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At least 217 records · Page 12Linked to original sources

Rare long-distance dispersal of the Island Night Lizard, Xantusia riversiana, maintains high diversity in a fragmented environment

The Island Night Lizard ( Xantusia riversiana ) is endemic to three of the Channel Islands off the coast of California, USA. Introduced species such as goats, sheep, and cats have profoundly affected the fauna and flora of the islands for over 150 years, but most of these non-native species have been recently removed. We measured the distribution of genetic diversity in Island Night Lizards across San Nicolas Island using DNA microsatellites to assess the impacts of historical habitat change on effective population size, gene flow, and population divergence; to provide baseline data for future monitoring of genetic diversity; and to provide recommendations to inform the restoration of degraded habitat. Despite a history of profound anthropogenic habitat disturbance, genetic diversity was high within sites, and there was no evidence of population bottlenecks. Divergence between sites was extraordinarily high, as expected for this sedentary species. Landscape resistance modeling using circuit theory showed that unsuitable habitat is relatively permeable to gene flow compared to suitable habitat, and yet populations separated by very short geographic distances remain genetically distinct. We found no evidence of a need for short-term intervention such as artificial translocations to maintain genetic diversity. Instead, we suggest that management should focus on maintaining, improving, and increasing habitat, especially in creating patches of habitat to link existing sites.

California

Relationships between diatom metrics based on species nutrient traits and agricultural land use

We assessed how diatom metrics were related to different ranges of agricultural land use. Diatom assemblage composition, nutrients, and landscape characteristics were determined at 232 sites in eight agriculturally dominated study areas of the continental United States. Two regional groups based on differences in diatom relations to human disturbance were determined. Changes in diatom species composition were related to nutrients,pH,and conductivity in the eastern study areas (due to more wetlands) and more exclusively to nutrients in the west-central study areas. Homogenization of diatom flora among streams was related to high agricultural disturbance at this transcontinental scale. Species traits were developed separately for the east and west central study groups and calculated two ways: indicator species analysis for taxa in low and high TN or TP conditions and weighted average partial least squares models of TN and TP concentration. These diatom metrics were significantly related to many indicators of agricultural land use in watersheds, especially percent row crops. Further analysis was conducted on only the west-central region due to its larger sample size.Overall, diatom metrics using species responses to N gradients were better related to agricultural land use than were species responses to P gradients. Most nutrient-based diatom metrics changed greatly in response to low ranges of percent row crops, but only a few high N diatom metrics responded to high row crop conditions. The greater response of diatoms to changes in low agriculture conditions may be due to past diatom evolution occurring when most waters had low nutrient conditions.

Alabama, Arkansas, Delaware, Florida, Georgia, Ida

Monitoring the dynamics of an invasive emergent macrophyte community using operational remote sensing data

Potamogeton crispus L. (curly pondweed) is a cosmopolitan aquatic macrophyte considered invasive in North America and elsewhere. Its range is expanding and, on individual water bodies, its coverage can be dynamic both within and among years. In this study, we evaluate the use of free and low-cost satellite remote sensing data to monitor a problematic emergent macrophyte community dominated by P. crispus . Between 2000 and 2006, we acquired eight satellite images of 24,000-ha Lake Sharpe, South Dakota (USA). During one of the dates for which satellite imagery was acquired, we sampled the lake for P. crispus and other emergent macrophytes using GPS and photography for documentation. We used cluster analysis to assist in classification of the satellite imagery and independently validated results using the field data. Resulting estimates of emergent macrophyte coverage ranged from less than 20 ha in 2002 to 245 ha in 2004. Accuracy assessment indicated 82% of image pixels were correctly classified, with errors being primarily due to failure to identify emergent macrophytes. These results emphasize the dynamic nature of P. crispus -dominated macrophyte communities and show how they can be effectively monitored over large areas using low-cost remote sensing imagery. While results may vary in other systems depending on water quality and local flora, such an approach could be applied elsewhere and for a variety of macrophyte communities.

Hydrobiologia

Eruptive and environmental processes recorded by diatoms in volcanically-dispersed lake sediments from the Taupo Volcanic Zone, New Zealand

Late Pleistocene diatomaceous sediment was widely dispersed along with volcanic ash (tephra) across and beyond New Zealand by the 25.4 ka Oruanui supereruption from Taupo volcano. We present a detailed analysis of the diatom populations in the Oruanui tephra and the newly discovered floras in two other eruptions from the same volcano: the 28.6 ka Okaia and 1.8 ka Taupo eruptions. For comparison, the diatoms were also examined in Late Pleistocene and Holocene lake sediments from the Taupo Volcanic Zone (TVZ). Our study demonstrates how these microfossils provide insights into the lake history of the TVZ since the Last Glacial Maximum. Morphometric analysis of Aulacoseira valve dimensions provides a useful quantitative tool to distinguish environmental and eruptive processes within and between individual tephras. The Oruanui and Okaia diatom species and valve dimensions are highly consistent with a shared volcanic source, paleolake and eruption style (involving large-scale magma-water interaction). They are distinct from lacustrine sediments sourced elsewhere in the TVZ. Correspondence analysis shows that small, intact samples of erupted lake sediment (i.e., lithic clasts in ignimbrite) contain heterogeneous diatom populations, reflecting local variability in species composition of the paleolake and its shallowly-buried sediments. Our analysis also shows a dramatic post-Oruanui supereruption decline in Cyclostephanos novaezelandiae, which likely reflects a combination of (1) reorganisation of the watershed in the aftermath of the eruption, and (2) overall climate warming following the Last Glacial Maximum. This decline is reflected in substantially lower proportions of C. novaezelandiae in the 1.8 ka Taupo eruption deposits, and even fewer in post-1.8 ka sediments from modern (Holocene) Lake Taupo. Our analysis highlights how the excellent preservation of siliceous microfossils in volcanic tephra may fingerprint the volcanic source region and retain a valuable record of volcanically-influenced environmental change.

Taupo Volcanic Zone

Life-history habitat matching in invading non-native plant species

We briefly reviewed the literature on habitat matching in invading non-native plant species. Then we hypothesized that the richness and cover of native annual and perennial plant species integrate complex local information of vegetation and soils that would help to predict invasion success by similarly adapted non-native plant species. We tested these ‘life-history habitat matching’ relationships in 603 0.1-ha plots, including 294 plots in Colorado, which were relatively high for the cover of native perennial plant species, and for 309 0.1-ha plots in southern Utah, which were relatively high in the cover of native annual plant species. We found strong positive relationships between the richness and foliar cover for both native and non-native species, whether they were annual or perennial species (0.34 > r 2 < 0.53; P < 0.0001). We also found significant positive relationships between the cover of native annual species at a site and the richness ( r 2 = 0.13; P < 0.0001) and the foliar cover ( r 2 = 0.06; P < 0.0001) of non-native annual species. The proportion of non-native annual species in the flora of a plot also increased significantly with the foliar cover of native annual species. Conversely, the richness and cover of non-native annual species were significantly negatively associated with the foliar cover of native perennial species ( r 2 = 0.05 and 0.06, respectively; P < 0.0001). The cover of non-native annual or perennial species was not significantly correlated with soil texture variables, %N, or %C. We conclude that there may be a high degree of life-history habitat matching by non-native annual species in these study sites. Information on native annual and perennial species richness and cover may help characterize the complex soils, climate, and disturbance environment in which similarly adapted non-native plant species establish and gain foliar cover.

Plant and Soil

Impacts of coastal land use and shoreline armoring on estuarine ecosystems: An introduction to a special issue

The nearshore land-water interface is an important ecological zone that faces anthropogenic pressure from development in coastal regions throughout the world. Coastal waters and estuaries like Chesapeake Bay receive and process land discharges loaded with anthropogenic nutrients and other pollutants that cause eutrophication, hypoxia, and other damage to shallow-water ecosystems. In addition, shorelines are increasingly armored with bulkhead (seawall), riprap, and other structures to protect human infrastructure against the threats of sea-level rise, storm surge, and erosion. Armoring can further influence estuarine and nearshore marine ecosystem functions by degrading water quality, spreading invasive species, and destroying ecologically valuable habitat. These detrimental effects on ecosystem function have ramifications for ecologically and economically important flora and fauna. This special issue of Estuaries and Coasts explores the interacting effects of coastal land use and shoreline armoring on estuarine and coastal marine ecosystems. The majority of papers focus on the Chesapeake Bay region, USA, where 50 major tributaries and an extensive watershed (~ 167,000 km 2 ), provide an ideal model to examine the impacts of human activities at scales ranging from the local shoreline to the entire watershed. The papers consider the influence of watershed land use and natural versus armored shorelines on ecosystem properties and processes as well as on key natural resources.

Estuaries and Coasts

Identifying and constraining marsh-type transitions in response to increasing erosion over the past century

Marsh environments, characterized by their flora and fauna, change laterally in response to shoreline erosion, water levels and inundation, and anthropogenic activities. The Grand Bay coastal system (USA) has undergone multiple large-scale geomorphic and hydrologic changes resulting in altered sediment supply, depositional patterns, and degraded barrier islands, leaving wetland salt marshes vulnerable to increased wave activity. Two shore-perpendicular transect sites, one along a low-activity shoreline and the other in a high activity area of the same bay-marsh complex, were sampled to investigate how the marshes within 50 m of the modern shoreline have responded to different levels of increased wave activity over the past century. Surface sediments graded finer and more organic with increased distance from the shoreline while cores generally exhibited a coarsening upwards grain-size trend; all cores contained multiple large sedimentological shifts. 210 Pb-based mass accumulation rates over the last two decades were greater than the long-term (centurial) average at each site with the fastest accumulation rates of 7.81 ± 1.58 and 7.79 ± 1.63 kg/m 2 /year at the sites nearest the shoreline. A shoreline change analysis of three time-slices (1848–2017, 1957–2017, 2016–2017) shows increased erosion at both sites since 1848 with modern rates of −0.95 and −0.88 m/year. Downcore sedimentology, mass accumulation rates, and shoreline change rates paired with foraminiferal biofacies and identification of local estuarine indicator species, Paratrochammina simplissima , aided in identifying paleo marsh types, their relative proximity to the shoreline, and sediment provenance. The high-energy marsh site transitioned from middle marsh to low marsh in the 1960s, and the low-energy marsh site transitioned later, at the end of the twentieth and early twenty-first century, due to its more protected location. Marsh type transition corresponds chronologically with the coarsening upwards grain-size trend observed and the degradation of Grand Batture Island; since its submergence, signatures of multiple storm event have been preserved downcore.

Mississippi

Development of ten microsatellite loci in the invasive giant African land snail, Achatina (=Lissachatina) fulica Bowdich, 1822

A suite of tetra-nucleotide microsatellite loci were developed for the invasive giant African land snail, Achatina (=Lissachatina) fulica Bowdich, 1822, from Ion Torrent next-generation sequencing data. Ten of the 96 primer sets tested amplified consistently in 30 snails from Miami, Florida, plus 12 individuals representative of their native East Africa, Indian and Pacific Ocean regions. The loci displayed moderate levels of allelic diversity (average 5.6 alleles/locus) and heterozygosity (average 42 %). Levels of genetic diversity were sufficient to produce unique multi-locus genotypes and detect phylogeographic structuring among regional samples. The invasive A. fulica can cause extensive damage to important food crops and natural resources, including native flora and fauna. The loci characterized here will be useful for determining the origins and tracking the spread of invasions, detecting fine-scale spatial structuring and estimating demographic parameters.

Conservation Genetics Resources

Impacts of mute swans ( Cygnus olor ) on submerged aquatic vegetation in Illinois River Valley backwaters

Wetland loss in North America has been considerable and well documented, and the establishment of exotic species in remaining wetlands can further reduce their ability to support native flora and fauna. In the Chesapeake Bay and Great Lakes ecosystems, exotic mute swans ( Cygnus olor ) have been found to negatively impact wetlands through degradation of submerged aquatic vegetation (SAV) communities. Mute swan populations have expanded into many areas of mid-continental North America outside the Great Lakes ecosystem, but the environmental impact of these populations is not well known. Mid-continental wetlands in North America differ in physical characteristics (e.g., size, depth, and permanency) and aquatic vegetation species composition compared to wetlands in other areas where mute swans have been studied and, thus, may be more or less susceptible to degradation from swan herbivory. To investigate the impact of mute swan herbivory on SAV communities in mid-continent wetlands, we used exclosures to prevent swans from foraging in 2 wetland complexes in central Illinois. Above-ground biomass of vegetation did not differ between exclosures and controls; however, mean below-ground biomass was greater in exclosures (52.0 g/m 2 , SE&thinsp;=&thinsp;6.0) than in controls (34.4 g/m 2 SE&thinsp;=&thinsp;4.0). Thus, although swan densities were lower in our study region compared to that of previous studies, we observed potentially detrimental impacts of swan herbivory on below-ground biomass of SAV. Our results indicate that both above-ground and below-ground impacts of herbivory should be monitored, and below-ground biomass may be most sensitive to swan foraging.

Illinois

A floodplain continuum for Atlantic coast rivers of the Southeastern US: Predictable changes in floodplain biota along a river's length

Floodplains are among the world’s economically-most-valuable, environmentally-most-threatened, and yet conceptually-least-understood ecosystems. Drawing on concepts from existing riverine and wetland models, and empirical data from floodplains of Atlantic Coast rivers in the Southeastern US (and elsewhere when possible), we introduce a conceptual model to explain a continuum of longitudinal variation in floodplain ecosystem functions with a particular focus on biotic change. Our hypothesis maintains that major controls on floodplain ecology are either external (ecotonal interactions with uplands or stream/river channels) or internal (wetland-specific functions), and the relative importance of these controls changes progressively from headwater to mid-river to lower-river floodplains. Inputs of water, sediments, nutrients, flora, and fauna from uplands-to-floodplains decrease, while the impacts of wetland biogeochemistry and obligate wetland plants and animals within-floodplains increase, along the length of a river floodplain. Inputs of water, sediment, nutrients, and fauna from river/stream channels to floodplains are greatest mid-river, and lower either up- or down-stream. While the floodplain continuum we develop is regional in scope, we review how aspects may apply more broadly. Management of coupled floodplain-river ecosystems would be improved by accounting for how factors controlling the floodplain ecosystem progressively change along longitudinal riverine gradients.

Wetlands

A minimum U-Pb age for Siberian flood-basalt volcanism

Establishing an accurate and precise age for Siberian flood-basalt volcanism is of great importance in evaluating causes for the unequaled mass extinction of flora and fauna at the Permian-Triassic boundary. We report a new, minimum U-Pb age obtained from zircon and baddeleyite from the mineralized Noril'sk I intrusion that cuts the lower third of this rapidly deposited, 3500-m-thick volcanic sequence near Noril'sk. This 251.2 ?? 0.3 (2??) Ma age is within analytical error of the SHRIMP U-Pb age for zircon from the Permian-Triassic boundary at Meishan, South China [251.1 ?? 3.6 Ma (2??)], and confirms Siberian basaltic volcanism as a possible contributor to the mass extinction.

Geochimica et Cosmochimica Acta

The terminal Eocene event and the Polish connection

The Eocene/Oligocene boundary in Europe is marked by major discontinuities in all environments: the “Grande Coupure” in continental mammals; the elimination of semitropical elements from high-latitude floras; the virtually complete replacement of the shallow-marine malacofauna; and an extraordinary downslope excursion of carbonate deposition in deep-ocean basins (drop in the CCD). These phenomena collectively represent the “Terminal Eocene Event” (TEE). In the Carpathian Mountains, the TEE is manifested in the thin but regionally persistent Globigerina Marl, a calcareous unit containing abundant cool-water microplankton that occurs within very thick, siliceous, bathyal flysch sequences. In southern Poland, the marl is of very latest Eocene age, within planktonic foraminifera zone P17, calcareous nannoplankton zone NP19/20, and the zone of the dinoflagellate Rhomdodinium perforatum . Zircons from bentonites bracketing the marl are dated by fission-track analysis; at Polany, two underlying bentonites are 41.7 and 39.8 Ma, and at Znamirowice two overlying bentonites are 34.6 and 28.9 Ma, in sequence. This accords with glauconite K/Ar ages in Western Europe by which the Eo/Oligocene boundary age is estimated at 37–38 Ma. Global correlations indicate that the TEE corresponds to a major glacio-eustatic regression with a duration of about 0.5 Ma, in which a large Antarctic ice cap was formed, the ocean circulation was permanently changed to the psychrospheric condition, and world climate shifted irreversibly towards the modern state.

Palaeogeography, Palaeoclimatology, Palaeoecology

Radiocarbon dates and late-Quaternary stratigraphy from Mamontova Gora, unglaciated central Yakutia, Siberia, U.S.S.R.

A fine exposure of perennially frozen ice-rich silt and associated flora and vertebrate fauna of late-Quaternary age exists at Mamontova Gora along the Aldan River in central Yakutia, Siberia, U.S.S.R. The silt deposit caps a 50-m-high terrace and consists of three units. An upper layer 1–2 m thick overlies a 10–15-m-thick brownish to black silt layer. The lower silt layer is greenish to gray and about 15 m thick. All the silt is well sorted with 60% of the particles falling between 0.005 and 0.5 mm in diameter and is generally chemically and mineralogically homogeneous. The middle unit contains may extinct vertebrate mammal remains and ice wedges. The lower unit contains little vegetation and no ice wedges. The silt is widespread and exists as a loamy blanket on terraces at various elevations on both sides of the lower Aldan River. The origin of the silt blanket of late-Quaternary age in central Yakutia has long been controversial. Various hypotheses have been suggested, including lacustrine and alluvial, as well as frost-action origins. It is sometimes referred to as loess-like loam. Péwé believes the silt at Mamontova Gora is loess, some of which has been retransported very short distances by water. The silt probably was blown from wide, braided, unvegetated flood plains of rivers draining nearby glaciers. The silt deposits are late Quaternary in age and probably associated with the Maximum glaciation (Samarov) and Sartan and Syryan glaciations of Wisconsinan age. On the basis of biostratigraphy, 10 radiocarbon dates, and their relation to the nearby glacial record, it is felt that the upper unit at Mamontova Gora is Holocene and the middle unit is Wisconsinan. The youngest date available from the middle unit at this particular location is 26,000 years. Dates greater than 56,000 years were obtained in the lower part of the middle unit. The lower unit is definitely beyond the range of radiocarbon dating and probably is older than the last interglacial. The sediment, fauna, ice wedges, stratigraphy, and age of perennially frozen slit deposits in central Alaska are remarkably similar to those of the deposits exposed in central Yakutia. Both areas consist of unglaciated rolling lowlands and river terraces surrounded by high mountains that were extensively glaciated in Pleistocene time. The glaciers extended from the high mountains to the edges of the ranges. In both regions, extensively braided, silt-charged rivers drained the mountains and flowed through the lowlands on their way to the sea. It follows that there should be a similar late-Quaternary history.

Siberia

Shock-induced microdeformations in quartz and other mineralogical indications of an impact event at the Cretaceous-Tertiary boundary

The event terminating the Cretaceous period and the Mesozoic era caused massive extinctions of flora and fauna worldwide. Theories of the nature of this event can be classed as endogenic (volcanic, climatic, etc.) or exogenic (extraterrestrial causes). Mineralogical evidence from the boundary clays and claystones strongly favor the impact of an extraterrestrial body as the cause of this event. Nonmarine KT boundary claystones are comprised of two separate layers—an upper layer composed of high-angle ejecta material (shocked quartz, altered glass and spinel) and a basal kaolinitic layer containing spherules, clasts, and altered glass, together with some shocked grains. Recognition of this dual-layered nature of the boundary clay is important for the determination of the timing and processes involved in the impact event and in the assignment and interpretation of geochemical signatures. Multiple sets of shock-induced microdeformations (planar features) in quartz grains separated from K T boundary clays provide compelling evidence of an impact event. This mineralogical manifestation of shock metamorphism is associated worldwide with a large positive anomaly of iridium in these boundary clays, which has also been considered indicative of the impact of a large extraterrestrial body. Global distributions of maximum sizes of shocked quartz grains from the boundary clays and the mineralogy of the ejecta components favor an impact on or near the North American continent. Spinel crystals (magnesioferrite) occur in the boundary clays as micrometer-sized octahedra or skeletal forms. Their composition differs from that of spinels found in terrestrial oceanic basalts. Magnesioferrite crystals are restricted to the high-angle ejecta layer of the boundary clays and their small size and skeletal morphology suggest that they are condensation products of a vaporized bolide. Hollow spherules ranging up to 1 mm in size are ubiquitously associated with the boundary clays. In nonmarine sections, where a high-angle ejecta layer and an underlying kaolinitic layer can be distinguished, the spherules are found only in the kaolinitic layer. The morphologies and surface features of these spherules suggest that they are original forms, and not secondary growths or algal bodies. These impact spherules closely resemble microtektites in size and shape. All of these features of the boundary clay are uniquely associated with impact, and cannot have been formed by volcanic or other terrestrial processes.

Colorado, Montana, New Mexico, Wyoming

Comparison of the petrography, palynology and paleobotany of the Stockton coal bed, West Virginia and implications for paleoenvironmental interpretations

The Stockton coal bed (Middle Pennsylvanian) is a relatively high ash coal composed primarily of moderately thin banded, sparsely thin banded, and nonbanded coal (splint and cannel coal). Comparisons of petrographic, palynologic, and paleobotanic data gathered from the same sample sets from a single column of the Stockton coal bed indicate that compositional correspondences among the sets exist regardless of coal type. Some correspondences are believed to exist because of original plant constituents and others because of the paleoenvironment of peat formation. Using some combination of these data is critical when interpreting paleoenvironmental conditions because (1) a direct correspondence is lacking between many of the data and (2) each of the three data sets provides a unique and important perspective on the paleomire. The Stockton paleomire in the area of this study supported a diverse flora that consisted of both small and arboreous lycopsids, small ferns and tree ferns, calamites, cordaites, and pteridosperms. There appear to have been two successions of Lycospora spore-dominated, vitrinite-rich, liptinite-poor peat formation, which were followed by inertinite-rich peat formation marked by a tree fern-dominant spore assemblage and abundant unidentifiable plant tissues. These are interpreted to be two water-laden or topogenous peat formational stages followed by slightly domed, better drained peat formation.

West Virginia

Comparison of the petrography, palynology, and paleobotany of the Little Fire Creek coal bed, southwestern Virginia, U.S.A.

Two continuous cores that penetrated the Lower Pennsylvanian Little Fire Creek coal bed in the Southwestern coal field in southwestern Virginia were sampled and X-ray radiographed to determine subunit distinctions. Comparison of petrographic, palynologic, and paleobotanic data from the same sample sets from the two cores allowed for comparison of compositional data within the Little Fire Creek coal bed. The proximate, petrographic, palynologic, and plant tissue data from two sets of samples indicate a high ash, gelocollinite- and liptinite-rich coal consisting of a relatively diverse paleoflora, including lycopsid trees, small lycopsids, tree ferns, small ferns, pteridosperms (seed ferns), and rare calamites and cordaites. The relatively very high ash yields (3–80 wt%), the relatively thin subunits (1–28 cm), and the large scale vertical variations in palynomorph floras suggest that the study area was at the edge of the paleopea-forming environment. As a result, most of the compositional correspondences are among those components indicative of degradation or decomposition.

Virginia

Changing patterns of Pennsylvanian coal-swamp vegetation and implications of climatic control on coal occurrence

Improved regional and interregional stratigraphic correlations of Pennsylvanian strata permit comparisons of vegetational changes in Euramerican coal swamps. The coal-swamp vegetation is known directly from in situ coal-ball peat deposits from more than 65 coals in the United States and Europe. Interpretations of coal-swamp floras on the basis of coal-ball peat studies are extended to broader regional and stratigraphic patterns by use of coal palynology. Objectives of the quantitative analyses of the vegetation in relation to coal are to determine the botanical constituents at the peat stage and their environmental implications for plant growth and peat accumulation. Morphological and paleoecological analyses provide a basis for deducing freshwater regimes of coal swamps. Changes in composition of Pennsylvanian coal-swamp vegetation are quire similar from one paralic coal region to another and show synchrony that is attributable to climate. Paleobotany and paleogeography of the Euramerican province indicate a moist tropical paleoclimate. Rainfall, runoff and evapotranspiration were the variable climatic controls in the distribution of coal-swamp vegetation, peat accumulation and coal resources. In relative terms of climatic wetness the Pennsylvanian Period is divisible into five intervals, which include two relatively drier intervals that developed during the Lower-Middle and Middle-Upper Pennsylvanian transitions. The climate during Early Pennsylvanian time was moderately wet and the median in moisture availability. Early Middle Pennsylvanian was drier, probably seasonally dry-wet; late Middle Pennsylvanian was the wettest in the Midcontinent; early Late Pennsylvanian was the driest; and late Late Pennsylvanian was probably the wettest in the Dunkard Basin. The five climatic intervals represent a general means of dividing coal resources within each region into groups with similar botanical constituents and environments of peat accumulation. Regional differences in basinal geology and climate were significant variables, but the synchronous control of paleoclimate was of primary importance.

International Journal of Coal Geology

Stratigraphic and interregional changes in Pennsylvanian coal-swamp vegetation: Environmental inferences

Quantitative analysis of Pennsylvanian coal-swamp vegetation provides a means of inferring organization and structure of communities. Distribution of these communities further provides inferences about environmental factors, including paleoclimate. Our observations are based on in situ, structurally preserved peat deposits in coal-ball concretions from 32 coal seams in the eastern one-half of the United States and from several seams in western Europe and on spore assemblages from more than 150 seams. There were three times of particularly significant and nearly synchronous vegetational changes in the Midcontinent and Appalachian coal regions during the Pennsylvanian Period. Each was different in kind and magnitude. The first marked changes occurred during the early part of the Middle Pennsylvanian with the fluctuating decline in the high level of lycopod dominance. The abundance of cordaites increased. There was a rise in the occurrences of the lycopod herbs to form intercalated marshlands and an overall increase in floral diversity. Changes ensuing from this time also include shifts in dominant species of lycopod trees and a sustained rise in abundance and diversity of tree-fern spores. The next significant time of change was during the middle part of the Middle Pennsylvanian, representing both a culmination of earlier trends and expansions of cordaites in the Midcontinent where there was a maximum change in species without net loss of diversity. Tree ferns and medullosan pteridosperms attained subdominant levels of abundance and diverse lycopod species dominated except in the Atokan-Desmoinesian transition of the Midcontinent. The third and sharpest break occurred near the Middle—Late Pennsylvanian boundary when extinctionsof the dominant, coal-swamp lycopods allowed development of tree-fern dominance. The Late Pennsylvanian coal swamps apparently were colonized or recolonized mainly by species from outside coal swamps rather than by the survivor populations of the Middle Pennsylvanian swamps. Paralleling the changes in floras through the Pennsylvanian are changes in preservational aspects of the peat. These include a decline in shoot/root ratios from approximately 1 to < 1 during the first time of vegetational changes and a rise in this ratio during the second; there was a parallel rise and fall in fusain abundance and a rise in wood/periderm ratios. The stratigraphic distribution of identified coal resources in the United States is interpreted as largely dependent on net changes in relative wetness of Pennsylvanian coal swamps, a pattern of drying during the first period of vegetational change, followed by a concomitant increase in continuous wet climate with brackish influence in the Midcontinent during the second; this was followed by a time of extreme moisture stress bringing on the third, and most severe, vegetational change.

International Journal of Coal Geology