after 1550 cal yr BP (400 cal yr CE). The increased influx of microscopic charcoal coincides with archaeological evidence for dramatic regional population growth in the northern Rio Grande region and probably reflects extensive cultural burning associated with the development of an agricultural economy. Although climate was an important driver of vegetation change and fire regimes in the mountains of the southwestern USA during the Holocene, our results suggest that human impacts on regional burning during the last 2000 years far exceeded earlier climate-driven variation.

Rocky Mountains↗

A test of the variable circular-plot method where exact density of a bird population was known

Variable circular-plot (VCP) counts are statistically more sound than point counts because they are adjusted for the probability of detecting birds at different distances and under different conditions. However, many ornithologists use point counts rather than VCP counts because they believe that assumptions of the VCP method are almost always violated, leading to poor results, and because earlier field tests using ad hoc analysis methods gave variable and relatively poor results. We conducted the first field test of the VCP method where the exact density of a forest bird was known as part of re-establishing the 'Oma'o Myadestes obscurus in former range. All 'Oma'o in the new population were intensively monitored by radio telemetry so that the number present during four VCP censuses was known. Excluding the first census, when three of the four detections were of the same individual, differences in VCP density estimates ranged from -34% to +24% (mean 0%) even though =18 'Oma'o were detected per survey. We review critical assumptions of the VCP method and make recommendations for data analysis based on our experience with the method on Pacific islands.

Pacific Conservation Biology↗

East African weathering dynamics controlled by vegetation-climate feedbacks

Tropical weathering has important linkages to global biogeochemistry and landscape evolution in the East African rift. We disentangle the influences of climate and terrestrial vegetation on chemical weathering intensity and erosion at Lake Malawi using a long sediment record. Fossil pollen, microcharcoal, particle size, and mineralogy data affirm that the detrital clays accumulating in deep water within the lake are controlled by feedbacks between climate and hinterland forest composition. Particle-size patterns are also best explained by vegetation, through feedbacks with lake levels, wildfires, and erosion. We develop a new source-to-sink framework that links lacustrine sedimentation to hinterland vegetation in tropical rifts. Our analysis suggests that climate-vegetation interactions and their coupling to weathering/erosion could threaten future food security and has implications for accurately predicting petroleum play elements in continental rift basins.

Geology↗

Level II scour analysis for Bridge 32 (BRNATH00470032) on Town Highway 47, crossing Locust Creek, Barnard, Vermont

This report provides the results of a detailed Level II analysis of scour potential at structure BRNATH00470032 on town highway 47 crossing Locust Creek, Barnard, Vermont (figures 1–8). A Level II study is a basic engineering analysis of the site, including a quantitative analysis of stream stability and scour (U.S. Department of Transportation, 1993). Results of a Level I scour investigation also are included in Appendix E of this report. A Level I investigation provides a qualitative geomorphic characterization of the study site. Information on the bridge, gleaned from Vermont Agency of Transportation (VTAOT) files, was compiled prior to conducting Level I and Level II analyses and is found in Appendix D. The site is in the Green Mountain section of the New England physiographic province of central Vermont in the town of Barnard. The 6.26-mi2 drainage area is in a predominantly rural and forested basin. In the vicinity of the study site, the banks have dense woody vegetation coverage except for the upstream right bank, which is grass and brush covered. In the study area, Locust Creek has an incised, sinuous channel with a slope of approximately 0.029 ft/ft, an average channel top width of 44 ft., and an average channel depth of 5 ft. The predominant channel bed material is gravel and cobbles (D50 is 91.7 mm or 0.301 ft). The geomorphic assessment at the time of the Level I and Level II site visit on October 12, 1994, indicated that the reach was stable. The town highway 47 crossing of Locust Creek is a 28-ft-long, one-lane bridge consisting of one 25-foot span concrete slab superstructure (Vermont Agency of Transportation, written commun., August 23, 1994). The bridge is supported by vertical, concrete abutments with concrete wingwalls. The channel is skewed approximately 10 degrees to the opening while the opening-skew-to-roadway is 35 degrees. The scour protection measures at the site were type-2 stone fill (less than 36 inches diameter) on the banks upstream, the upstream wingwalls, the downstream right wingwall, and the downstream right bank. The downstream left wingwall and left bank are protected with type-3 stone fill (less than 48 inches diameter). Additional details describing conditions at the site are included in the Level II Summary and Appendices D and E. Scour depths and rock rip-rap sizes were computed using the general guidelines described in Hydraulic Engineering Circular 18 (Richardson and others, 1993). Total scour at a highway crossing is comprised of three components: 1) long-term streambed degradation; 2) contraction scour (due to accelerated flow caused by a reduction in flow area at a bridge) and; 3) local scour (caused by accelerated flow around piers and abutments). Total scour is the sum of the three components. Equations are available to compute depths for contraction and local scour and a summary of these computed results follow. Contraction scour for all modelled flows ranged from 1.4 to 2.2 feet. The worst-case contraction scour occurred at the 500-year discharge. Abutment scour ranged from 10.3 to 15.0 feet. The worst-case abutment scour also occurred at the 500-year discharge. Additional information on scour depths and depths to armoring are included in the section titled “Scour Results”. Scoured-streambed elevations, based on the calculated depths, are presented in tables 1 and 2. A cross-section of the scour computed at the bridge is presented in figure 8. Scour depths were calculated assuming an infinite depth of erosive material and a homogeneous particle-size distribution. It is generally accepted that the Froehlich equation (abutment scour) gives “excessively conservative estimates of scour depths” (Richardson and others, 1993, p. 47). Usually, computed scour depths are evaluated in combination with other information including (but not limited to) historical performance during flood events, the geomorphic stability assessment, existing scour protection measures, and the results of the hydraulic analyses. Therefore, scour depths adopted by VTAOT may differ from the computed values documented herein.

Vermont↗

Level II scour analysis for Bridge 1 (CANATH00010001) on Town Highway 1, crossing Halls Stream, Canaan, Vermont

This report provides the results of a detailed Level II analysis of scour potential at structure CANATH00010001 on town highway 1 crossing Halls Stream, Canaan, Vermont (figures 1–8). A Level II study is a basic engineering analysis of the site, including a quantitative analysis of stream stability and scour (U.S. Department of Transportation, 1993). A Level I study is included in Appendix E of this report. A Level I study provides a qualitative geomorphic characterization of the study site. Information on the bridge, gleaned from Vermont Agency of Transportation (VTAOT) files, was compiled prior to conducting Level I and Level II analyses and can be found in Appendix D. The site is in the White Mountain section of the New England physiographic province of northeastern Vermont in the town of Canaan. The 89.5-mi2 drainage area is in a predominantly rural and forested basin. In the vicinity of the study site, the banks have tree, shrub and brush, and grass vegetation coverage. In the study area, Halls Stream has a sinuous channel with a slope of approx-imately 0.0012 ft/ft, an average channel top width of 109 ft and an average channel depth of 4 ft. The predominant channel bed materials are sand and gravel (D50 is 5.03 mm or 0.0165 ft). The geomorphic assessment at the time of the Level I and Level II site visit on October 27, 1994, indicated that the reach was laterally unstable. The lateral instability was evident due to a wide point-bar and cut-banks with undermining of bank material, slumping, fallen bank vegetation evident in the upstream channel. The town highway 1 crossing of Halls Stream is a 99-ft-long, two-lane bridge consisting of one 33-foot and two 31-foot concrete T-beam spans (Vermont Agency of Transportation, written communication, August 5, 1994). The bridge is supported by vertical, concrete abutments with spill-through embankments in front of each abutment wall. The channel is skewed approximately 10 degrees to the opening while the opening-skew-to-roadway is zero degrees. There are two piers in the channel at this site. Field notes and the channel survey at the bridge indicate that the streambed elevation is higher on the downstream right sides of each pier and lower on the downstream left sides. This asymmetrical streambed condition suggests a flow attack angle may influence scour on each pier. Furthermore, field observations suggest that the flow attack angle is higher for the right pier (pier 2) than the left pier (pier 1). The scour protection measures at the site were type-2 stone fill (less than 36 inches diameter) on both upstream banks and both downstream road embankments. Type-3 stone fill (less than 48 inches diameter) was found on the spill-through slopes of each abutment and both downstream banks. The stone fill protection on the spill-through embankment of the right abutment was noted as slumped with some of the fill material evident in the channel immediately downstream of the bridge. Additional details describing conditions at the site are included in the Level II Summary and Appendices D and E. Scour depths and rock rip-rap sizes were computed using the general guidelines described in Hydraulic Engineering Circular 18 (Richardson and others, 1995). Total scour at a highway crossing is comprised of three components: 1) long-term streambed degradation; 2) contraction scour (due to accelerated flow caused by a reduction in flow area at a bridge) and; 3) local scour (caused by accelerated flow around piers and abutments). Total scour is the sum of the three components. Equations are available to compute depths for contraction and local scour and a summary of the results of these computations follows. Contraction scour for all modelled flows ranged from 8.0 to 8.8 ft. The worst-case contraction scour occurred at the 500-year discharge. Abutment scour ranged from 8.9 to 17.3 ft. The worst-case abutment scour occurred at the 500-year discharge. For the two piers, scour ranged from 11.1 to 15.8. The worst-case pier scour occurred for pier2 at the incipient overtopping discharge. Additional information on scour depths and depths to armoring are included in the section titled “Scour Results”. Scoured-streambed elevations, based on the calculated scour depths, are presented in tables 1 and 2. A cross-section of the scour computed at the bridge is presented in figure 8. Scour depths were calculated assuming an infinite depth of erosive material and a homogeneous particle-size distribution. It is generally accepted that the Froehlich equation (abutment scour) gives “excessively conservative estimates of scour depths” (Richardson and others, 1995, p. 47). Usually, computed scour depths are evaluated in combination with other information including (but not limited to) historical performance during flood events, the geomorphic stability assessment, existing scour protection measures, and the results of the hydraulic analyses. Therefore, scour depths adopted by VTAOT may differ from the computed values documented herein.

Vermont↗

Hydrogeology and Water Quality of the Pepacton Reservoir Watershed in Southeastern New York. Part 3. Responses of Stream Base-Flow Chemistry to Hydrogeologic Factors and Nonpoint-Sources of Contamination

Base-flow samples were collected seasonally from 20 small streams in the 372-square-mile Pepacton Reservoir watershed to evaluate the effects of hydrogeologic factors and nonpoint sources of contamination on the chemical composition of ground-water discharge. The reservoir provides part of New York City?s water supply. The subbasins represented one of three general types of land use, each with at least 45 percent forested area (mostly on the hillsides): farmed (dairy) land, formerly farmed land with low-density residential development, or forested land with little or no development. The subbasin areas ranged from 0.38 to 10.23 square miles. All streams were sampled in December 2000 and in May, July, and October 2001. Three of the sites were designated as landuse- index sites and were sampled as many as five additional times during the study. No samples exceeded state or federal drinking-water standards for chloride, sodium, nitrate, orthophosphate, herbicides, or herbicide degradates. The chemical composition of base-flow samples was classified into major-ion water types, which were broadly defined as naturally occurring and road-salt-affected water types. About one-third of the base-flow samples were roadsalt- affected types. Natural water types were differentiated as dilute or evolved. Dilute waters have bicarbonate and sulfate as dominant anions and evolved waters have only bicarbonate as a dominant anion. Dilute water types indicate relatively short ground-water residence times or contact with unreactive aquifer material. Evolved waters have either longer ground-water residence time or contact with more reactive aquifer material than dilute ground waters. The larger subbasins with wider valley-bottom areas were more likely to have evolved water types than small subbasins with little floodplain development. Positive correlations between selected constituents and the intensity of nonpoint sources emphasize the connection between land use, shallow ground-water quality, and stream base-flow water quality. Chloride and sodium, which are relatively conservative constituents, showed strong linear relations with annual estimates of road-salt application during all four sampling periods. Nonconservative constituents, such as the nutrients nitrate and orthophosphate, showed linear relations with manure production rate among farmed basins, but only at specific times of the year because of losses through biologic activity. Nitrate showed the strongest relation in winter because losses to biological activity were at a minimum. Orthophosphate showed the strongest relation in early summer, when hydrologic and chemical conditions appear to favor release from sediments. Atmospheric nitrogen deposition is an additional source of nitrogen that can be released from mature or stressed forested basins. Detections of herbicides (atrazine, metolachlor, simazine) and herbicide degradates ( Metolachlor ESA, alachlor ESA, deethylatrazine) in base flow were closely correlated with subbasins in which corn was grown during the study. Atrazine was detected at the farmed index site only in early summer, after application and two rain storms. This detection corresponded to the peak orthophosphate concentration. In contrast, metolachlor ESA was detected in nearly all farmedindex- subbasin samples and peaked in late summer, when percent base-flow contributions from farmed valley-bottom areas were likely highest. The implications of this study are that seasonal and more frequent base-flow surveys of water chemistry from small stream basins can help refine the understanding of local hydrogeologic systems and define the effects of nonpointsource contamination on base-flow water quality. The concentration of most nonpoint sources in valley-bottom or lower-hillside areas helped indicate the relative contributions of water from hillside and valley-bottom areas at different times of year. The positive correlations between the intensity of nonpoint-source activities and nonpoint-source constituents in base flow underscores the link between land use (nonpoint sources), ground-water quality, and surface-water quality.

Scientific Investigations Report↗

Competitive interactions and resource partitioning between northern spotted owls and barred owls in western Oregon

The federally threatened northern spotted owl ( Strix occidentalis caurina ) is the focus of intensive conservation efforts that have led to much forested land being reserved as habitat for the owl and associated wildlife species throughout the Pacific Northwest of the United States. Recently, however, a relatively new threat to spotted owls has emerged in the form of an invasive competitor: the congeneric barred owl ( S. varia ). As barred owls have rapidly expanded their populations into the entire range of the northern spotted owl, mounting evidence indicates that they are displacing, hybridizing with, and even killing spotted owls. The range expansion by barred owls into western North America has made an already complex conservation issue even more contentious, and a lack of information on the ecological relationships between the 2 species has hampered recovery efforts for northern spotted owls. We investigated spatial relationships, habitat use, diets, survival, and reproduction of sympatric spotted owls and barred owls in western Oregon, USA, during 2007–2009. Our overall objective was to determine the potential for and possible consequences of competition for space, habitat, and food between these previously allopatric owl species. Our study included 29 spotted owls and 28 barred owls that were radio-marked in 36 neighboring territories and monitored over a 24-month period. Based on repeated surveys of both species, the number of territories occupied by pairs of barred owls in the 745-km 2 study area (82) greatly outnumbered those occupied by pairs of spotted owls (15). Estimates of mean size of home ranges and core-use areas of spotted owls (1,843 ha and 305 ha, respectively) were 2–4 times larger than those of barred owls (581 ha and 188 ha, respectively). Individual spotted and barred owls in adjacent territories often had overlapping home ranges, but interspecific space sharing was largely restricted to broader foraging areas in the home range with minimal spatial overlap among core-use areas. We used an information-theoretic approach to rank discrete-choice models representing alternative hypotheses about the influence of forest conditions, topography, and interspecific interactions on species-specific patterns of nighttime resource selection. Spotted owls spent a disproportionate amount of time foraging on steep slopes in ravines dominated by old (>120 yr) conifer trees. Barred owls used available forest types more evenly than spotted owls, and were most strongly associated with patches of large hardwood and conifer trees that occupied relatively flat areas along streams. Spotted and barred owls differed in the relative use of old conifer forest (greater for spotted owls) and slope conditions (steeper slopes for spotted owls), but we found no evidence that the 2 species differed in their use of young, mature, and riparian-hardwood forest types. Mean overlap in proportional use of different forest types between individual spotted owls and barred owls in adjacent territories was 81% (range = 30–99%). The best model of habitat use for spotted owls indicated that the relative probability of a location being used was substantially reduced if the location was within or in close proximity to a core-use area of a barred owl. We used pellet analysis and measures of food-niche overlap to determine the potential for dietary competition between spatially associated pairs of spotted owls and barred owls. We identified 1,223 prey items from 15 territories occupied by spotted owls and 4,299 prey items from 24 territories occupied by barred owls. Diets of both species were dominated by nocturnal mammals, but diets of barred owls included many terrestrial, aquatic, and diurnal prey species that were rare or absent in diets of spotted owls. Northern flying squirrels ( Glaucomys sabrinus ), woodrats ( Neotoma fuscipes , N. cinerea ), and lagomorphs ( Lepus americanus , Sylvilagus bachmani ) were primary prey for both owl species, accounting for 81% and 49% of total dietary biomass for spotted owls and barred owls, respectively. Mean dietary overlap between pairs of spotted and barred owls in adjacent territories was moderate (42%; range = 28–70%). Barred owls displayed demographic superiority over spotted owls; annual survival probability of spotted owls from known-fate analyses (0.81, SE = 0.05) was lower than that of barred owls (0.92, SE = 0.04), and pairs of barred owls produced an average of 4.4 times more young than pairs of spotted owls over a 3-year period. We found a strong, positive relationship between seasonal (6-month) survival probabilities of both species and the proportion of old (>120 yr) conifer forest within individual home ranges, which suggested that availability of old forest was a potential limiting factor in the competitive relationship between these 2 species. The annual number of young produced by spotted owls increased linearly with increasing distance from a territory center of a pair of barred owls, and all spotted owls that attempted to nest within 1.5 km of a nest used by barred owls failed to successfully produce young. We identified strong associations between the presence of barred owls and the behavior and fitness potential of spotted owls, as shown by changes in movements, habitat use, and reproductive output of spotted owls exposed to different levels of spatial overlap with territorial barred owls. When viewed collectively, our results support the hypothesis that interference competition with barred owls for territorial space can constrain the availability of critical resources required for successful recruitment and reproduction of spotted owls. Availability of old forests and associated prey species appeared to be the most strongly limiting factors in the competitive relationship between these species, indicating that further loss of these conditions can lead to increases in competitive pressure. Our findings have broad implications for the conservation of spotted owls, as they suggest that spatial heterogeneity in vital rates may not arise solely because of differences among territories in the quality or abundance of forest habitat, but also because of the spatial distribution of a newly established competitor. Experimental removal of barred owls could be used to test this hypothesis and determine whether localized control of barred owl numbers is an ecologically practical and socio-politically acceptable management tool to consider in conservation strategies for spotted owls.

Oregon↗

Ground Water Atlas of the United States: Segment 2, Arizona, Colorado, New Mexico, Utah

This chapter of the Ground Water Atlas of the United States describes the aquifers in Arizona, Colorado, New Mexico, and Utah. These four States, which comprise Segment 2 of this Atlas, are located in the Southwestern United States and extend from the rolling grasslands of the Great Plains on the east across the Rocky Mountains and Continental Divide to the desert basins of the Southwest. The 425,000-square-mile area ranges in altitude from about 14,400 feet above sea level in the Rocky Mountains of Colorado to about 100 feet near the lower Colorado River in southwestern Arizona. All the ground water in Segment 2 ultimately is derived from infiltration of precipitation, which varies considerably with the altitude and topography of the area. The Great Plains Physiographic Province of the Central United States extends into eastern Colorado and New Mexico (fig. 1), where flat to rolling prairie (fig. 2) with scattered hills and bluffs gradually rises westward to 5,000 to 7,000 feet above sea level and abruptly gives way to the frontal ranges of the Rocky Mountains in the Southern Rocky Mountain and Basin and Range Physiographic Provinces. West of the frontal ranges in Colorado and northern New Mexico are additional and higher mountain ranges generally oriented north-south but with many spurs and extensions oriented in other directions. The many ranges of the Rocky Mountains are separated by valleys and high mountain parks (fig. 3). Colorado contains about three-fourths of the Nation's land area above 10,000 feet and has 53 mountain peaks higher than 14,000 feet. Most of these high peaks are located near the Continental Divide (fig. 1), which extends approximately north-south through central Colorado and western New Mexico. The altitude of the divide decreases in southern New Mexico to less than 4,500 feet in a few areas. Farther westward, the mountains are less prevalent and are interspersed with broad structural basins. These basins and the broad valleys of the middle Colorado River and its tributaries form the irregular intermontane topography of the Colorado Plateaus Physiographic Province (fig. 4). Plateaus and high mesas are formed where the surface has been dissected by rugged canyons carved by the Colorado River and its tributaries (fig. 5). The largest of these canyons-the Grand Canyon-extends about 220 miles southwestward from the mouth of the Little Colorado River in Arizona and ranges from 4 to 18 miles in width and from 2,700 to 5,700 feet in depth below the rim. Small mountain ranges and intervening broad desert valleys of the Basin and Range Physiographic Province are prevalent to the west and south of the Colorado Plateaus in western Utah, southern Arizona, and southern New Mexico (fig. 1). These mountain ranges generally protrude 3,000 to 6,000 feet above the surrounding valley floor (fig. 6) and commonly extend from 20 to 50 miles in a north or northwesterly direction. Small parts of the Middle Rocky Mountains and Wyoming Basin Physiographic Provinces extend into northwestern Colorado and northeastern Utah (fig. 1). The topography, geology, and hydrology of the two areas are described in Chapter 1 of this Atlas. Four of the Nation's major river systems have headwaters in the mountainous areas of Segment 2. The South Platte River of the Missouri River system drains the eastern slope of northern Colorado; the Arkansas River and its tributary, the Canadian River, drain southeastern Colorado and northeastern New Mexico; the Rio Grande and its tributary, the Pecos River, drain south-central Colorado and central New Mexico; and the Colorado River and its tributaries drain Arizona, eastern Utah, Northwestern New Mexico, and western Colorado (fig. 1). Western Utah is drained by numerous streams that terminate in local desert basins, the Great Salt Lake, or other local lakes and reservoirs. Because the Great Salt Lake lies in the Great Basin, which is the largest closed basin in North America, it has no outlet to the sea. The salinity of the lake water is about 20 percent or about 6 times the salinity of seawater. Most of Segment 2 is sparsely populated. The average population density of counties is less than 8 persons per square mile in about 65 percent of the four-State area (fig. 7). Population densities range from less than 0.5 person per square mile in a few rural counties to more than 4 ,000 persons per square mile in populous urban areas. The 1990 population of the four States was about 10 million; almost 70 percent of this population was in Arizona and Colorado. Most land in Segment 2 is undeveloped forest grassland, or desert shrubland, much of which is used for livestock grazing. Land used for production of commercial crops primarily is in eastern Colorado and eastern New Mexico.

Arizona, Colorado, New Mexico, Utah↗

Trees have similar growth responses to first-entry fires and reburns following long-term fire exclusion

Managing fire ignitions for resource benefit decreases fuel loads and reduces the risk of high-severity fire in fire-suppressed dry conifer forests. However, the reintroduction of low-severity wildfire can injure trees, which may decrease their growth after fire. Post-fire growth responses could change from first-entry fires to reburns, as first-entry fires reduce fuel loads and the vulnerability among trees to fire effects, which may result in trees sustaining less damage during reburns. To determine whether trees had growth responses that varied from first-entry fires to reburns, we cored 87 ponderosa pine trees in the Gila Wilderness, New Mexico, USA that experienced 3–5 fires between 1950 and 2012 following long-term fire-exclusion and 67 unburned control trees from the Gila and Apache-Sitgreaves National Forests. We assessed tree growth response to fire by comparing tree-ring growth among burned and unburned trees from two years before to two years after fires. We compared growth between burned and unburned trees using a bootstrapping procedure to calculate annual median tree-ring width index values with 95 % confidence intervals. We compared post-fire growth after first-entry fires and reburns following long-term fire-exclusion. Burned trees had similar growth responses following first-entry fires and reburns, with lower growth during the fire year through two years post-fire compared to unburned controls. Burned tree growth returned to expected rates following these immediate post-fire growth reductions. Interestingly, trees had lower growth during the year before and the year of reburns compared to the first-entry fire, reflecting greater aridity before reburns. Greater aridity may have contributed to larger-than-expected growth reductions following reburns, which could explain similar growth responses to first-entry fires and reburns. Our results indicate that trees had consistent short-term growth responses to low-severity fires following long-term fire-exclusion. As trees retained vigor after multiple fires, managing fires for resource benefit is an effective approach to reduce the likelihood of high-severity fire without long-term negative effects on tree growth.

Forest Ecology and Management↗

Assessment of fire behavior and management options in subalpine vegetation on Mauna Kea, Hawai'i

Fire is a major threat to habitat for the endangered Palila (Loxioides bailleui) within subalpine vegetation on Mauna Kea volcano, Hawai‘i. The presence of large amounts of fine fuel from grasses, dry climate, and human ignition sources produces a significant risk of wildfire in this area year-round. The purpose of this report is to provide information on fuels and potential fire behavior that will contribute to fire management of Palila habitat. Recommended actions will contribute to the conservation of these native forests and facilitate restoration in degraded areas. To assess the effects of grass invasion on fuel conditions and potential fire danger, we quantified vegetation and fuels across an elevation gradient from grasslands into sub-alpine forests on the west slope of Mauna Kea. Our results indicated that grass cover was reduced under tree canopy in plots below ~2,500 m elevation, but at higher elevations grass cover was higher under trees than in the open. However, tree canopy cover below 2,500 m elevation was not high enough overall (~25% on average) to result in significant reductions in fine fuels at the landscape level. Sampling directly under and away from tree crowns at multiple elevations suggested that below ~2,500 m, the presence of tree canopy cover can reduce grass fuels significantly. Furthermore, moisture content of live surface fuels was increased under tree canopy compared with open areas. These results suggest that restoration of forest cover may have the potential to alter grass fuels in ways that decrease the threat of fire in some subalpine forests. Fire behavior estimates based on fuel data from grasslands, mixed forest and māmane forest indicated the need for fuelbreaks of at least 20-30 m to limit fire spread in most areas. In many cases, breaks as wide as 40 m are required to limit fire spread risk under extreme weather conditions. Based on our fuels data and fire behavior predictions, recommended actions include: (1) construction of new or expansion of existing fuelbreaks to immediately reduce fire risk to the most sensitive areas adjacent to the core Palila population on the southwest slope and the translocated Palila population on the north slope of Mauna Kea, (2) enhancement of forest restoration activities to increase fuel moisture and reduce grass fuel loads (3) installation of water sources (diptanks) in both areas to decrease firefighter response time, and (4) increased public education and awareness with regard to fire danger on Mauna Kea.

Hawaii↗

Landscape correlates of breeding bird richness across the United States mid-Atlantic region

Using a new set of landscape indicator data generated by the U.S.EPA, and a comprehensive breeding bird database from the National Breeding Bird Survey, we evaluated associations between breeding bird richness and landscape characteristics across the entire mid-Atlantic region of the United States. We evaluated how these relationships varied among different groupings (guilds) of birds based on functional, structural, and compositional aspects of individual species demographics. Forest edge was by far the most important landscape attribute affecting the richness of the lumped specialist and generalist guilds; specialist species richness was negatively associated with forest edge and generalist richness was positively associated with forest edge. Landscape variables (indicators) explained a greater proportion of specialist species richness than the generalist guild (46% and 31%, respectively). The lower value in generalists may reflect freer-scale distributions of open habitat that go undetected by the Landsat satellite, open habitats created by roads (the areas from which breeding bird data are obtained), and the lumping of a wide variety of species into the generalist category. A further breakdown of species into 16 guilds showed considerable variation in the response of breeding birds to landscape conditions; forest obligate species had the strongest association with landscape indicators measured in this study (55% of the total variation explained) and forest generalists and open ground nesters the lowest (17% of the total variation explained). The variable response of guild species richness to landscape pattern suggests that one must consider species' demographics when assessing the consequences of landscape change on breeding birds.Using a new set of landscape indicator data generated by the U.S. EPA, and a comprehensive breeding bird database from the National Breeding Bird Survey, we evaluated associations between breeding bird richness and landscape characteristics across the entire mid-Atlantic region of the United States. We evaluated how these relationships varied among different groupings (guilds) of birds based on functional, structural, and compositional aspects of individual species demographics. Forest edge was by far the most important landscape attribute affecting the richness of the lumped specialist and generalist guilds; specialist species richness was negatively associated with forest edge and generalist richness was positively associated with forest edge. Landscape variables (indicators) explained a greater proportion of specialist species richness than the generalist guild (46% and 31%, respectively). The lower value in generalists may reflect finer-scale distributions of open habitat that go undetected by the Landsat satellite, open habitats created by roads (the areas from which breeding bird data are obtained), and the lumping of a wide variety of species into the generalist category. A further breakdown of species into 16 guilds showed considerable variation in the response of breeding birds to landscape conditions; forest obligate species had the strongest association with landscape indicators measured in this study (55% of the total variation explained) and forest generalists and open ground nesters the lowest (17% of the total variation explained). The variable response of guild species richness to landscape pattern suggests that one must consider species' demographics when assessing the consequences of landscape change on breeding birds.

Conference Paper↗

Linking plant and animal functional diversity with an experimental community restoration in a Hawaiian lowland wet forest

Testing how plant restoration influences animal taxonomic and functional diversity can shift restoration projects beyond mainly plant community considerations. We incorporated multi-trophic interactions into restoration by describing an ongoing functional trait-based restoration experiment in Hawaiian lowland tropical wet forest (Liko Nā Pilina Experiment), where litter arthropods are examined from a functional perspective thereby linking plants and higher trophic levels. We hypothesized that (1) communities with greater plant functional trait diversity would have cascading effects through food webs, increasing animal diversity and network complexity, and (2) increases in animal species and network complexity would be stronger for restoration efforts in plant communities with more complementary functional traits than those with more redundant traits. We examined experimental treatments of planted communities with the same species richness but with different plant functional trait profiles based on (1) rates of expected carbon turnover (slow or moderate), and (2) the similarity of their functional trait measurements (redundant or complementary), as determined by functional dispersion calculations. Initial data on arthropod communities and leaf litter decomposition rates revealed linkages between plant functional traits and arthropod community diversity. Overall, we argue that a more comprehensive evaluation of restoration accounts for both functional diversity and the multi-trophic nature of animal and plant communities. Developing restoration projects based on plant functional traits that influence both plant and invertebrate species provides a new paradigm, and the incorporation of both native and non-native (but non-invasive) plants shows promise in restoring ecosystem function in disturbed lowland tropical forests.

Hawaii↗

Second U.S. Geological Survey Wildland Fire Workshop: Los Alamos, New Mexico, October 31-November 3, 2000

In the past decade, fire-management policies have shifted to embrace the reduction of long-term building of excessive fuel levels in the Nation's forests and rangelands. In 1995, the Departments of Agriculture and the Interior issued the Federal Wildland-Fire Policy Statement (Appendix A) identifying the critical role that fire can play in the management of forests and rangelands. As a result of revised policy, Federal agencies have increased the acres treated to reduce fire risk through "prescribed" fires and thinning techniques from 500,000 acres in 1995 to more than 2.4 million acres in 2001.

Open-File Report↗