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Research about North Carolina, Tennessee

Source-linked reports with geographic coverage including North Carolina, Tennessee.

At least 19 recordsLinked to original sources

Development of genomic markers for monitoring and research on plethodontid salamanders

Despite the importance of plethodontid salamanders and their vulnerability to ongoing environmental change, they are inherently difficult to monitor due to their cryptic nature. Recent advances in genomics have created new opportunities for monitoring of populations and their responses to environmental perturbations. In this study, we developed a new target capture-based genomic panel for the purposes of genetic monitoring in plethodontid salamanders. We demonstrate its utility in several distantly related species and present an example application in two representative species with co-occurring distributions but different ecological attributes and expected patterns of population structure: Plethodon jordani and Desmognathus wrighti . Although the number of successfully assembled loci declined with phylogenetic distance from the original reference species ( Desmognathus spp), we obtained high-quality data from thousands of loci from species in all four genera tested ( Desmognathus , Plethodon , Eurycea , and Gyrinophilus ), which span the deepest split in Plethodontidae. Landscape genetic analyses detected weak but statistically significant geographic structure in P. jordani , and much stronger geographic structure in D. wrighti , as expected based on the lower population density and likely lower dispersal ability of D. wrighti . Our target capture panel is broadly applicable across salamanders in Plethodontidae and has the potential to provide data for a wide range of phylogenetic, biogeographic, and population genetics research questions.

North Carolina, Tennessee

Genetic structure of restored Brook Trout populations in the Southern Appalachian Mountains indicates successful reintroductions

Wildlife reintroduction is an important conservation tool for threatened species, yet identifying appropriate source populations poses a challenge. In particular, the possibility of outbreeding depression is cited as a constraint limiting the range of candidate source populations for translocation. When multiple source lineages are mixed during reintroduction, genetic monitoring is necessary to evaluate whether sources contribute equally to subsequent generations and whether they are interbreeding as expected. Moreover, statistical analysis of genetic data should account for complex life histories that might affect the timescale of admixture and genetic drift. Here, we use samples collected over a 23-year period and a stochastic age-structured model to analyze the genetic mixing process in reintroduced Brook Trout ( Salvelinus fontinalis ) populations in the Southern Appalachians. Each restored population was seeded with two to three source populations. Previous research inferred reproductive isolation between source populations leading to a proposal of splitting the species into multiple taxa. In contrast, we found patterns of ancestry that were consistent with random mating and no advantage for one source lineage over any other. Brook Trout from different source streams are mixing as expected in the restoration sites. This result does not support the hypothesis that Brook Trout in the Southern Appalachian Mountains includes several distinct species. Mixing different sources from the same watershed seems to be an effective way to increase genetic diversity of reintroduced populations while minimizing risk to source populations.

North Carolina, Tennessee

Fuels and vegetation changes in southwestern, unburned portions of Great Smoky Mountains National Park, USA, 2003-2019

Overstory basal area, ericaceous shrub cover ( Kalmia latifolia L. and Rhododendron maximum L.), and fuels (i.e., woody fuel loads and depths and O Horizon thickness) were assessed within Great Smoky Mountains National Park, USA, in 2003 − 2004. Due to recent wildfire activity within the southern Appalachian Mountain region (including Great Smoky Mountains National Park), the potential spread and expansion of ericaceous shrubs, and the impacts of the hemlock woolly adelgid ( Adelges tsugae Annand) on eastern hemlock ( Tsuga canadensis (L.) Carrière), these same ecosystem components were again assessed in 2019. Elevation and moisture regime (xeric, intermediate, and mesic) were included in this assessment as potential influential factors. An evaluation of repeated measurements from 40 plots suggested that O Horizon thickness did not change significantly over the 16-year period, but increased as elevation increased, and moisture regime (xeric O Horizon thickness > mesic O Horizon thickness) was a significant, related factor. The sum of 1-, 10-, and 100-h fuel loads (fuels less < 7.6 cm diameter) increased, whereas woody fuel depth decreased over the 16-year period. No significant changes in 1000-h fuel loads (> 7.6 cm diameter), total woody fuel loads, ericaceous shrub cover, total basal area, or live T. canadensis basal area were observed. Live T. canadensis basal area decreased with increasing elevation. Dead, standing T. canadensis basal area increased from 2003–2019, and that increase was most pronounced as elevation increased on xeric and intermediate sites. Overall, we found that: 1. hypothesized increases in total woody fuel loads and ericaceous shrub cover were not present; and 2. elevation and moisture regime were most related to observed changes in vegetation and fuel condition.

North Carolina, Tennessee

National Park Service Vegetation Mapping Inventory Program: Great Smoky Mountains National Park vegetation mapping project

The National Park Service (NPS) Vegetation Mapping Inventory (VMI) Program is an effort to classify, describe, and map existing vegetation communities in national park units throughout the United States. The NPS VMI Program is managed by the NPS Natural Resource Stewardship and Science Inventory and Monitoring Program and provides baseline vegetation information to natural resource managers, researchers, and ecologists. The U.S. Geological Survey Upper Midwest Environmental Sciences Center, NatureServe, and NPS Great Smoky Mountains National Park (GRSM, also referred to as the “Park”) have completed vegetation classification and mapping of GRSM, including the Foothills Parkway, for the NPS VMI Program. Mappers, ecologists, and botanists collaborated to affirm vegetation types of GRSM and to determine how best to map the vegetation types by using aerial imagery. A vegetation classification developed in 2003 by NatureServe and the NPS served as a foundation to further classify and map the vegetation types of the Park. Data from an additional 10 vegetation plots supported vegetation types either rare or not documented in the 2003 classification. Data from 203 verification sites were collected to test the field key to vegetation types and the application of vegetation types to a sample set of map polygons. Furthermore, data from 972 accuracy assessment (AA) sites were collected (of which 966 were used to test accuracy of the vegetation map layer). This GRSM vegetation mapping project identified 112 vegetation types consisting of 105 association types in the U.S. National Vegetation Classification (USNVC), 2 “park-special” types, 1 “map-special” type, and 4 cultural types in the USNVC. To map the vegetation and land cover of GRSM, 52 map classes were developed. Of these 52 map classes, 46 represent natural (including ruderal) vegetation types, most of which types are recognized in the USNVC. For the remaining 6 of the 52 map classes, 4 represent USNVC cultural types for agricultural and developed areas, and 2 represent non-USNVC types for nonvegetated open water and nonvegetated rock. Features were interpreted from viewing four-band digital aerial imagery using digital onscreen three-dimensional stereoscopic workflow systems in geographic information systems; digital aerial imagery was collected during September 23–October 30, 2015. The interpreted data were digitally and spatially referenced, thus making the spatial-database layers usable in a geographic information system. Polygon units were mapped to either a 0.5- or 0.25- hectare (ha) minimum mapping unit, depending on vegetation type. A geodatabase containing several feature-class layers and tables provides the locations and data of USNVC vegetation types (vegetation map layer), vegetation plots, verification sites, AA sites, project boundary extent, and aerial image centers and flight lines. Covering 210,875 ha, the feature-class layer and related tables for the vegetation map layer provide 34,084 polygons of detailed attribute data when special modifiers are not considered (average polygon size of 6.2 ha) and 36,589 polygons of detailed attribute data when special modifiers are considered (average polygon size of 5.8 ha). Each map polygon is assigned a map-class code and name and, when applicable, are linked to USNVC classification tables within the geodatabase. The vegetation map extent includes the administrative boundary for GRSM and the Foothills Parkway. A summary report, generated from the vegetation map layer, concludes that the 46 map classes representing natural (including ruderal) vegetation types apply to 99.2% of polygons (33,797 polygons; average size of 6.2 ha) and cover 98.6% of the Park (207,971.4 ha). Further broken down, map classes representing natural vegetation types indicate that the Park is 97.7% forest and woodland (205,882.5 ha), 0.6% shrubland (1,174.6 ha), and 0.4% herbaceous (914.3 ha). Map classes representing cultural vegetation types apply to 0.8% of polygons (259 polygons; average size of 4.9 ha) and cover 0.6% of the Park (1,277.4 ha). Map classes representing nonvegetation open and flowing water and unvegetated rock apply to 0.08% of polygons (28 polygons; average size of 58.1 ha) and cover 0.8% of the Park (1,625.9 ha). A thematic AA study was completed of map classes representing the natural (including ruderal) vegetation types of the Park. Initial AA results were discussed with NPS staff from the Park. Following input from NPS staff on how to handle map classes that fell below accuracy standards, adjustments were made to the vegetation map layer. Final results indicate an overall accuracy of 80.64% (kappa index of 79.96% for chance agreements) based on data from 966 of the 972 AA sites. Most individual map-class themes exceed the NPS VMI Program standard of 80% with a 90% confidence interval. The GRSM vegetation mapping project delivers many geospatial and vegetation data products, including an in-depth project report discussing methods and results, which includes map classification and map-class descriptions. This suite of products also includes descriptions and a field key to vegetation types; a database of vegetation plots, verification sites, and AA sites; digital images of field sites; field data sheets; digital aerial imagery; hardcopy and digital maps; a geodatabase of vegetation and land cover (map layer), field sites (vegetation plots, verification sites, and AA sites), aerial imagery index, project boundary, and metadata; and a contingency table listing AA results. Geospatial products are projected in the Universal Transverse Mercator, Zone 17 North, by using the North American Datum of 1983. Information on the NPS VMI Program and completed mapping projects are on the internet at https://www.nps.gov/im/vegetation-inventory.htm.

North Carolina, Tennessee

Survival and conflict behavior of American black bears after rehabilitation

Wildlife agencies face difficult situations when orphaned or injured American black bear ( Ursus americanus ) cubs (<12 months old) or yearlings (≥12 and <24 months old) are captured. One option is bear rehabilitation, the care and feeding of cubs or yearlings in a semi‐natural environment, followed by release. Unfortunately, the survival and movements of bears released from rehabilitation facilities are often poorly documented and the ultimate reasons for success or failure poorly understood. Our goal was to assess survival and post‐release conflict of orphaned bear cubs and yearlings following release from a rehabilitation facility, Appalachian Bear Rescue (ABR), in Townsend, Tennessee, USA, from 2015–2016. We predicted that rehabilitated bears would survive at similar rates, die from similar causes, and engage in similar conflict behavior to wild conspecifics. We equipped 42 black bear cubs and yearlings from ABR with global positioning system‐collars and released them in Great Smoky Mountains National Park or Cherokee National Forest, Tennessee and North Carolina, USA. Estimated annual survival using known‐fate methods for all released bears was 0.93 ± 0.06 [SE]). Survival for 13 bears released as cubs was 0.64 ± 0.14, whereas none of the bears released as yearlings died within 1 year after release ( n = 29). Survival of rehabilitated bears was similar to or higher than published rates for wild conspecifics. Three of 42 bears (7.1%) released from ABR engaged in conflict behavior up to 1 year following release, and those had spent time involved in conflict behavior with their mothers (e.g., approaching humans) prior to being orphaned. Despite not having the typical post‐natal experience with their mothers, the bears in our study appeared to behave and survive similarly to their wild conspecifics. Rehabilitation is effective for managing orphaned or injured bears. Best survival occurred for bears released as yearlings; however, managers can maximize cub survival through fall releases when plentiful wild foods are available.

North Carolina, Tennessee

Activity patterns and temporal predator avoidance of white-tailed deer (Odocoileus virginianus) during the fawning season

In the presence of a predator, prey may alter their temporal activity patterns to reduce the risk of an encounter that may induce injury or death. Prey perception of predation risk and antipredator responses may increase in the presence of dependent offspring. We conducted a camera trap study during summer 2015 in North Carolina and Tennessee, USA to evaluate temporal avoidance of a predator (coyote Canis latrans ) by white-tailed deer ( Odocoileus virginianus ). We analyzed activity patterns of bucks, does, and nursery groups (i.e., groups that included fawns) relative to those of coyotes to determine the coefficient of overlap (Δ) using a kernel density estimator. We found that bucks and does had similar Δ with coyotes [Δ 1 = 0.729 (0.629–0.890) and Δ 1 = 0.686 (0.558–0.816, respectively] and exhibited crepuscular activity patterns comparable to those of coyotes. However, nursery groups displayed a dramatically different activity pattern: unimodal activity was concentrated in the middle of the day with little overlap with coyote activity [Δ 1 = 0.362 (0.176–0.491)]. Because adult deer are rarely prey for coyotes, whereas fawns are common prey during summer, the shift in activity patterns of nursery groups demonstrates a behavioral shift likely aimed at avoiding coyote predation on fawns.

North Carolina, Tennessee

Relationships between indicators of acid-base chemistry and fish assemblages in streams of the Great Smoky Mountains National Park

The acidity of many streams in the Great Smoky Mountains National Park (GRSM) has increased significantly since pre-industrial (∼1850) times due to the effects of highly acidic atmospheric deposition in poorly buffered watersheds. Extensive stream-monitoring programs since 1993 have shown that fish and macroinvertebrate assemblages have been adversely affected in many streams across the GRSM. Matching chemistry and fishery information collected from 389 surveys performed at 52 stream sites over a 22-year period were assessed using logistic regression analysis to help inform the U.S. Environmental Protection Agency’s assessment of the environmental impacts of emissions of oxides of nitrogen (NO x ) and sulfur (SO x ). Numerous logistic equations and associated curves were derived that defined the relations between acid neutralizing capacity (ANC) or pH and different levels of community richness, density, and biomass; and density and biomass of brook trout, rainbow trout, and small prey (minnow) populations in streams of the GRSM. The equations and curves describe the status of fish assemblages in the GRSM under contemporary emission levels and deposition loads of nitrogen (N) and sulfur (S) and provide a means to estimate how newly proposed (and various alternative) target deposition loads, which strongly influence stream ANC, might affect key ecological indicators. Several examples using ANC, community richness, and brook trout density are presented to illustrate the steps needed to predict how future changes in stream chemistry (resulting from different target deposition loads of N and S) will affect the probabilities of observing specific levels of selected biological indicators in GRSM streams. The implications of this study to the regulation of NO x and SO x emissions, water quality, and fisheries management in streams of the GRSM are discussed, but also qualified by the fact that specific examples provided need to be further explored before recommendations concerning their use as ecological indicators could be proposed.

North Carolina, Tennessee

Karst hydrogeology of Tuckaleechee Cove and the western Great Smoky Mountains, Tennessee and North Carolina

The geology of Great Smoky Mountains National Park (GRSM) in Tennessee and North Carolina is dominated by siliciclastics and metamorphic strata. However, in the western portion of GRSM, a series of carbonate fensters (windows) expose the Lower Ordovician–age section of the Knox Group, a series of dolomite and limestone units that are partially marbleized as a result of contact metamorphism from the Great Smoky fault. The fensters create opportunities for allogenic recharge to occur at points along the contact of the surrounding insoluble strata with the underlying soluble carbonates. The combination of chemically aggressive surface recharge and vertical relief has resulted in the formation of deep caves, many of which have active streams and water resources. Though the karst is limited in extent and the number of caves is fairly small, the significance of the resources is substantial, with several of the caves in the area over 150 m in depth and at least two being major bat hibernacula. In 2017, the U.S. Geological Survey (USGS) began a study to better understand the hydrologic behavior of these karst systems through hydrologic and geochemical monitoring, groundwater tracing using fluorescent dyes, and seepage runs. Stage and water-quality instrumentation was installed in two caves in GRSM, the main stream of Bull Cave, and in a sump pool in Whiteoak Blowhole, at 173 m and 70 m below land surface, respectively. Following setup of the cave sites, dye injections were conducted to determine discharge points for four of the deep cave systems on Rich Mountain and Turkeypen ridge. Results show water in these systems has an extremely rapid travel time, with tracers detected from caves to springs in less than 24 h for each of the systems. This field guide describes the complex geology, regional hydrogeology, and unique landscape characterized by high-gradient subterranean streams, carbonate fensters, and deep caves of the GRSM karst.

North Carolina, Tennessee

The Southern Appalachian Brook Trout management conundrum: What should restoration look like in the 21st Century?

Brook Trout Salvelinus fontinalis in the southern Appalachian portion of their range have been isolated in remote headwater systems for millennia. Recent genetic investigations indicate extremely low allelic diversity, heterozygosity and effective population sizes in many streams. In populations restored using multiple source stocks, limited introgression has been observed despite source stocks being collected from streams within the same subwatershed. It remains unclear if pre- and/or post-reproductive isolating mechanisms are restricting effective gene flow among source stocks in restored streams. Objectives of this study were to: 1) identify environmental variables contributing to assortative mating, and 2) use common garden crossings to determine if wild type brood stock crossings resulted in physiologically viable offspring. We observed markedly different fertilization success rates within-population (66.7%) and betweenpopulation (91.7%) from the 42 crosses (N=18 control, N=24 treatment). Moreover, we observed significant (P < 0.05) differences between within-population and between-population groups in each of our linear mixed effects global models for each trial stage of development (i.e., fertilization rate, eyed egg rate, and hatch rates). Tukey’s HSD comparisons revealed only one significantly (P < 0.003) different fertilization rate among the forty five pairwise comparisons in each of our three stages of trails. In addition, we observed differential peaks of gamete production within and among source stream brood stock, despite common garden conditions, that appeared to have limited fertilization success rates between interstream and control groups. Despite differential peak gamete timing, intrastream crosses performed equally, and, in some instances, better than those between control groups. Our results suggest differential responses to shared environmental conditions (i.e., temperature and/or photoperiod) may contribute to mismatched spawning phenology (i.e., gamete production timing) among restoration founder stocks leading to introgression (i.e., genetic admixture). The application of contemporary genetic techniques could help determine if these possible local adaptations are genetically fixed or may break down over time in restored populations with mixed source stocks. These findings demonstrate the need to apply contemporary conservation genetics tools to future wild trout restoration projects using translocated source stock towards the goal of “genetically-robust”, naturally reproducing populations with the ability to cope with current and future perturbations.

North Carolina, Tennessee

Reevaluating the age of the Walden Creek Group and the kinematic evolution of the western Blue Ridge, southern Appalachians

An integrated synthesis of existing datasets (detailed geologic mapping, geochronologic, paleontologic, geophysical) with new paleontologic and geochemical investigations of rocks previously interpreted as part of the Neoproterozoic Walden Creek Group in southeastern Tennessee suggest a necessary reevaluation of the kinematics and structural architecture of the Blue Ridge Foothills. The western Blue Ridge of Tennessee, North Carolina, and Georgia is composed of numerous northwest-directed early and late Paleozoic thrust sheets, which record pronounced variation in stratigraphic/structural architecture and timing of metamorphism. The detailed spatial, temporal, and kinematic relationships of these rocks have remained controversial. Two fault blocks that are structurally isolated between the Great Smoky and Miller Cove-Greenbrier thrust sheets, here designated the Maggies Mill and Citico thrust sheets, contain Late Ordovician-Devonian conodonts and stable isotope chemostratigraphic signatures consistent with a mid-Paleozoic age. Geochemical and paleontological analyses of Walden Creek Group rocks northwest and southeast of these two thrust sheets, however, are more consistent with a Late Neoproterozoic (550&ndash;545 Ma) depositional age. Consequently, the structural juxtaposition of mid-Paleozoic rocks within a demonstrably Neoproterozoic-Cambrian succession between the Great Smoky and Miller Cove-Greenbrier thrust sheets suggests that a simple foreland-propagating thrust sequence model is not applicable in the Blue Ridge Foothills. We propose that these younger rocks were deposited landward of the Ocoee Supergroup, and were subsequently plucked from the Great Smoky fault footwall as a horse, and breached through the Great Smoky thrust sheet during Alleghanian emplacement of that structure.

North Carolina, Tennessee

Geologic map of the Great Smoky Mountains National Park region, Tennessee and North Carolina

The geology of the Great Smoky Mountains National Park region of Tennessee and North Carolina was studied from 1993 to 2003 as part of a cooperative investigation by the U.S. Geological Survey with the National Park Service (NPS). This work resulted in a 1:100,000-scale geologic map derived from mapping that was conducted at scales of 1:24,000 and 1:62,500. The geologic data are intended to support cooperative investigations with the NPS, the development of a new soil map by the Natural Resources Conservation Service, and the All Taxa Biodiversity Inventory. In response to a request by the NPS, we mapped previously unstudied areas, revised the geology where problems existed, and developed a map database for use in interdisciplinary research, land management, and interpretive programs for park visitors.

North Carolina, Tennessee

40 Ar/ 39 Ar age-spectrum data for hornblende, biotite, white mica, and K-feldspar samples from metamorphic rocks in the Great Smoky Mountains of North Carolina and Tennessee

This report contains reduced 40 Ar/ 39 Ar data of hornblende, biotite, white mica and (or) sericite, and potassium-feldspar mineral separates and phyllite groundmass samples from metamorphic rocks of the Great Smoky Mountains in North Carolina and Tennessee. Included in this report are information on the location of the samples and a brief description of the samples. The data contained herein are not interpreted in a geological context, and care should be taken by users unfamiliar with argon isotopic data in the use of these results. No geological meaning is implied for any of the apparent ages presented below, and many of the individual apparent ages are not geologically meaningful. This report is primarily a detailed source document for subsequent publications that will integrate these data into a geological context. All the samples in this report were collected in and around the Great Smoky Mountain National Park in western North Carolina and eastern Tennessee.

North Carolina, Tennessee

Comparison of breeding bird and vegetation communities in primary and secondary forests of Great Smoky Mountains National Park

We compared breeding bird communities and vegetation characteristics at paired point locations in primary (undisturbed) and mature secondary forest (70-100 years old) sites in Great Smoky Mountains National Park, USA to understand how sites logged prior to creation of the park compare to undisturbed sites following 70 years of protection from human disturbance. We found that bird and vegetation communities are currently similar, but retain some differences in species composition. Rank abundance curves for primary and secondary forest bird communities showed very similar patterns of species dominance. Species composition was also similar on the two sites which shared 24 of the 25 most frequently recorded species. Nonetheless, comparisons of density estimates derived from distance sampling showed three bird species were more abundant on primary forest sites and that one bird species was significantly more abundant on secondary forest sites. Notably, comparisons based on raw counts (unadjusted for potential differences in detectability) produced somewhat different results. Analyses of vegetation samples for the paired sites also showed relative similarity, but with some differences between primary and secondary forests. Primary forest sites had more large trees (trees greater than 50 cm diameter at breast height) and late successional species. Primary forest sites had a denser tall shrub layer while secondary forest sites had a denser canopy layer. Nonetheless, tree species richness, basal area of live trees and number of standing snags did not differ between primary and secondary forest sites. Results indicate that breeding bird communities on sites within the park that were logged commercially 70 years ago are currently quite similar to bird communities on sites with no history of human disturbance. Similarities between the bird communities on previously disturbed and undisturbed sites in Great Smoky Mountains National Park may exceed those on more fragmented landscapes because large patches of primary forest, adjacent to commercially logged sites, remained in the park when it was established in 1935. These patches of primary forest may have served as source areas for commercially logged sites.

North Carolina, Tennessee

Bait stations, hard mast, and black bear population growth in Great Smoky Mountains National Park

Bait-station surveys are used by wildlife managers as an index to American black bear ( Ursus americanus ) population abundance, but the relationship is not well established. Hard mast surveys are similarly used to assess annual black bear food availability which may affect mortality and natality rates. We used data collected in Great Smoky Mountains National Park (GSMNP) from 1989 to 2003 to determine whether changes in the bait-station index (&Delta;BSI) were associated with estimated rates of bear population growth (&lambda;) and whether hard mast production was related to bear visitation to baits. We also evaluated whether hard mast production from previous years was related to &lambda;. Estimates of &lambda; were based on analysis of capture-recapture data with the Pradel temporal symmetry estimator. Using the Akaike's Information Criterion (AIC), our analysis revealed no direct relationship between &Delta;BSI and &lambda;. A simulation analysis indicated that our data were adequate to detect a relationship had one existed. Model fit was marginally improved when we added total oak mast production of the previous year as an interaction term suggesting that the BSI was confounded with environmental variables. Consequently the utility of the bait-station survey as a population monitoring technique is questionable at the spatial and temporal scales we studied. Mast survey data, however, were valuable covariates of &lambda;. Population growth for a given year was negatively related to oak mast production 4 and 5 years prior. That finding supported our hypothesis that mast failures can trigger reproductive synchrony, which may not be evident from the trapped sample until years later.

North Carolina, Tennessee

Generalized geologic map of bedrock lithologies and surficial deposits in the Great Smoky Mountains National Park region, Tennessee and North Carolina

The geology of the Great Smoky Mountain National Park (GSMNP) region of Tennessee and North Carolina was studied from 1993 to 2003 as part of a cooperative investigation with the National Park Service (NPS). This work has been compiled as a 1:100,000-scale map derived from mapping done at 1:24,000 and 1:62,500 scale. The geologic data are intended to support cooperative investigations with NPS, the development of a new soil map by the Natural Resources Conservation Service, and the All Taxa Biodiversity Inventory ( http://www.discoverlifeinamerica.org/ ). At the request of NPS, we mapped areas previously not visited, revised the geology where stratigraphic and structural problems existed, and developed a map database for use in interdisciplinary research, land management, and interpretive programs for park visitors.

North Carolina, Tennessee

Geologic map of the Great Smoky Mountains National Park region, Tennessee and North Carolina

The geology of the Great Smoky Mountain National Park (GSMNP) region of Tennessee and North Carolina was studied from 1993 to 2003 as part of a cooperative investigation with the National Park Service (NPS). This work has been compiled as a 1:100,000-scale map derived from mapping done at 1:24,000 and 1:62,500 scale. The geologic data are intended to support cooperative investigations with NPS, the development of a new soil map by the Natural Resources Conservation Service, and the All Taxa Biodiversity Inventory ( http://www.discoverlifeinamerica.org/ ). At the request of NPS, we mapped areas previously not visited, revised the geology where stratigraphic and structural problems existed, and developed a map database for use in interdisciplinary research, land management, and interpretive programs for park visitors.

North Carolina, Tennessee

Investigating species co-occurrence patterns when species are detected imperfectly

1. Over the last 30 years there has been a great deal of interest in investigating patterns of species co-occurrence across a number of locations, which has led to the development of numerous methods to determine whether there is evidence that a particular pattern may not have occurred by random chance. 2. A key aspect that seems to have been largely overlooked is the possibility that species may not always be detected at a location when present, which leads to 'false absences' in a species presence/absence matrix that may cause incorrect inferences to be made about co-occurrence patterns. Furthermore, many of the published methods for investigating patterns of species co-occurrence do not account for potential differences in the site characteristics that may partially (at least) explain non-random patterns (e.g. due to species having similar/different habitat preferences). 3. Here we present a statistical method for modelling co-occurrence patterns between species while accounting for imperfect detection and site characteristics. This method requires that multiple presence/absence surveys for the species be conducted over a reasonably short period of time at most sites. The method yields unbiased estimates of probabilities of occurrence, and is practical when the number of species is small (< 4). 4. To illustrate the method we consider data collected on two terrestrial salamander species, Plethodon jordani and members of the Plethodon glutinosus complex, collected in the Great Smoky Mountains National Park, USA. We find no evidence that the species do not occur independently at sites once site elevation has been allowed for, although we find some evidence of a statistical interaction between species in terms of detectability that we suggest may be due to changes in relative abundances.

North Carolina, Tennessee