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Historical rock falls in Yosemite National Park, California (1857-2011)

Inventories of rock falls and other types of landslides are valuable tools for improving understanding of these events. For example, detailed information on rock falls is critical for identifying mechanisms that trigger rock falls, for quantifying the susceptibility of different cliffs to rock falls, and for developing magnitude-frequency relations. Further, inventories can assist in quantifying the relative hazard and risk posed by these events over both short and long time scales. This report describes and presents the accompanying rock fall inventory database for Yosemite National Park, California. The inventory database documents 925 events spanning the period 1857–2011. Rock falls, rock slides, and other forms of slope movement represent a serious natural hazard in Yosemite National Park. Rock-fall hazard and risk are particularly relevant in Yosemite Valley, where glacially steepened granitic cliffs approach 1 km in height and where the majority of the approximately 4 million yearly visitors to the park congregate. In addition to damaging roads, trails, and other facilities, rock falls and other slope movement events have killed 15 people and injured at least 85 people in the park since the first documented rock fall in 1857. The accompanying report describes each of the organizational categories in the database, including event location, type of slope movement, date, volume, relative size, probable trigger, impact to humans, narrative description, references, and environmental conditions. The inventory database itself is contained in a Microsoft Excel spreadsheet (Yosemite_rock_fall_database_1857-2011.xlsx). Narrative descriptions of events are contained in the database, but are also provided in a more readable Adobe portable document format (pdf) file (Yosemite_rock_fall_database_narratives_1857-2011.pdf) available for download separate from the database.

California

Staircase Falls Rockfall on December 26, 2003, and Geologic Hazards at Curry Village, Yosemite National Park, California

Since 1857, several hundred rockfalls, rockslides, and debris flows have been observed in Yosemite National Park. At 12:45 a.m. on December 26, 2003, a severe winter storm triggered a rockfall west of Glacier Point in Yosemite Valley. Rock debris moved quickly eastward down Staircase Falls toward Curry Village. As the rapidly moving rock mass reached talus at the bottom of Staircase Falls, smaller pieces of flying rock penetrated occupied cabins. Physical characterization of the rockfall site included rockfall volume, joint patterns affecting initial release of rock and the travel path of rockfall, factors affecting weathering and weakening of bedrock, and hydrology affecting slope stability within joints. Although time return intervals are not predictable, a three-dimensional rockfall model was used to assess future rockfall potential and risk. Predictive rockfall and debris-flow methods suggest that landslide hazards beneath these steep cliffs extend farther than impact ranges defined from surface talus in Yosemite Valley, leaving some park facilities vulnerable.

Open-File Report

Investigation and hazard assessment of the 2003 and 2007 Staircase Falls rock falls, Yosemite National Park, California, USA

Since 1857 more than 600 rock falls, rock slides, debris slides, and debris flows have been documented in Yosemite National Park, with rock falls in Yosemite Valley representing the majority of the events. On 26 December 2003, a rock fall originating from west of Glacier Point sent approximately 200 m 3 of rock debris down a series of joint-controlled ledges to the floor of Yosemite Valley. The debris impacted talus near the base of Staircase Falls, producing fragments of flying rock that struck occupied cabins in Curry Village. Several years later on 9 June 2007, and again on 26 July 2007, smaller rock falls originated from the same source area. The 26 December 2003 event coincided with a severe winter storm and was likely triggered by precipitation and/or frost wedging, but the 9 June and 26 July 2007 events lack recognizable triggering mechanisms. We investigated the geologic and hydrologic factors contributing to the Staircase Falls rock falls, including bedrock lithology, weathering, joint spacing and orientations, and hydrologic processes affecting slope stability. We improved upon previous geomorphic assessment of rock-fall hazards, based on a shadow angle approach, by using STONE, a three-dimensional rock-fall simulation computer program. STONE produced simulated rock-fall runout patterns similar to the mapped extent of the 2003 and 2007 events, allowing us to simulate potential future rock falls from the Staircase Falls detachment area. Observations of recent rock falls, mapping of rock debris, and simulations of rock fall runouts beneath the Staircase Falls detachment area suggest that rock-fall hazard zones extend farther downslope than the extent previously defined by mapped surface talus deposits.

Natural Hazards and Earth System Sciences

Yosemite Falls - A new perspective

Yosemite Falls with their spectacular drop of 2,425 feet (including the Upper Fall, middle cascades, and Lower Fall) are world-renowned and an icon for Yosemite Valley (Fig. 1). They are truly unmatched and were recognized as such early on.

California

Habitat suitability of patch types: a case study of the Yosemite toad

Understanding patch variability is crucial in understanding the spatial population structure of wildlife species, especially for rare or threatened species. We used a well-tested maximum entropy species distribution model (Maxent) to map the Yosemite toad (Anaxyrus (= Bufo) canorus) in the Sierra Nevada mountains of California. Twenty-six environmental variables were included in the model representing climate, topography, land cover type, and disturbance factors (e.g., distances to agricultural lands, fire perimeters, and timber harvest areas) throughout the historic range of the toad. We then took a novel approach to the study of spatially structured populations by applying the species-environmental matching model separately for 49 consistently occupied sites of the Yosemite toad compared to 27 intermittently occupied sites. We found that the distribution of the entire population was highly predictable (AUC = 0.95±0.03 SD), and associated with low slopes, specific vegetation types (wet meadow, alpine-dwarf shrub, montane chaparral, red fir, and subalpine conifer), and warm temperatures. The consistently occupied sites were also associated with these same factors, and they were also highly predictable (AUC = 0.95±0.05 SD). However, the intermittently occupied sites were associated with distance to fire perimeter, a slightly different response to vegetation types, distance to timber harvests, and a much broader set of aspect classes (AUC = 0.90±0.11 SD). We conclude that many studies of species distributions may benefit by modeling spatially structured populations separately. Modeling and monitoring consistently-occupied sites may provide a realistic snapshot of current species-environment relationships, important climatic and topographic patterns associated with species persistence patterns, and an understanding of the plasticity of the species to respond to varying climate regimes across its range. Meanwhile, modeling and monitoring of widely dispersing individuals and intermittently occupied sites may uncover environmental thresholds and human-related threats to population persistence.

Frontiers of Earth Science

Triggering mechanisms and depositional rates of postglacial slope-movement processes in the Yosemite Valley, California

We examined information collected from 395 reports of slope-movement events during about the past 150 years in Yosemite National Park, central Sierra Nevada, California, to identify the most prevalent types of slope movements and their triggering mechanisms. Rock slides and rock falls have been more numerous than debris slides, debris flows, and miscellaneous slumps. Rock falls have produced the largest cumulative volume of deposits. About half of slope movements had unreported or unrecognized triggering events. Earthquakes and rain storms individually accounted for the greatest cumulative volumes of deposits from recognized triggers of all types of historical slope movements; snowmelt, human activities and freeze-thaw conditions accounted for only a small proportion of the volumes from reported triggers. A comparison of the historical and postglacial average annual rates of deposition from slope-movement processes in a portion of the Yosemite Valley indicates that, during the period 1851-1992, slope-movement processes have been producing about half the average rate of deposits than during the past 15,000 years.

Geomorphology

Twentieth-century decline of large-diameter trees in Yosemite National Park, California, USA

Studies of forest change in western North America often focus on increased densities of small-diameter trees rather than on changes in the large tree component. Large trees generally have lower rates of mortality than small trees and are more resilient to climate change, but these assumptions have rarely been examined in long-term studies. We combined data from 655 historical (1932-1936) and 210 modern (1988-1999) vegetation plots to examine changes in density of large-diameter trees in Yosemite National Park (3027 km 2 ). We tested the assumption of stability for large-diameter trees, as both individual species and communities of large-diameter trees. Between the 1930s and 1990s, large-diameter tree density in Yosemite declined 24%. Although the decrease was apparent in all forest types, declines were greatest in subalpine and upper montane forests (57.0% of park area), and least in lower montane forests (15.3% of park area). Large-diameter tree densities of 11 species declined while only 3 species increased. Four general patterns emerged: (1) Pinus albicaulis, Quercus chrysolepis, and Quercus kelloggii had increases in density of large-diameter trees occur throughout their ranges; (2) Pinus jeffreyi, Pinus lambertiana, and Pinus ponderosa, had disproportionately larger decreases in large-diameter tree densities in lower-elevation portions of their ranges; (3) Abies concolor and Pinus contorta, had approximately uniform decreases in large-diameter trees throughout their elevational ranges; and (4) Abies magnifica, Calocedrus decurrens, Juniperus occidentalis, Pinus monticola, Pseudotsuga menziesii, and Tsuga mertensiana displayed little or no change in large-diameter tree densities. In Pinus ponderosa-Calocedrus decurrens forests, modern large-diameter tree densities were equivalent whether or not plots had burned since 1936. However, in unburned plots, the large-diameter trees were predominantly A. concolor, C. decurrens, and Q. chrysolepis, whereas P. ponderosa dominated the large-diameter component of burned plots. Densities of large-diameter P. ponderosa were 8.1 trees ha -1 in plots that had experienced fire, but only 0.5 trees ha -1 in plots that remained unburned. ?? 2009 Elsevier B.V. All rights reserved.

Forest Ecology and Management

Quantifying 40 years of rockfall activity in Yosemite Valley with historical Structure-from-Motion photogrammetry and terrestrial laser scanning

Rockfalls and rockslides are often dominant geomorphic processes in steep bedrock landscapes, but documenting their occurrence can be challenging, requiring frequent monitoring and well resolved spatial data. Repeat application of remote sensing methods such as Terrestrial Laser Scanning (TLS) and Structure-from-Motion (SfM) photogrammetry can detect even very small rockfalls, but typically these acquisitions span only years and may not record rockfall activity representative of longer-term rates of cliff erosion. Inventory databases can extend rockfall records, but are commonly incomplete and prone to observation bias. We employed TLS and SfM on two adjacent cliffs (El Capitan and Middle Brother) in Yosemite Valley, integrating semi-annual data collections from 2010 to 2017 with “historical” (archival) SfM models derived from oblique photographs taken in 1976. Comparing the 1976 SfM models against more recent data allows for more accurate and precise rockfall detection and volume measurement over a 40-year period. Change detection indicates that 235 rockfalls occurred from the two cliffs, more than twice as many events as are recorded in Yosemite's inventory database. Although individual rockfall volumes reported in the inventory database vary from those measured by SfM-TLS, reported cumulative volumes are similar to measured volumes, likely because the large-volume events that account for most of the cumulative volume tend to be widely observed and well-documented. Volume-frequency relationships indicate that the cliffs erode predominantly by less frequent, larger-volume rockfalls, at rates of 0.9 to 1.7 mm/yr. Our study demonstrates how integrated SfM and TLS measurements, especially utilizing SfM models derived from historical imagery, allow detection and quantification of rockfalls spanning several decades, complementing and improving inventory databases, informing rockfall hazard assessment, and providing longer-term rates of cliff erosion.

California

Quantifying the fire regime distributions for severity in Yosemite National Park, California, USA

This paper quantifies current fire severity distributions for 19 different fire-regime types in Yosemite National Park, California, USA. Landsat Thematic Mapper remote sensing data are used to map burn severity for 99 fires (cumulatively over 97 000 ha) that burned in Yosemite over a 20-year period. These maps are used to quantify the frequency distributions of fire severity by fire-regime type. A classification is created for the resultant distributions and they are discussed within the context of four vegetation zones: the foothill shrub and woodland zone; the lower montane forest zone; the upper montane forest zone and the subalpine forest zone. The severity distributions can form a building block from which to discuss current fire regimes across the Sierra Nevada in California. This work establishes a framework for comparing the effects of current fires on our landscapes with our notions of how fires historically burned, and how current fire severity distributions differ from our desired future conditions. As this process is refined, a new set of information will be available to researchers and land managers to help understand how fire regimes have changed from the past and how we might attempt to manage them in the future.

California

Using landscape genomics to delineate future adaptive potential for climate change in the Yosemite toad (Anaxyrus canorus)

An essential goal in conservation biology is delineating population units that maximize the probability of species persisting into the future and adapting to future environmental change. However, future-facing conservation concerns are often addressed using retrospective patterns that could be irrelevant. We recommend a novel landscape genomics framework for delineating future “Geminate Evolutionary Units” (GEUs) in a focal species: (1) identify loci under environmental selection, (2) model and map adaptive conservation units that may spawn future lineages, (3) forecast relative selection pressures on each future lineage, and (4) estimate their fitness and likelihood of persistence using geo-genomic simulations. Using this process, we delineated conservation units for the Yosemite toad ( Anaxyrus canorus ), a U.S. federally threatened species that is highly vulnerable to climate change. We used a genome-wide dataset, redundancy analysis, and Bayesian association methods to identify 24 candidate loci responding to climatic selection ( R 2 ranging from 0.09 to 0.52), after controlling for demographic structure. Candidate loci included genes such as MAP3K5, involved in cellular response to environmental change. We then forecasted future genomic response to climate change using the multivariate machine learning algorithm Gradient Forests. Based on all available evidence, we found three GEUs in Yosemite National Park, reflecting contrasting adaptive optima: YF-North (high winter snowpack with moderate summer rainfall), YF-East (low to moderate snowpack with high summer rainfall), and YF-Low-Elevation (low snowpack and rainfall). Simulations under the RCP 8.5 climate change scenario suggest that the species will decline by 29% over 90 years, but the highly diverse YF-East lineage will be least impacted for two reasons: (1) geographically it will be sheltered from the largest climatic selection pressures, and (2) its standing genetic diversity will promote a faster adaptive response. Our approach provides a comprehensive strategy for protecting imperiled non-model species with genomic data alone and has wide applicability to other declining species.

California

Pleistocene glacial cycles drove lineage diversification and fusion in the Yosemite toad (Anaxyrus canorus)

Species endemic to alpine environments can evolve via steep ecological selection gradients between lowland and upland environments. Additionally, many alpine environments have faced repeated glacial episodes over the past two million years, fracturing these endemics into isolated populations. In this “glacial pulse” model of alpine diversification, cycles of allopatry and ecologically divergent glacial refugia play a role in generating biodiversity, including novel admixed (“fused”) lineages. We tested for patterns of glacial pulse lineage diversification in the Yosemite toad ( Anaxyrus [ Bufo ] canorus ), an alpine endemic tied to glacially influenced meadow environments. Using double‐digest RADseq on populations densely sampled from a portion of the species range, we identified nine distinct lineages with divergence times ranging from 18 to 724 thousand years ago (ka), coinciding with multiple Sierra Nevada glacial events. Three lineages have admixed origins, and demographic models suggest these fused lineages have persisted throughout past glacial cycles. Directionality indices supported the hypothesis that some lineages recolonized Yosemite from east of the ice sheet, whereas other lineages remained in western refugia. Finally, refugial niche reconstructions suggest that low‐ and high‐elevation lineages have convergently adapted to similar climatic niches. Our results suggest glacial cycles and refugia may be important crucibles of adaptive diversity across deep evolutionary time.

California

Predicting patterns of non-native plant invasions in Yosemite National Park, California, USA

One of the major issues confronting management of parks and reserves is the invasion of non-native plant species. Yosemite National Park is one of the largest and best-known parks in the United States, harbouring significant cultural and ecological resources. Effective management of non-natives would be greatly assisted by information on their potential distribution that can be generated by predictive modelling techniques. Our goal was to identify key environmental factors that were correlated with the percent cover of non-native species and then develop a predictive model using the Genetic Algorithm for Rule-set Production technique. We performed a series of analyses using community-level data on species composition in 236 plots located throughout the park. A total of 41 non-native species were recorded which occurred in 23.7% of the plots. Plots with non-natives occurred most frequently at low- to mid-elevations, in flat areas with other herbaceous species. Based on the community-level results, we selected elevation, slope, and vegetation structure as inputs into the GARP model to predict the environmental niche of non-native species. Verification of results was performed using plot data reserved from the model, which calculated the correct prediction of non-native species occurrence as 76%. The majority of the western, lower-elevation portion of the park was predicted to have relatively low levels of non-native species occurrence, with highest concentrations predicted at the west and south entrances and in the Yosemite Valley. Distribution maps of predicted occurrences will be used by management to: efficiently target monitoring of non-native species, prioritize control efforts according to the likelihood of non-native occurrences, and inform decisions relating to the management of non-native species in postfire environments. Our approach provides a valuable tool for assisting decision makers to better manage non-native species, which can be readily adapted to target non-native species in other locations.

Diversity and Distributions

Short-term response of Holcus lanatus L. (Common Velvetgrass) to chemical and manual control at Yosemite National Park, USA

One of the highest priority invasive species at both Yosemite and Sequoia and Kings Canyon national parks is Holcus lanatus L. (common velvetgrass), a perennial bunchgrass that invades mid-elevation montane meadows. Despite velvetgrass being a high priority species, there is little information available on control techniques. The goal of this project was to evaluate the short-term response of a single application of common chemical and manual velvetgrass control techniques. The study was conducted at three montane sites in Yosemite National Park. Glyphosate spot-spray treatments were applied at 0.5, 1.0, 1.5, and 2.0% concentrations, and compared with hand pulling to evaluate effects on cover of common velvetgrass, cover of other plant species, and community species richness. Posttreatment year 1 cover of common velvetgrass was 12.1% ± 1.6 in control plots, 6.3% ± 1.5 averaged over the four chemical treatments (all chemical treatments performed similarly), and 13.6% ± 1.7 for handpulled plots. This represents an approximately 50% reduction in common velvetgrass cover in chemically- treated plots recoded posttreatment year 1 and no statistically significant reduction in hand pulled plots compared with controls. However, there was no treatment effect in posttreatment year 2, and all herbicide application rates performed similarly. In addition, there were no significant treatment effects on nontarget species or species richness. These results suggest that for this level of infestation and habitat type, (1) one year of hand pulling is not an effective control method and (2) glyphosate provides some level of control in the short-term without impact to nontarget plant species, but the effect is temporary as a single year of glyphosate treatment is ineffective over a two-year period.

California

Geologic map of Yosemite National Park and vicinity, California

This digital map database represents the general distribution of bedrock and surficial deposits of the Yosemite National Park vicinity. It was produced directly from the file used to create the print version in 1989. The Yosemite National Park region is comprised of portions of 15 7.5 minute quadrangles. The original publication of the map in 1989 included the map, described map units and provided correlations, as well as a geologic summary and references, all on the same sheet. The database delineates map units that are identified by general age and lithology following the stratigraphic nomenclature of the U.S. Geological Survey. The scale of the source maps limits the spatial resolution (scale) of the database to 1:125,000 or smaller.

IMAP

Rock-fall hazards in the Yosemite Valley

This report with map shows the areas of highest rock-fall hazard in four selected parts of Yosemite Valley, California, defined by the National Park Service. Two specific levels of hazard are recognized and identified from rock falls ranging in size from individual boulders to moderate-sized events with volumes of 100,000 m3. These different levels of hazard are associated with areas within the base of the talus (red line in Plate 1) and those normally beyond the base of the talus which we refer to as the rock-fall shadow (yellow line in Plate 1). Rock falls of even greater size, exceeding 100,000 m3, referred to as rock avalanches, are a potential hazard, but the extent of this hazard is not easily defined. Rock avalanches are considered to be much less likely to occur because of the relatively few prehistoric rock-fall avalanche deposits that have been recognized in the Yosemite Valley. With the configuration of the steep valley walls and the relatively narrow valley, it should be noted that there are no absolutely safe or zero probability areas from large rock avalanches. This study has shown in map form where rock-fall hazard exists and has given general indication of the expected frequency of these events; however, the study does not quantify the probability at any specific location, nor evaluate the risk to people or facilities to such events.

Open-File Report

Rock-fall potential in the Yosemite Valley, California

We used two methods of estimating rock-fall potential in the Yosemite Valley, California based on (1) physical evidence of previous rock-fall travel, in which the potential extends to the base of the talus, and (2) theoretical potential energy considerations, in which the potential can extend beyond the base of the talus, herein referred to as the rock-fall shadow. Rock falls in the valley commonly range in size from individual boulders of less than 1 m3 to moderate-sized falls with volumes of about 100,000 m3. Larger rock falls exceeding 100,000 m3, referred to as rock avalanches, are considered to be much less likely to occur based on the relatively few prehistoric rock-fall avalanche deposits in the Yosemite Valley. Because the valley has steep walls and is relatively narrow, there are no areas that are absolutely safe from large rock avalanches. The map shows areas of rock-fall potential, but does not predict when or how frequently a rock fall will occur. Consequently, neither the hazard in terms of probability of a rock fall at any specific location, nor the risk to people or facilities to such events can be assessed from this map.

California

Ground-water resources and water-supply alternatives in the Wawona area of Yosemite National Park, California

Planning efforts to implement the 1980 General Management Plan, which recommends relocating park administrative facilities and employee housing from Yosemite Valley in Yosemite National Park, California, have focused on the availability of water at potential relocation sites within the park. Ground-water resources and water-supply alternatives in the Wawona area, one of several potential relocation sites, were evaluated between June 1991 and October 1993. Ground water flowing from Biledo Spring near the headwaters of Rainier Creek, about 5 miles southeast of Wawona, is probably the most reliable source of good quality ground water for Wawona. A dilute calcium bicarbonate ground water flows from the spring at about 250 gallons per minute. No Giardia was detected in a water sample collected from Biledo Spring in July 1992. The concentration of dissolved 222 radon at Biledo Spring was 420 picoCuries per liter, exceeding the primary drinking-water standard of 300 picoCuries per liter proposed by the U.S. Environmental Protection Agency. This concentration, however, was considerably lower than the concentrations of dissolved 222 radon measured in ground water at Wawona. The median value for 15 wells sampled at Wawona was 4,500 picoCuries per liter. Water-quality samples from 45 wells indicate that ground water in the South Fork Merced River valley at Wawona is segregated vertically. Shallow wells produce a dilute calcium sodium bicarbonate water that results from chemical dissolution of minerals as water flows through fractured granitic rock from hillside recharge areas toward the valley floor. Tritium concentrations indicate that ground water in the shallow wells originated as precipitation after the 1960's when testing of atmospheric nuclear devices stopped. Ground water from the deep flowing wells in the valley floor is older sodium calcium chloride water. This older water probably originated either as precipitation during a climatically cooler period or as precipitation from altitudes between 1,400 and 3,700 feet higher than precipitation that recharged the local shallow ground-water-flow system. Chloride and associated cations in the deepground-water-flow system may result from upward leakage of saline ground water or from leaching of saline fluid inclusions in the granitic rocks. Water-level and pressure-gage measurements for 38 wells in the South Fork Merced River valley also indicate that the ground water in the valley is segregated vertically. Hydraulic head in deep fractures is as much as 160 feet above the valley floor. Vertical hydraulic gradients between the shallow and deep systems are as high as 4.5 feet per foot in one of two test holes drilled for this study. Measurements of in situ stress in the two test holes indicate that the vertical segregation of ground water may be related to pressures in the earth that squeeze horizontal fractures closed at depth. Fractures within a few hundred feet of land surface are poorly connected to fractures deeper beneath the valley. About 100 privately owned wells currently are in use at Wawona; but, the ground-water-flow system may not be an adequate source of good quality ground water for relocated park facilities. Yields from existing wells are low (median 4-5 gallons per minute) and traditional methods for locating high-yielding wells in fractured rocks have not been successful in this area. Concentrations of dissolved 222 radon (median 4,500 picoCuries per liter) are high, and the development of deep ground water could cause deeper saline water to migrate upward into producing wells. The South Fork Merced River, the primary source of water supply for Wawona, does not meet current demands during late summer and autumn. Data collected between 1958 and 1968 indicate that 25 percent of the time discharge of the South Fork River at Wawona during the dry season (August through October) was less than 2 cubic feet per second-the discharge rate at which the National Park Service is restricted from withdrawing water from the river. the river, however, could be relied on for additional water supply if facilities were constructed to divert and store water during periods of high flow for use later in the year.

California

Biological, habitat, and water quality conditions in the upper Merced River drainage, Yosemite National Park, California, 1993-1996

Four studies were done in the upper Merced River drainage in Yosemite National Park and nearby areas from 1993 to 1996. First, monitoring studies of benthic algae, benthic invertebrates, fish, and habitat were undertaken at sites near Happy Isles and Pohono bridges from 1993 to 1995 as part of the National Water-Quality Assessment Program of the U.S. Geological Survey. Second, an ecological survey of benthic algae, benthic invertebrates, fish, and habitat was done in the upper Merced River drainage in 1994. Third, a special study of benthic algae, habitat, and water quality was done in the reach of the Merced River within Yosemite Valley to deter-mine whether human activities were having measurable effects on the ecosystem. Fourth, baseline data on benthic algae, benthic invertebrates, and habitat were collected in 1996 at four sites, two of which were undergoing extensive streambank restoration activities. Comparisons of the baseline data with future collections could be used to assess the effects of streambank restoration on aquatic biota. The general conclusion from these studies is that water quality in the upper Merced River was very good from 1993-1996, despite high levels of human activities in some areas. Fish communities did not appear to be a useful indicator of habitat and water quality because of low species richness and the apparent importance of physical barriers in determining species distributions. Measurements of fish densities and size-distributions might be useful, but would be logistically difficult. Benthic algae and benthic invertebrates do appear to be useful in monitoring environmental conditions. Benthic algae may be more sensitive than benthic invertebrates to small environmental differences within years. Benthic algae were also more responsive than benthic invertebrates to differences in discharge between years. Thus, benthic invertebrates may be more useful in comparing environmental conditions between years, independent of discharge conditions.

Water-Resources Investigations Report