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Research about Florida

Source-linked reports with geographic coverage including Florida.

At least 541 records · Page 30Linked to original sources

Late Holocene sea- and land-level change on the U.S. southeastern Atlantic Coast

Late Holocene relative sea-level (RSL) reconstructions can be used to estimate rates of land-level (subsidence or uplift) change and therefore to modify global sea-level projections for regional conditions. These reconstructions also provide the long-term benchmark against which modern trends are compared and an opportunity to understand the response of sea level to past climate variability. To address a spatial absence of late Holocene data in Florida and Georgia, we reconstructed ~ 1.3 m of RSL rise in northeastern Florida (USA) during the past ~ 2600 years using plant remains and foraminifera in a dated core of high salt-marsh sediment. The reconstruction was fused with tide-gauge data from nearby Fernandina Beach, which measured 1.91 ± 0.26 mm/year of RSL rise since 1900 CE. The average rate of RSL rise prior to 1800 CE was 0.41 ± 0.08 mm/year. Assuming negligible change in global mean sea level from meltwater input/removal and thermal expansion/contraction, this sea-level history approximates net land-level (subsidence and geoid) change, principally from glacio-isostatic adjustment. Historic rates of rise commenced at 1850–1890 CE and it is virtually certain ( P = 0.99) that the average rate of 20th century RSL rise in northeastern Florida was faster than during any of the preceding 26 centuries. The linearity of RSL rise in Florida is in contrast to the variability reconstructed at sites further north on the U.S. Atlantic coast and may suggest a role for ocean dynamic effects in explaining these more variable RSL reconstructions. Comparison of the difference between reconstructed rates of late Holocene RSL rise and historic trends measured by tide gauges indicates that 20th century sea-level trends along the U.S. Atlantic coast were not dominated by the characteristic spatial fingerprint of melting of the Greenland Ice Sheet.

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Comment on "Donders, T.H. 2014. Middle Holocene humidity increase in Florida: climate or sea-level? Quaternary Science Reviews 103:170-174."

Donders (2014) has recently proposed that the climate of Florida became progressively wetter over the past 5000 years in response to a marked strengthening of the El Niño regime. This reconstruction is largely based on a re-analysis of pollen records from regions north of Lake Okeechobee ( Fig. 1 ) using a new set of pollen transfer functions. Donders concluded that a latitudinal gradient in precipitation prevailed across Florida since the mid Holocene, but the overall trend was toward progressively wetter conditions from 5000 cal BP to the present. Donders (2014) also proposed that this climatic trend extended across South Florida despite contrary paleo-records from the Everglades. In particular he singled out the Northeast Shark River Slough (NESRS) record of Glaser et al. (2013) as an atypical local signal of paleo-environmental change that was biased by a misinterpretation of the ecology of pine and Amaranthaceae (Amaranth family). In response to this direct critique of our paleo-environmental interpretation, we wish to point out that: Our interpretation of the NESRS sedimentary sequence (site 4, Fig. 1 A) was based on multiple lines of evidence that all indicate a shift from wetter to drier (i.e. less wet) conditions that occurred after 2800 cal BP. A similar climatic shift from wetter to less wet conditions was reported for this time interval from other sites in the Everglades ( Willard et al., 2006 and Willard and Bernhardt, 2011 ) and also from the Caribbean region to the east ( Hodell et al., 1991 , Hodell et al., 1995 and Higuera-Gundy et al., 1999 ). The NESRS site is located in an area where runoff collected from a 10,000 km 2 wetland prior to 1900 AD when most of the drainage from the Everglades was channeled to the sea through the narrow Shark River Slough. The sedimentary sequence at the NESRS site should therefore contain an integrated record of hydrological change across the pre-historical Everglades ( Glaser et al., 2012 ). In addition this essentially non-forested wetland is well suited to accumulate a representative sample of the regional pollen rain in its sediments (e.g. Jaccobson and Bradshaw, 1981 and Prentice, 1985 ).

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Bahamas connection: residence areas selected by breeding female loggerheads tagged in Dry Tortugas National Park, USA

Background Delineation of home ranges, residence and foraging areas, and migration corridors is important for understanding the habitat needs for a given species. Recently, many population segments of Northwest Atlantic loggerhead sea turtles ( Caretta caretta ) were designated as endangered or threatened; the smallest subpopulation is in the Dry Tortugas. Foraging and residence areas for this subpopulation have not been defined outside the Gulf of Mexico. Here, for Dry Tortugas loggerheads that traveled to the Bahamas, we use a combination of switching state-space modeling (SSM) and home-range estimators to determine migration period, spatially delineate and describe residence areas, and examine inter-annual home-range repeatability. Results In 5,973 tracking days, migration dates for Dry Tortugas loggerheads traveling to the Bahamas occurred during July–September, with turtles tracked twice showing remarkably similar migration paths and timing of departure from nesting sites. Core-use residence areas for 19 loggerheads ranged from 3.7 to 179.5 km 2 (mean ± 1 SD = 56.2 ± 49.5 km 2 ). For three turtles, we found inter-annual home-range repeatability, with centroids of core areas only 0.7–2.9 km apart and significant overlap of inter-annual 50% kernel contours. Conclusions We demonstrate a previously unknown link between Dry Tortugas nesting beaches and Bahamas residence areas; 17/39 (43.6%) of nesting loggerheads tagged in and tracked from the Dry Tortugas take up residence at sites in the Bahamas. Residence area estimates for these turtles were similar in size to previous foraging area estimates for two turtles tracked to the Bahamas in other studies. We show inter-annual residence area repeatability, and that residence areas of different individuals generally did not overlap. We suggest that these loggerheads possibly establish territories.

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Brumation of introduced Black and White Tegus, Tupinambis merianae (Squamata: Teiidae), in southern Florida

An established population of Tupinambis merianae (Black and White Tegu) in southeastern Florida threatens the Everglades ecosystem. Understanding the behavioral ecology of Black and White Tegus could aid in management and control plans. Black and White Tegus are seasonally active and brumate during the winter in their native range, but brumation behavior is largely unstudied in either the native or the invasive range. We describe the first observations of Black and White Tegu brumation in southeastern Florida after monitoring 5 free-ranging, adult male Black and White Tegus through an inactive season using radiotelemetry and automated cameras. Duration of brumation averaged 137 days, beginning in September and ending by February. One of the 5 Black and White Tegus emerged to bask regularly during brumation, which to our knowledge represents the first documented instance of a free-ranging Black and White Tegu basking during brumation. These preliminary findings provide a basis for future research of brumation behavior.

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The quality of our Nation's waters: Water quality in the Upper Floridan aquifer and overlying surficial aquifers, southeastern United States, 1993-2010

About 10 million people rely on groundwater from the Upper Floridan and surficial aquifers for drinking water. The Upper Floridan aquifer also is of primary importance to the region as a source of water for irrigation and as a source of crystal clear water that discharges to springs and streams providing recreational and tourist destinations and unique aquatic habitats. The reliance of the region on the Upper Floridan aquifer for drinking water and for the tourism and agricultural economies highlights the importance of long-term management to sustain the availability and quality of these resources.

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Depth-dependent groundwater quality sampling at City of Tallahassee test well 32, Leon County, Florida, 2013

Public-supply wells sometimes produce water of less than desirable quality because contaminants can migrate to the open interval of wells through preferential pathways. If these pathways can be identified, zones that produce poor quality water can be excluded during the well-construction process. The U.S. Geological Survey has developed geophysical testing methods that can be used to delineate zones of high permeability in test wells. Once the highly permeable zones are identified, water-quality data can be collected from each zone to identify whether any of the zones produce water of poor quality. The zones producing poor quality water can then be cased off in the final well design so that they do not contribute flow to the production well, reducing subsequent water-treatment costs. A test well was drilled by the City of Tallahassee to assess the suitability of the site for the installation of a new well for public water supply. The test well is in Leon County in north-central Florida. The U.S. Geological Survey delineated high-permeability zones in the Upper Floridan aquifer, using borehole-geophysical data collected from the open interval of the test well. A composite water sample was collected from the open interval during high-flow conditions, and three discrete water samples were collected from specified depth intervals within the test well during low-flow conditions. Water-quality, source tracer, and age-dating results indicate that the open interval of the test well produces water of consistently high quality throughout its length. The cavernous nature of the open interval makes it likely that the highly permeable zones are interconnected in the aquifer by secondary porosity features.

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Changes in the saltwater interface corresponding to the installation of a seepage barrier near Lake Okeechobee, Florida

In 2011, the U.S. Geological Survey and the U.S. Army Corps of Engineers began monitoring the saltwater interface near Lake Okeechobee to evaluate changes in interface depth that could possibly be related to the repair of the Herbert Hoover Dike. A seepage barrier (or cut-off wall), installed by the U.S. Army Corps of Engineers, is a wall of grout designed to protect the Herbert Hoover Dike from internal erosion caused by the piping of water. The seepage barrier prevents water from flowing through or immediately under the dike by diverting the flow below the dike, into the surficial aquifer system. The seepage barrier extends below the saltwater interface in some areas. Monitoring consisted of collecting water samples and time series electromagnetic-induction log (TSEMIL) datasets from 10 well clusters, each of which have 1 shallow and 1 deep monitoring well, with 5- to 10-foot- (ft) long-screened intervals. The deep wells are 120 to 187 ft deep, and the shallow wells are 44 to 100 ft deep. Changes in the depth of the saltwater interface were identified that correspond closely to the depth of the bottom of the seepage barrier. These changes may have been the consequence of changes in groundwater flow initiated by the seepage barrier installation. In areas of the dike where a seepage barrier had not been installed, or where the bottom of the seepage barrier is well above the saltwater interface, monitoring detected no changes in the depth of the saltwater interface. At five of the monitoring-well cluster locations, a long-screened well was also installed for monitoring and comparison purposes. These long-screened wells are 160 to 200 ft deep, and have open intervals ranging from 145 to 185 ft in length. Water samples were collected at depth intervals of about 5 to 10 ft, using 3-ft-long straddle packers to isolate each sampling interval. The results of monitoring conducted using these long-screened interval wells were generally too variable to identify any changes that might be associated with the seepage barrier. Samples from one of these long-screened interval wells failed to detect the saltwater interface evident in samples and TSEMIL datasets from a collocated well cluster. This failure may have been caused by downward flow of freshwater from above the saltwater interface in the well bore.

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Field observations of artificial sand and oil agglomerates

Oil that comes into the surf zone following spills, such as occurred during the 2010 Deepwater Horizon ( DWH ) blowout, can mix with local sediment to form heavier-than-water sand and oil agglomerates ( SOAs ), at times in the form of mats a few centimeters thick and tens of meters long. Smaller agglomerates that form in situ or pieces that break off of larger mats, sometimes referred to as surface residual balls ( SRBs ), range in size from sand-sized grains to patty-shaped pieces several centimeters ( cm ) in diameter. These mobile SOAs can cause beach oiling for extended periods following the spill, on the scale of years as in the case of DWH . Limited research, including a prior effort by the U.S. Geological Survey ( USGS ) investigating SOA mobility, alongshore transport, and seafloor interaction using numerical model output, focused on the physical dynamics of SOAs . To address this data gap, we constructed artificial sand and oil agglomerates ( aSOAs ) with sand and paraffin wax to mimic the size and density of genuine SOAs . These aSOAs were deployed in the nearshore off the coast of St. Petersburg, Florida, during a field experiment to investigate their movement and seafloor interaction. This report presents the methodology for constructing aSOAs and describes the field experiment. Data acquired during the field campaign, including videos and images of aSOA movement in the nearshore (1.5-meter and 0.5-meter water depth) and in the swash zone, are also presented in this report.

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The Everglades Depth Estimation Network (EDEN) surface-water model, version 2

The Everglades Depth Estimation Network (EDEN) is an integrated network of water-level gages, interpolation models that generate daily water-level and water-depth data, and applications that compute derived hydrologic data across the freshwater part of the greater Everglades landscape. The U.S. Geological Survey Greater Everglades Priority Ecosystems Science provides support for EDEN in order for EDEN to provide quality-assured monitoring data for the U.S. Army Corps of Engineers Comprehensive Everglades Restoration Plan. The EDEN surface-water model, version 2 (V2), interpolates water-level data from a network of 240 gages to generate gridded daily water-level surfaces for the freshwater domain of the Everglades. When these spatiotemporal continuous surfaces are combined with EDEN’s digital elevation model of ground surface, derived hydrologic data provide scientists and water managers working in the Everglades with data necessary to analyze ecological and biotic responses to hydrologic changes in the Everglades. Derived datasets include water depth, recession rates, days since last dry, water-surface slopes, and hydroperiod. The V2 model includes enhancements from the previous model (version 1; V1) to accommodate changes in the water-level gage network, adjustments to water-level data, improved understanding of the flow dynamics (particularly near canals), and installation of an elevation benchmark network. Enhancements to the V2 model included Expansion of the EDEN domain: The model domain was expanded to include a part of southern Big Cypress National Preserve and northwestern Everglades National Park upstream of the marsh mangrove wetlands, thus completing the coastal connection along the southwestern boundary of the model; and Development of subdomain models: To account for insufficient water-control structure gage data at some subbasin boundaries, subdomain models were developed for five subdomains, and the resulting water-level surfaces were merged to generate the final water-level surface. Model performance statistics show a general improvement in the V2 model as compared to the V1 model. Overall, the root mean squared error (RMSE) was reduced by 2.42 centimeters (cm) to 4.68 cm. In Water Conservation Area 3A North and Water Conservation Area 3B, the RMSE was reduced by 10.88 and 9.15 cm, respectively. In addition to evaluating model performance statistics, 2-cm water-level maps were generated and evaluated for irregular contours that would indicate a potential problem either with data input or water-level estimates. Three applications of the EDEN-modeled water surfaces and other EDEN datasets are presented in the report to show how scientists and resource managers are using EDEN datasets to analyze biological and ecological responses to hydrologic changes in the Everglades. The biological responses of two important Everglades species, alligators and wading birds, to changes in hydrology are described. The effects of hydrology on fire dynamics in the Everglades are also discussed.

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Hydrologic record extension of water-level data in the Everglades Depth Estimation Network (EDEN), 1991-99

The real-time Everglades Depth Estimation Network (EDEN) has been established to support a variety of scientific and water management purposes. The expansiveness of the Everglades, limited number of gaging stations, and extreme sensitivity of the ecosystem to small changes in water depth have created a need for accurate water-level and water-depth maps. The EDEN water-surface elevation model uses data from approximately 240 gages in the Everglades to create daily continuous interpolations of the water-surface elevation and water depth for the freshwater portion of the Everglades from 2000 to the present (2014). These maps provide hydrologic data previously unavailable for assessing biological and ecological studies. Ecologists working in the Everglades expressed a need to the EDEN project team for daily EDEN water-level surfaces from 1990 to 1999. The additional 10 years of surfaces will provide ecologists and resource managers with two decades (1991–2011) of surfaces to analyze hydrologic dynamics. Before 2000, many of the EDEN gages used to generate water surfaces were not in operation. These datasets were extended to provide estimations of hydrologic time-series histories. The general approach to the record extension (hindcasts) was to (1) create a database of available data from 1990 to the present; (2) use dynamic cluster analysis to group stations with similar hydrologic behaviors for subareas of the Everglades with a large number of stations; (3) use results from the cluster analysis to select candidate explanatory variables; (4) develop linear regression or artificial neural network models to extend water-level records; and (5) evaluate record extensions by using model performance statistics and comparison of water-surface maps for similar hydrologic conditions for the hindcasted period (1991–99) and measured period (2000–11). To hindcast and fill data records, 214 empirical models were developed—189 are linear regression models and 25 are artificial neural network models. The coefficient of determination (R 2 ) for 163 of the models is greater than 0.80 and the median percent model error (root mean square error divided by the range of the measured data) is 5 percent. To evaluate the performance of the hindcast models as a group, contour maps of modeled water-level surfaces at 2-centimeter (cm) intervals were generated using the hindcasted data. The 2-cm contour maps were examined for selected days to verify that water surfaces from the EDEN model are consistent with the input data. The biweekly 2-cm contour maps did show a higher number of issues during days in 1990 as compared to days after 1990. May 1990 had the lowest water levels in the Everglades of the 21-year dataset used for the hindcasting study. To hindcast these record low conditions in 1990, many of the hindcast models would require large extrapolations beyond the range of the predictive quality of the models. For these reasons, it was decided to limit the hindcasted data to the period January 1, 1991, to December 31, 1999. Overall, the hindcasted and gap-filled data are assumed to provide reasonable estimates of station-specific water-level data for an extended historical period to inform research and natural resource management in the Everglades.

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Seismic-sequence stratigraphy and geologic structure of the Floridan aquifer system near "Boulder Zone" deep wells in Miami-Dade County, Florida

The U.S. Geological Survey, in cooperation with the Miami-Dade Water and Sewer Department, acquired, processed, and interpreted seismic-reflection data near the North and South District “Boulder Zone” Well Fields to determine if geologic factors may contribute to the upward migration of injected effluent into that upper part of the Floridan aquifer system designated by the U.S. Environmental Protection Agency as an underground source of drinking water. The depth of the Boulder Zone at the North and South District “Boulder Zone” Well Fields ranges from about 2,750 to 3,300 feet below land surface (ft bls), whereas overlying permeable zones used as alternative drinking water supply range in depth from about 825 to 1,580 ft bls at the North and South District “Boulder Zone” Well Fields. Seismic-sequence stratigraphy and geologic structures imaged on seismic-reflection profiles created for the study describe the part of the Floridan aquifer system overlying and within the Boulder Zone. Features of the Floridan aquifer system underlying the Boulder Zone were not studied because seismic-reflection profiles acquired near the North and South District “Boulder Zone” Well Fields lacked adequate resolution at such depths. Stratigraphic analysis of seismic-reflection data collected from the study area was mainly applied to the Floridan aquifer system and used to identify four provisional seismic sequences, which extend vertically from near the base of the Floridan aquifer system upward to the lower part of the intermediate confining unit. These four seismic sequences compose a framework in which each sequence includes a major permeable unit of the Floridan aquifer system; from shallowest to deepest, these units are the Upper Floridan aquifer, Avon Park permeable zone, uppermost major permeable zone of the Lower Floridan aquifer, and Boulder Zone. The relations between seismic-sequence stratigraphy and hydrostratigraphy allow for detailed mapping of permeable zones and semiconfining units of the Floridan aquifer system at a level of resolution never before accomplished using well data alone. In addition to the preceding seismic-reflection analysis, interpretation of geophysical well log data from four effluent injection wells at the North District “Boulder Zone” Well Field delineated a narrow karst collapse structure beneath the injection facility that extends upward about 900 ft from the top of the Boulder Zone to about 125 ft above the top of the uppermost major permeable zone of the Lower Floridan aquifer. No karst collapse structures were identified in the seismic-reflection profiles acquired near the North District “Boulder Zone” Well Field. However, karst collapse structures at the level of the lowermost major permeable zone of the Lower Floridan aquifer at the South District “Boulder Zone” Well Field are present at three locations, as indicated by seismic-reflection data acquired in the C–1 Canal bordering the south side of the injection facility. Results from the North District “Boulder Zone” Well Field well data indicate that a plausible hydraulic connection between faults and stratiform permeability zones may contribute to the upward transport of effluent, terminating above the base of the deepest U.S. Environmental Protection Agency designated underground source of drinking water at the North District “Boulder Zone” Well Field.

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Spread of the Rat Lungworm (Angiostrongylus cantonensis) in Giant African Land Snails (Lissachatina fulica) in Florida, USA

The rat lungworm ( Angiostrongylus cantonensis ) is a parasitic nematode that causes rat lungworm disease. It is the leading cause of eosinophilic meningitis and is a zoonotic health risk. We confirmed the presence of A. cantonensis using species-specific, quantitative PCR in 18 of 50 (36%) giant African land snails ( Lissachatina fulica ) collected from Miami, Florida in May 2013. These snails were collected from seven of 21 core areas that the Florida Department of Agriculture and Consumer Services monitor weekly. Rat lungworms have not previously been identified in these areas. Duplicate DNA extractions of foot muscle tissue from each snail were tested. Of the seven core areas we examined, six were positive for A. cantonensis and prevalence of infection ranged from 27% to 100%. Of the 18 positive snails, only five were positive in both extractions. Our results confirm an increase in the range and prevalence of rat lungworm infection in Miami. We also emphasize the importance of extracting sufficient host tissue to minimize false negatives.

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Defining conservation targets on a landscape-scale

Conservation planning, the process of deciding how to protect, conserve, enhance and(or) minimize loss of natural and cultural resources, is a fundamental process to achieve conservation success in a time of rapid environmental change. Conservation targets, the measurable expressions of desired resource conditions, are an important tool in biological planning to achieve effective outcomes. Conservation targets provide a focus for planning, design, conservation action, and collaborative monitoring of environmental trends to guide landscape-scale conservation to improve the quality and quantity of key ecological and cultural resources. It is essential to have an iterative and inclusive method to define conservation targets that is replicable and allows for the evaluation of the effectiveness of conservation targets over time. In this document, we describe a process that can be implemented to achieve landscape-scale conservation, which includes defining conservation targets. We also describe what has been accomplished to date (September 2015) through this process for the Peninsular Florida Landscape Conservation Cooperative (PFLCC).

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Breeding loggerhead marine turtles Caretta caretta in Dry Tortugas National Park, USA, show high fidelity to diverse habitats near nesting beaches

We used satellite telemetry to identify in-water habitat used by individuals in the smallest North-west Atlantic subpopulation of adult nesting loggerhead turtles Caretta caretta during the breeding season. During 2010, 2011 and 2012 breeding periods, a total of 20 adult females used habitats proximal to nesting beaches with various levels of protection within Dry Tortugas National Park. We then used a rapid, high-resolution, digital imaging system to map habitat adjacent to nesting beaches, revealing the diversity and distribution of available benthic cover. Turtle behaviour showing measurable site-fidelity to these diverse habitats has implications for managing protected areas and human activities within them. Protecting diverse benthic areas adjacent to loggerhead turtle nesting beaches here and elsewhere could provide benefits for overall biodiversity conservation.

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Genetic structure of Florida green turtle rookeries as indicated by mitochondrial DNA control region sequences

Green turtle ( Chelonia mydas ) nesting has increased dramatically in Florida over the past two decades, ranking the Florida nesting aggregation among the largest in the Greater Caribbean region. Individual beaches that comprise several hundred kilometers of Florida’s east coast and Keys support tens to thousands of nests annually. These beaches encompass natural to highly developed habitats, and the degree of demographic partitioning among rookeries was previously unresolved. We characterized the genetic structure of ten Florida rookeries from Cape Canaveral to the Dry Tortugas through analysis of 817 base pair mitochondrial DNA ( mtDNA ) control region sequences from 485 nesting turtles. Two common haplotypes, CM-A1.1 and CM-A3.1, accounted for 87 % of samples, and the haplotype frequencies were strongly partitioned by latitude along Florida’s Atlantic coast. Most genetic structure occurred between rookeries on either side of an apparent genetic break in the vicinity of the St. Lucie Inlet that separates Hutchinson Island and Jupiter Island, representing the finest scale at which mtDNA structure has been documented in marine turtle rookeries. Florida and Caribbean scale analyses of population structure support recognition of at least two management units: central eastern Florida and southern Florida. More thorough sampling and deeper sequencing are necessary to better characterize connectivity among Florida green turtle rookeries as well as between the Florida nesting aggregation and others in the Greater Caribbean region.

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Predicting spatial and temporal distribution of Indo-Pacific lionfish ( Pterois volitans ) in Biscayne Bay through habitat suitability modeling

Invasive species may exhibit higher levels of growth and reproduction when environmental conditions are most suitable, and thus their effects on native fauna may be intensified. Understanding potential impacts of these species, especially in the nascent stages of a biological invasion, requires critical information concerning spatial and temporal distributions of habitat suitability. Using empirically supported environmental variables (e.g., temperature, salinity, dissolved oxygen, rugosity, and benthic substrate), our models predicted habitat suitability for the invasive lionfish ( Pterois volitans ) in Biscayne Bay, Florida. The use of Geographic Information Systems (GIS) as a platform for the modeling process allowed us to quantify correlations between temporal (seasonal) fluctuations in the above variables and the spatial distribution of five discrete habitat quality classes, whose ranges are supported by statistical deviations from the apparent best conditions described in prior studies. Analysis of the resulting models revealed little fluctuation in spatial extent of the five habitat classes on a monthly basis. Class 5, which represented the area with environmental variables closest to the best conditions for lionfish, occupied approximately one-third of Biscayne Bay, with subsequent habitats declining in area. A key finding from this study was that habitat suitability increased eastward from the coastline, where higher quality habitats were adjacent to the Atlantic Ocean and displayed marine levels of ambient water quality. Corroboration of the models with sightings from the USGS-NAS database appeared to support our findings by nesting 79 % of values within habitat class 5; however, field testing (i.e., lionfish surveys) is necessary to confirm the relationship between habitat classes and lionfish distribution.

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