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At least 199 records · Page 11Linked to original sources

Fix success and accuracy of GPS radio collars in old-growth temperate coniferous forests

Global Positioning System (GPS) telemetry is used extensively to study animal distribution and resource selection patterns but is susceptible to biases resulting from data omission and spatial inaccuracies. These data errors may cause misinterpretation of wildlife habitat selection or spatial use patterns. We used both stationary test collars and collared free-ranging American black bears ( Ursus americanus ) to quantify systemic data loss and location error of GPS telemetry in mountainous, old-growth temperate forests of Olympic National Park, Washington, USA. We developed predictive models of environmental factors that influence the probability of obtaining GPS locations and evaluated the ability of weighting factors derived from these models to mitigate data omission biases from collared bears. We also examined the effects of microhabitat on collar fix success rate and examined collar accuracy as related to elevation changes between successive fixes. The probability of collars successfully obtaining location fixes was positively associated with elevation and unobstructed satellite view and was negatively affected by the interaction of overstory canopy and satellite view. Test collars were 33% more successful at acquiring fixes than those on bears. Fix success rates of collared bears varied seasonally and diurnally. Application of weighting factors to individual collared bear fixes recouped only 6% of lost data and failed to reduce seasonal or diurnal variation in fix success, suggesting that variables not included in our model contributed to data loss. Test collars placed to mimic bear bedding sites received 16% fewer fixes than randomly placed collars, indicating that microhabitat selection may contribute to data loss for wildlife equipped with GPS collars. Horizontal collar errors of >800 m occurred when elevation changes between successive fixes were >400 m. We conclude that significant limitations remain in accounting for data loss and error inherent in using GPS telemetry in coniferous forest ecosystems and that, at present, resource selection patterns of large mammals derived from GPS telemetry should be interpreted cautiously.

Washington↗

Identification of novel hepaciviruses and Sylvilagus-associated viruses via metatranscriptomics in North American lagomorphs

Cottontails ( Sylvilagus spp.) and jackrabbits ( Lepus spp.) within the Leporidae family are native to North America and are found in a wide range of habitats, including deserts, forests, and grasslands. Although there is a growing body of research describing the arrival of the highly virulent rabbit haemorrhagic disease virus 2 (RHDV2, GI.2) on this continent, and its impact on native lagomorphs, information about the natural virome and microbiome of healthy and deceased American lagomorphs is relatively limited. In this study, we used a meta-transcriptomics approach to conduct whole pathogen profiling on healthy and deceased animals in the USA. We analysed 48 matched liver and lung sample pools from apparently healthy cottontails and jackrabbits in Texas and an additional 48 liver samples from deceased animals from nine other US states. This approach enabled the discovery of three distinct new viruses and revealed additional new insights into the lung and liver microbiomes of North American lagomorphs. Of the three new viruses, a tetnovirus and a novel picorna-like virus were likely of insect origin and therefore considered environmental contaminants. Of particular interest was a new species of hepacivirus, with around 50% sequence identity to a known hepacivirus from a xeric four-striped grass rat ( Rhabdomys pumilio ). Phylogenetic analysis from 41 individual hepacivirus genomes recovered from our lagomorph samples revealed two distinct clades, corresponding with different cottontail species. No hepaciviruses were detected in any of the jackrabbit samples. This is the first description of a hepacivirus in lagomorphs. Our findings extend the Hepacivirus genus, provide new insights into its evolution, and describe the first baseline on microbial diversity in North American lagomorphs, an important step towards understanding the role of potential pathogens for population management and conservation.

Arizona, California, Iowa, Massachusetts, Montana,↗

Characterization of errors in a coupled snow hydrology-microwave emission model

Traditional approaches to the direct estimation of snow properties from passive microwave remote sensing have been plagued by limitations such as the tendency of estimates to saturate for moderately deep snowpacks and the effects of mixed land cover within remotely sensed pixels. An alternative approach is to assimilate satellite microwave emission observations directly, which requires embedding an accurate microwave emissions model into a hydrologic prediction scheme, as well as quantitative information of model and observation errors. In this study a coupled snow hydrology [Variable Infiltration Capacity (VIC)] and microwave emission [Dense Media Radiative Transfer (DMRT)] model are evaluated using multiscale brightness temperature (TB) measurements from the Cold Land Processes Experiment (CLPX). The ability of VIC to reproduce snowpack properties is shown with the use of snow pit measurements, while TB model predictions are evaluated through comparison with Ground-Based Microwave Radiometer (GBMR), air-craft [Polarimetric Scanning Radiometer (PSR)], and satellite [Advanced Microwave Scanning Radiometer for the Earth Observing System (AMSR-E)] TB measurements. Limitations of the model at the point scale were not as evident when comparing areal estimates. The coupled model was able to reproduce the TB spatial patterns observed by PSR in two of three sites. However, this was mostly due to the presence of relatively dense forest cover. An interesting result occurs when examining the spatial scaling behavior of the higher-resolution errors; the satellite-scale error is well approximated by the mode of the (spatial) histogram of errors at the smaller scale. In addition, TB prediction errors were almost invariant when aggregated to the satellite scale, while forest-cover fractions greater than 30% had a significant effect on TB predictions. ?? 2008 American Meteorological Society.

Journal of Hydrometeorology↗

Geomagnetic monitoring in the mid-Atlantic United States

Near historic battlegrounds of the American Civil War, southeast of Fredericksburg, Virginia, on a secluded grassy glade surrounded by forest, a specially designed observatory records the Earth’s changing magnetic field. This facility, the Fredericksburg Magnetic Observatory, is 1 of 14 observatories the U.S. Geological Survey Geomagnetism Program operates at various locations across the United States and its Territories as a service to the Nation and in support of a diversity of governmental, academic, and commercial scientific projects.

Virginia↗

Declines in low-elevation subalpine tree populations outpace growth in high-elevation populations with warming

Species distribution shifts in response to climate change require that recruitment increase beyond current range boundaries. For trees with long life spans, the importance of climate-sensitive seedling establishment to the pace of range shifts has not been demonstrated quantitatively. Using spatially explicit, stochastic population models combined with data from long-term forest surveys, we explored whether the climate-sensitivity of recruitment observed in climate manipulation experiments was sufficient to alter populations and elevation ranges of two widely distributed, high-elevation North American conifers. Empirically observed, warming-driven declines in recruitment led to rapid modelled population declines at the low-elevation, ‘warm edge’ of subalpine forest and slow emergence of populations beyond the high-elevation, ‘cool edge’. Because population declines in the forest occurred much faster than population emergence in the alpine, we observed range contraction for both species. For Engelmann spruce, this contraction was permanent over the modelled time horizon, even in the presence of increased moisture. For limber pine, lower sensitivity to warming may facilitate persistence at low elevations – especially in the presence of increased moisture – and rapid establishment above tree line, and, ultimately, expansion into the alpine. Synthesis . Assuming 21st century warming and no additional moisture, population dynamics in high-elevation forests led to transient range contractions for limber pine and potentially permanent range contractions for Engelmann spruce. Thus, limitations to seedling recruitment with warming can constrain the pace of subalpine tree range shifts.

Journal of Ecology↗

Dams, floodplain land use, and riparian forest conservation in the semiarid Upper Colorado River Basin, USA

Land and water resource development can independently eliminate riparian plant communities, including Fremont cottonwood forest (CF), a major contributor to ecosystem structure and functioning in semiarid portions of the American Southwest. We tested whether floodplain development was linked to river regulation in the Upper Colorado River Basin (UCRB) by relating the extent of five developed land-cover categories as well as CF and other natural vegetation to catchment reservoir capacity, changes in total annual and annual peak discharge, and overall level of mainstem hydrologic alteration (small, moderate, or large) in 26 fourth-order subbasins. We also asked whether CF appeared to be in jeopardy at a regional level. We classified 51% of the 57,000 ha of alluvial floodplain examined along >2600 km of mainstem rivers as CF and 36% as developed. The proportion developed was unrelated to the level of mainstem hydrologic alteration. The proportion classified as CF was also independent of the level of hydrologic alteration, a result we attribute to confounding effects from development, the presence of time lags, and contrasting effects from flow alteration in different subbasins. Most CF (68% by area) had a sparse canopy (???5% cover), and stands with >50% canopy cover occupied <1% of the floodplain in 15 subbasins. We suggest that CF extent in the UCRB will decline markedly in the future, when the old trees on floodplains now disconnected from the river die and large areas change from CF to non-CF categories. Attention at a basinwide scale to the multiple factors affecting cottonwood patch dynamics is needed to assure conservation of these riparian forests. ?? 2007 Springer Science+Business Media, LLC.

Environmental Management↗

A stand-replacing fire history in upper montane forests of the southern Rocky Mountains

Dendroecological techniques were applied to reconstruct stand-replacing fire history in upper montane forests in northern New Mexico and southern Colorado. Fourteen stand-replacing fires were dated to 8 unique fire years (1842–1901) using four lines of evidence at each of 12 sites within the upper Rio Grande Basin. The four lines of evidence were ( i ) quaking aspen ( Populus tremuloides Michx.) inner-ring dates, ( ii ) fire-killed conifer bark-ring dates, ( iii ) tree-ring width changes or other morphological indicators of injury, and ( iv ) fire scars. The annual precision of dating allowed the identification of synchronous stand-replacing fire years among the sites, and co-occurrence with regional surface fire events previously reconstructed from a network of fire scar collections in lower elevation pine forests across the southwestern United States. Nearly all of the synchronous stand-replacing and surface fire years coincided with severe droughts, because climate variability created regional conditions where stand-replacing fires and surface fires burned across ecosystems. Reconstructed stand-replacing fires that predate substantial Anglo-American settlement in this region provide direct evidence that stand-replacing fires were a feature of high-elevation forests before extensive and intensive land-use practices (e.g., logging, railroad, and mining) began in the late 19th century.

Canadian Journal of Forest Research↗

Multilocus phylogeography and systematic revision of North American water shrews (genus: Sorex )

North American water shrews, which have traditionally included Sorex alaskanus , S. bendirii , and S. palustris , are widely distributed through Nearctic boreal forests and adapted for life in semiaquatic environments. Molecular mitochondrial signatures for these species have recorded an evolutionary history with variable levels of regional divergence, suggesting a strong role of Quaternary environmental change in speciation processes. We expanded molecular analyses, including more-comprehensive rangewide sampling of specimens representing North American water shrew taxa, except S. alaskanus , and sequencing of 4 independent loci from the nuclear and mitochondrial genomes. We investigated relative divergence of insular populations along the North Pacific Coast, and newly recognized diversity from southwestern montane locations, potentially representing refugial isolates. Congruent independent genealogies, lack of definitive evidence for contemporary gene flow, and high support from coalescent species trees indicated differentiation of 4 major geographic lineages over multiple glacial cycles of the late Quaternary, similar to a growing number of boreal taxa. Limited divergence of insular populations suggested colonization following the last glacial. Characterization of southwestern montane diversity will require further sampling but divergence over multiple loci is indicative of a relictual sky-island fauna. We have reviewed and revised North American water shrew taxonomy including the recognition of 3 species within what was previously known as S. palustris . The possibility of gene flow between most distantly related North American water shrew lineages coupled with unresolved early diversification of this group and other sibling species reflects a complex but potentially productive system for investigating speciation processes.

Journal of Mammalogy↗

Chemical weathering in the Loch Vale Watershed, Rocky Mountain National Park, Colorado

Mineralogic, hydrologic, and geochemical data were used to determine the source of solutes to surface waters draining the Loch Vale Watershed (LVWS), an alpine-subalpine drainage located in the Front Range of Colorado. The flux of dissolved solids from LVWS is primarily controlled by interactions between snowmelt and materials derived from the local bedrock; the biomass has only a minor effect on solute budgets except for ammonium. LVWS is underlain by Precambrian granite and gneiss, the major minerals include quartz, microcline, plagioclase, biotite, and sillimanite. Small amounts of calcite were found along hydrothermally altered zones in the bedrock. Mass balance calculations indicate that the weathering of calcite contributes nearly 40% of the cations derived within the basin. The importance of calcite weathering in LVWS is a result of its chemical reactivity and the high rate of physical erosion in this alpine environment. The average cationic denudation rate in the drainage (390 eq/ha/yr) is similar to long-term rates in forested Adirondack watersheds (500–600 eq/ha/yr), but much lower than the average for the North American Continent (3800 eq/ha/yr). Surface waters in LVWS are susceptible to acidification should acid deposition from the atmosphere increase.

Colorado↗

Population declines in North American birds that migrate to the neotropics

Using data from the North American Breeding Bird Survey, we determined that most neotropical migrant bird species that breed in forests of the eastern United States and Canada have recently (1978-1987) declined in abundance after a period of stable or increasing populations. Most permanent residents and temperate-zone migrants did not show a general pattern of decrease during this period. Field data from Mexico were used to classify a subset of the neotropical migrants as using forest or scrub habitats during winter. Population declines during 1978-1987 were significantly greater among the forest-wintering species, while populations of scrub-wintering species increased. The same subset of neotropical migrants also showed overall declines in forest-breeding species, but no significant differences existed between species breeding in forest and scrub habitats. Neotropical migrant species that primarily use forested habitats in either wintering or breeding areas are declining, but a statistically significant association between habitat and population declines was detected only in the tropics.

Proceedings of the National Academy of Sciences↗

Decadal shifts in the population growth, regeneration, and health of Taxodium distichum in swamps of the Cache River Watershed, Illinois

Population projection models based on long-term trends in regeneration and tree survival can be used to predict the future stability of swamp forest species using water management. Population growth and regeneration of a foundational tree species in North American cypress swamps ( Taxodium distichum ) were compared in the Cache River watershed of southern Illinois USA over several decades. This study examined T. distichum population growth in several regional swamps within the Cache River watershed along a moisture gradient to model growth, stability, or decline based on data of life history stage transitions, using data from the following time periods: the 1990s, 2002–2011, and 2012–2022. Using data from the 1990s, T. distichum populations in Crawford Tract in Buttonland Swamp were projected to increase over time because of the high frequency of juvenile stages at elevations with growing season drawdown and winter flooding. The situation in Crawford Tract had changed by 2019–2021 because T. distichum seedlings had not transitioned (i.e., saplings and young trees were absent) although no old-growth trees had died during or after the 1990s. In other regional swamps, projection models were constructed over two decades (2006–2021) along a dry-to-wet flood gradient including Section 8 Woods, Deer Pond, Wildcat Bluff, Snake Hole, and Heron Pond. Populations in swamps on the drier end of the gradient had more support from juvenile stages. In a tree health survey in 2022 within the impounded interior of Buttonland Swamp (not Crawford Tract), seventy percent of the old-growth T. distichum trees were stressed or declining. Coupled with the fact that no seedlings or saplings were observed, the populations in the interior of the swamp were not stable. Many factors could further stress these ancient T. distichum including interactions of impoundment with chemicals, disease, and changes in air temperature and precipitation. Overall, the seedling abundance in these swamps increased in these environments from 2012 to 2021, except in the most flooded swamps. From a management perspective, drier conditions were more conducive to the success of the earlier life history stages of T. distichum in these old-growth forests.

Illinois↗

Ants of the national park of American Samoa

American Samoa makes up the eastern end of the Samoan Archipelago. On the islands of Tutuila, Taʽū and Ofu, the National Park of American Samoa (NPSA) protects about 4,000 ha of coastal, mid-slope and ridge-top forest. While the ant fauna of the Samoan Archipelago is considered relatively well documented, much of NPSA has never been surveyed for ants, leaving the fauna and its distribution poorly known. To address this shortfall, we systematically surveyed ants within the Tutuila and Taʽū units of NPSA using standard methods (hand collecting, litter sifting, and baits) at 39 sites within six vegetation types ranging from 8 to 945 m elevation. Forty-four ant species were identified, 19 of which are exotic to the Samoan Archipelago. Two notoriously aggressive species, Anoplolepis gracilipes and Pheidole megacephala were detected at two and seven sites, respectively. Both of these species largely excluded all other ants from bait, although their impact on ant community composition is unclear. A suite of habitat variables measured at each site was assessed to explain park-wide ant distributions. Of eight variables evaluated, only elevation was associated with ant community structure, as the ratio of native to exotic ant species increased significantly with elevation on Tutuila. Our survey documented two species not previously reported from American Samoa. Strumigenys eggersi, detected at 12 sites, appears to be a new immigrant to the Pacific Basin. A species of Pheidole was collected that likely represents an undescribed species. Solenopsis geminata, an aggressive species first reported on Tutuila in 2002, was not detected during our survey.

America Samoa↗

Evaluating legacy effects of hyperabundant white-tailed deer (Odocoileus virginianus) in forested stands of Harriman and Bear Mountain State Parks, New York

Executive Summary White-tailed deer (Odocoileus virginianus) are among the most impactful herbivores in the eastern United States. Legacy forest effects, those accrued from intense herbivory over time, manifest as low seedling regeneration, high cover of plant species that are infrequently browsed by deer, presence or expansion of nonnative or invasive plant species, few herbaceous species, and diminished capacity for recovery. Interfering vegetation (that is, species that increase in cover and density due to avoidance by deer, such as American beech sprouts, Pennsylvania sedge, and hay-scented fern) increase competition for light and hinder recruitment of trees into the forest canopy. The lower Hudson Valley in New York has been heavily browsed by white-tailed deer since the early 20th century. The region has some of the lowest tree regeneration rates in New York State as a result of deer browsing and subsequent increases in interfering vegetation. The U.S. Geological Survey and the State University of New York College of Environmental Science and Forestry studied sites where deer hunting is permitted (case sites) and nearby sites where hunting is currently prohibited (control sites) to assess and identify forest structure and composition differences. Instead of using deer exclosures, which are time-consuming and expensive to install and maintain, we used a case-control study because such studies are well-suited to effects with long latency and rare outcomes. Case-control studies seek to describe the relation between an outcome of interest (in this study, forest understory recovery from chronic herbivory) and forest condition. We inferred recovery by comparing these characteristics on adjacent sites in the lower Hudson Valley with similar forest communities and land uses but different deer population management histories. Case plots were on lands where deer management has taken place annually for several decades. Control plots were on lands where deer populations have not been consistently managed to lowered abundance. We accounted for differences in forest recovery not attributable to deer by first matching case and control plots along several important environmental gradients (slope, aspect, elevation, moisture, canopy openness). By controlling for these gradients, we looked for associations between measured forest conditions and deer herbivory reduction through population management. We surveyed more than 200 plots in upland forest types across case and control sites where we assessed forest condition by estimating density (number per unit area) and composition and cover (percent) of important vegetation constituents in ground, shrub, subcanopy, and canopy layers of the forest. We recorded 37 tree species, 22 shrub species, 57 herbaceous species, and 19 species of grasses and sedges in our plot surveys, including a number of nonnative and invasive plants. We also estimated the ages of a number of common canopy trees by counting rings from cores extracted from individual stems. Effects of more than 100 years of chronic deer browsing manifested in low herbaceous ground cover and little to no tree recruitment (saplings) on lands without deer management. In contrast, sustained deer management resulted in forests with conditions that indicated substantial recovery from chronic herbivory in the ground, shrub, and subcanopy layers. Sites with ongoing deer management exhibited greater ground cover of tree seedlings and herbs and less ground cover of interfering vegetation and nonnative species. The well-developed sub-canopy layer of small trees, saplings, and tall shrubs on sites with deer management indicates a high potential for sapling recruitment to the canopy of the future forest. Of the 25 subcanopy trees sampled on control sites, most were more than 100 years old, indicating little to no regeneration in areas sampled for more than 100 years. The forest canopy, a relic of land uses of bygone days, requires a source of young trees to replace itself as older trees die. Without an abundant layer of young trees in the subcanopy, a forest cannot be sustained over time. Reduction in deer herbivory promotes forest recovery and could benefit Harriman and Bear Mountain State Parks (the control sites for the study), but removal of interfering vegetation may be necessary to mitigate legacy effects where they currently hinder ground layer recovery. To successfully promote a more desirable forest condition that includes elimination of nonnative plant species, promotion of tree recruitment into the forest canopy, and development of diverse and abundant herbaceous cover in ground layer vegetation, future management decisions could include information on herbivory reduction and management of interfering vegetation where necessary.

New York↗

The effects of ENSO and the North American monsoon on mast seeding in two Rocky Mountain conifer species

We aimed to disentangle the patterns of synchronous and variable cone production (i.e. masting) and its relationship to climate in two conifer species native to dry forests of western North America. We used cone abscission scars to reconstruct ca 15 years of recent cone production in Pinus edulis and Pinus ponderosa , and used redundancy analysis to relate time series of annual cone production to climate indices describing the North American monsoon and the El Niño Southern Oscillation (ENSO). We show that the sensitivity to climate and resulting synchrony in cone production varies substantially between species. Cone production among populations of P. edulis was much more spatially synchronous and more closely related to large-scale modes of climate variability than among populations of P. ponderosa . Large-scale synchrony in P. edulis cone production was associated with the North American monsoon and we identified a dipole pattern of regional cone production associated with ENSO phase. In P. ponderosa , these climate indices were not strongly associated with cone production, resulting in asynchronous masting patterns among populations. This study helps frame our understanding of mast seeding as a life-history strategy and has implications for our ability to forecast mast years in these species.

Philosophical Transactions of the Royal Society B:↗

Modeled effects of soil acidification on long-term ecological and economic outcomes for managed forests in the Adirondack region (USA)

Sugar maple ( Acer saccharum ) is among the most ecologically and economically important tree species in North America, and its growth and regeneration is often the focus of silvicultural practices in northern hardwood forests. A key stressor for sugar maple (SM) is acid rain, which depletes base cations from poorly-buffered forest soils and has been associated with much lower SM vigor, growth, and recruitment. However, the potential interactions between forest management and soil acidification &ndash; and their implications for the sustainability of SM and its economic and cultural benefits &ndash; have not been investigated. In this study, we simulated the development of 50 extant SM stands in the western Adirondack region of NY (USA) for 100 years under different soil chemical conditions and silvicultural prescriptions. We found that interactions between management prescription and soil base saturation will strongly shape the ability to maintain SM in managed forests. Below 12% base saturation, SM did not regenerate sufficiently after harvest and was replaced mainly by red maple ( Acer rubrum ) and American beech ( Fagus grandifolia ). Loss of SM on acid-impaired sites was predicted regardless of whether the shelterwood or diameter-limit prescriptions were used. On soils with sufficient base saturation, models predicted that SM will regenerate after harvest and be sustained for future rotations. We then estimated how these different post-harvest outcomes, mediated by acid impairment of forest soils, would affect the potential monetary value of ecosystem services provided by SM forests. Model simulations indicated that a management strategy focused on syrup production &ndash; although not feasible across the vast areas where acid impairment has occurred &ndash; may generate the greatest economic return. Although pollution from acid rain is declining, its long-term legacy in forest soils will shape future options for sustainable forestry and ecosystem stewardship in the northern hardwood forests of North America.

New York↗

Influence of the diversion of Bear River into Bear Lake (Utah and Idaho) on the environment of deposition of carbonate minerals

Bear River, the largest river in the Great Basin, had some of its flow diverted into Bear Lake through a series of canals constructed between 1911 and 1918, turning Bear Lake into a reservoir. The prediversion lake had an unusually high Mg2+ : Ca2+ ratio (38 by weight), which resulted in precipitation of CaCO3 as aragonite. The amount and mineralogy of the carbonate did not change immediately after the diversion, but during the middle of the 20th century, high-Mg calcite began to precipitate. In contrast, at the time of diversion there were very distinct changes in the isotopic composition of the carbonate that clearly define the time of diversion within the sediments. Sediment-trap studies show that the CaCO3 polymorph that precipitates in the epilimnion of the lake today is high-Mg calcite. Samples in sediment traps placed 2 m above the bottom of the lake in 45 m of water (bottom traps) contain predominantly aragonite and quartz, with lesser amounts of high-Mg and low-Mg calcite and dolomite. Isotopic evidence from the sediments shows that the aragonite that accumulated in the bottom sediment traps and that constitutes the bulk of postdiversion sediments is reworked and redistributed from shallow sites to deep sites. ?? 2007, by the American Society of Limnology and Oceanography, Inc.

Limnology and Oceanography↗

Western North American shorebirds

Shorebirds are a diverse group that includes oystercatchers, stilts, avocets, plovers, and sandpipers. They are familiar birds of seashores, mudflats, tundra, and other wetlands, but they also occur in deserts, high mountains, forests, and agricultural fields. Widespread loss and alteration of these habitats, especially wetlands and grasslands during the past 150 years, coupled with unregulated shooting at the turn of the century, resulted in population declines and range contractions of several species throughout North America. In the western portion of the continent, efforts to monitor the status and trends of shorebirds have been in effect for only the past 15-25 years and for only a few species. Methods exist to monitor population trends for most shorebirds, but only broadscale, international efforts, relying largely on volunteer help, will accomplish this. In this article we address shorebirds primarily in western North America, the region west of the Continental Divide from northern Alaska to southern Mexico. The 12 states, a Canadian province and territory, and the western portion of Mexico within this region represent about 25% of the North American landmass (Fig. 1). Western North America includes portions of three broad ecological domains: the Polar Domain, encompassing the tundra and boreal forests that cover most of Greenland, Canada, and Alaska; the Humid Temperate Domain, including the humid midlatitude forests and shrublands within the United States, southern portions of the Canadian prairie provinces, and along the west coast of North America; and the Dry Domain, encompassing the short-grass prairies, sagebrush provinces, and deserts (Fig. 1; Bailey 1978, 1989).

Book chapter↗