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Geology topics

Jon E. Keeley

Publications and source records attributed to Jon E. Keeley.

At least 37 records · Page 2Linked to original sources
USGS2016

Exotic annual Bromus invasions: Comparisons among species and ecoregions in the western United States

Exotic annual Bromus species are widely recognized for their potential to invade, dominate, and alter the structure and function of ecosystems. In this chapter, we summarize the invasion potential, ecosystem threats, and management strategies for different Bromus species within each of five ecoregions of the western United States. We characterize invasion potential and threats in terms of ecosystem resistance to Bromus invasion and ecosystem resilience to disturbance with an emphasis on the importance of fire regimes. We also explain how soil temperature and moisture regimes can be linked to patterns of resistance and resilience and provide a conceptual framework that can be used to evaluate the relative potential for invasion and ecological impact of the dominant exotic annual Bromus species in the western United States.

Book chapter↗
USGS2016

Resprouting and seeding hypotheses: A test of the gap-dependent model using resprouting and obligate seeding subspecies of Arctostaphylos

Ecological factors favoring either postfire resprouting or postfire obligate seeding in plants have received considerable attention recently. Three ecological models have been proposed to explain patterns of these two life history types. In this study, we test these three models using data from California chaparral. We take an innovative approach to testing these models by not testing community or landscape patterns, but instead, investigating vegetation structure characteristic of four pairs of resprouting and (non-resprouting) obligate seeding subspecies of Arctostaphylos (Ericaceae), a dominant and diverse shrub genus in California chaparral. Data were analyzed for percentage bare ground, elevation, annual precipitation, number of fires, and time between fires and were compared independently for each subspecies pair. Results were consistently supportive of the gap-dependent model suggesting that obligate seeders are favored when post-disturbance gaps are large. Results were inconclusive or contrary to expectations for both of the other two models.

Plant Ecology↗
USGS2016

Conservation issues: California chaparral

California chaparral, a sclerophyllous shrub-dominated plant community shaped by a Mediterranean-type climate and infrequent, high-intensity fire, is one of the most biodiverse and threatened habitats on Earth. Distinct forms of chaparral, distinguished by differing species composition, geography, and edaphic characteristics, can cover thousands of hectares with dense vegetation or be restricted to smaller communities identified by the presence of endemic species. To maintain the biodiversity of chaparral, protective land management actions will be required to mitigate the loss due to the impacts of human population growth, development, climate change, and increased fire frequencies.

Book chapter↗
USGS2016

Historical reconstructions of California wildfires vary by data source

Historical data are essential for understanding how fire activity responds to different drivers. It is important that the source of data is commensurate with the spatial and temporal scale of the question addressed, but fire history databases are derived from different sources with different restrictions. In California, a frequently used fire history dataset is the State of California Fire and Resource Assessment Program (FRAP) fire history database, which circumscribes fire perimeters at a relatively fine scale. It includes large fires on both state and federal lands but only covers fires that were mapped or had other spatially explicit data. A different database is the state and federal governments’ annual reports of all fires. They are more complete than the FRAP database but are only spatially explicit to the level of county (California Department of Forestry and Fire Protection – Cal Fire) or forest (United States Forest Service – USFS). We found substantial differences between the FRAP database and the annual summaries, with the largest and most consistent discrepancy being in fire frequency. The FRAP database missed the majority of fires and is thus a poor indicator of fire frequency or indicators of ignition sources. The FRAP database is also deficient in area burned, especially before 1950. Even in contemporary records, the huge number of smaller fires not included in the FRAP database account for substantial cumulative differences in area burned. Wildfires in California account for nearly half of the western United States fire suppression budget. Therefore, the conclusions about data discrepancies and the implications for fire research are of broad importance.

California↗
USGS2016

Dispersal limitation does not control high elevational distribution of alien plant species in the southern Sierra Nevada, California

Patterns of elevational distribution of alien plant species in the southern Sierra Nevada of California were used to test the hypothesis that alien plant species invading high elevations around the world are typically climate generalists capable of growing across a wide elevational range. The Sierra Nevada has been heavily impacted for more than a century and a half, first by heavy grazing up into high elevation meadows, followed by major logging, and finally, by impacts associated with recreational use. The comparative elevational patterns of distribution and growth form were compared for native and alien plant species in the four families (Asteraceae, Brassicaceae, Fabaceae, and Poaceae) that contribute the majority of naturalized aliens in the study area. The distribution of realized climatic niche breadth, as measured by elevational range of occurrence, was virtually identical for alien and native species, with both groups showing a roughly Gaussian distribution peaking with species whose range covers a span of 1500–1999 m. In contrast to alien species, which only rarely occurred at higher elevations, native species showed a distribution of upper elevation limits peaking at 3000–3499 m, an elevation that corresponds to the zone of upper montane and subalpine forests. Consistent with a hypothesis of abiotic limitations, only a few alien species have been ecologically successful invaders at subalpine and alpine elevations above 2500 m. The low diversity of aliens able to become established in these habitats is unlikely due to dispersal limitations, given the long history of heavy grazing pressure at high elevations across this region. Instead, this low diversity is hypothesized to be a function of life history traits and multiple abiotic stresses that include extremes of cold air and soil temperature, heavy snowfall, short growing seasons, and low resource availability. These findings have significant implications for resource managers.

California↗
USGS2016

Setting priorities for private land conservation in fire-prone landscapes: Are fire risk reduction and biodiversity conservation competing or compatible objectives?

Although wildfire plays an important role in maintaining biodiversity in many ecosystems, fire management to protect human assets is often carried out by different agencies than those tasked for conserving biodiversity. In fact, fire risk reduction and biodiversity conservation are often viewed as competing objectives. Here we explored the role of management through private land conservation and asked whether we could identify private land acquisition strategies that fulfill the mutual objectives of biodiversity conservation and fire risk reduction, or whether the maximization of one objective comes at a detriment to the other. Using a fixed budget and number of homes slated for development, we simulated 20 years of housing growth under alternative conservation selection strategies, and then projected the mean risk of fires destroying structures and the area and configuration of important habitat types in San Diego County, California, USA. We found clear differences in both fire risk projections and biodiversity impacts based on the way conservation lands are prioritized for selection, but these differences were split between two distinct groupings. If no conservation lands were purchased, or if purchases were prioritized based on cost or likelihood of development, both the projected fire risk and biodiversity impacts were much higher than if conservation lands were purchased in areas with high fire hazard or high species richness. Thus, conserving land focused on either of the two objectives resulted in nearly equivalent mutual benefits for both. These benefits not only resulted from preventing development in sensitive areas, but they were also due to the different housing patterns and arrangements that occurred as development was displaced from those areas. Although biodiversity conflicts may still arise using other fire management strategies, this study shows that mutual objectives can be attained through land-use planning in this region. These results likely generalize to any place where high species richness overlaps with hazardous wildland vegetation.

California↗
USGS2016

Chaparral

One of the most dynamic California ecosystems is chaparral. Dominated by evergreen, sclerophyllous shrubs and small trees, chaparral is the most extensive vegetation type in the state (Figure 1). The nearly impenetrable tangle of stiff branches of this unusual vegetation inhibits exploration, and as a consequence the public know little about its natural history and unique characteristics. This under-valued ecosystem is recognized instead by the threat of its extensive, high-intensity canopy-burning wildfires that characterize the dry summer and fall seasons of the state. Because urban areas frequently share borders or intermix with chaparral, societal interests often conflict with conservation of this ecosystem, and understanding its history and dynamics are a key to appreciating its importance. In this chapter we emphasize the principal structure and dynamics of this important ecosystem. The long summer rainless period has strong impacts on all organisms and, importantly, the droughts influence the fire regime that characterizes chaparral. An ecosystem currently characterized by a specific drought and wildfire regime can expect significant impacts from climate change. Because of its dominance at lower elevations, chaparral also is frequently found at or near the boundaries of urban developments and metropolitan centers. Attempts to suppress fire also affect chaparral dynamics in the long absence of fire. The conflicts between the impacts of chaparral wildfire and human life and structures has been an aspect of California’s history since the beginning, but as development encroaches ever more into chaparral regions, the conflicts have increased. Consequently, understanding this vegetation is important not only because of its significance in understanding ecological evolution and the ecological services provided by chaparral, but also because of its direct impacts on human settlements.

Book chapter↗
USGS2015

Faunal responses to fire in chaparral and sage scrub in California, USA

Impact of fire on California shrublands has been well studied but nearly all of this work has focused on plant communities. Impact on and recovery of the chaparral fauna has received only scattered attention; this paper synthesizes what is known in this regard for the diversity of animal taxa associated with California shrublands and outlines the primary differences between plant and animal responses to fire. We evaluated the primary faunal modes of resisting fire effects in three categories: 1) endogenous survival in a diapause or diapause-like stage, 2) sheltering in place within unburned refugia, or 3) fleeing and recolonizing. Utilizing these patterns in chaparral and sagescrub, as well as some studies on animals in other mediterranean-climate ecosystems, we derived generalizations about how plants and animals differ in their responses to fire impacts and their post fire recovery. One consequence of these differences is that variation in fire behavior has a much greater potential to affect animals than plants. For example, plants recover from fire endogenously from soil-stored seeds and resprouts, so fire size plays a limited role in determining recovery patterns. However, animals that depend on recolonization of burned sites from metapopulations may be greatly affected by fire size. Animal recolonization may also be greatly affected by regional land use patterns that affect colonization corridors, whereas such regional factors play a minimal role in plant community recovery. Fire characteristics such as rate of spread and fire intensity do not appear to play an important role in determining patterns of chaparral and sage scrub plant recovery after fire. However, these fire behavior characteristics may have a profound role in determining survivorship of some animal populations as slow-moving, smoldering combustion may limit survivorship of animals in burrows, whereas fast-moving, high intensity fires may affect survivorship of animals in above ground refugia or those attempting to flee. Thus, fire regime characteristics may have a much greater effect on postfire recovery of animal communities than plant communities in these shrubland ecosystems.

Fire Ecology↗
USGS2015

Fire activity as a function of fire–weather seasonal severity and antecedent climate across spatial scales in southern Europe and Pacific western USA

Climate has a strong influence on fire activity, varying across time and space. We analyzed the relationships between fire–weather conditions during the main fire season and antecedent water-balance conditions and fires in two Mediterranean-type regions with contrasted management histories: five southern countries of the European Union (EUMED)(all fires); the Pacific western coast of the USA (California and Oregon, PWUSA)(national forest fires). Total number of fires (≥1 ha), number of large fires (≥100 ha) and area burned were related to mean seasonal fire weather index (FWI), number of days over the 90th percentile of the FWI, and to the standardized precipitation-evapotranspiration index (SPEI) from the preceding 3 (spring) or 8 (autumn through spring) months. Calculations were made at three spatial aggregations in each area, and models related first-difference (year-to-year change) of fires and FWI/climate variables to minimize autocorrelation. An increase in mean seasonal FWI resulted in increases in the three fire variables across spatial scales in both regions. SPEI contributed little to explain fires, with few exceptions. Negative water-balance (dry) conditions from autumn through spring (SPEI8) were generally more important than positive conditions (moist) in spring (SPEI3), both of which contributed positively to fires. The R2 of the models generally improved with increasing area of aggregation. For total number of fires and area burned, the R2 of the models tended to decrease with increasing mean seasonal FWI. Thus, fires were more susceptible to change with climate variability in areas with less amenable conditions for fires (lower FWI) than in areas with higher mean FWI values. The relationships were similar in both regions, albeit weaker in PWUSA, probably due to the wider latitudinal gradient covered in PWUSA than in EUMED. The large variance explained by some of the models indicates that large-scale seasonal forecast could help anticipating fire activity in the investigated areas.

California, Oregon↗
USGS2015

Towards understanding resprouting at the global scale

Understanding and predicting plant response to disturbance is of paramount importance in our changing world. Resprouting ability is often considered a simple qualitative trait and used in many ecological studies. Our aim is to show some of the complexities of resprouting while highlighting cautions that need be taken in using resprouting ability to predict vegetation responses across disturbance types and biomes. There are marked differences in resprouting depending on the disturbance type, and fire is often the most severe disturbance because it includes both defoliation and lethal temperatures. In the Mediterranean biome, there are differences in functional strategies to cope with water deficit between resprouters (dehydration avoiders) and nonresprouters (dehydration tolerators); however, there is little research to unambiguously extrapolate these results to other biomes. Furthermore, predictions of vegetation responses to changes in disturbance regimes require consideration not only of resprouting, but also other relevant traits (e.g. seeding, bark thickness) and the different correlations among traits observed in different biomes; models lacking these details would behave poorly at the global scale. Overall, the lessons learned from a given disturbance regime and biome (e.g. crown-fire Mediterranean ecosystems) can guide research in other ecosystems but should not be extrapolated at the global scale.

New Phytologist↗
USGS2015

Attacking invasive grasses

In grasslands fire may play a role in the plant invasion process, both by creating disturbances that potentially favour non-native invasions and as a possible tool for controlling alien invasions. Havill et al. ( Applied Vegetation Science , 18, 2015, this issue) determine how native and non-native species respond to different fire regimes as a first step in understanding the potential control of invasive grasses.

Applied Vegetation Science↗
USGS2015

Location, timing and extent of wildfire vary by cause of ignition

The increasing extent of wildfires has prompted investigation into alternative fire management approaches to complement the traditional strategies of fire suppression and fuels manipulation. Wildfire prevention through ignition reduction is an approach with potential for success, but ignitions result from a variety of causes. If some ignition sources result in higher levels of area burned, then ignition prevention programmes could be optimised to target these distributions in space and time. We investigated the most common ignition causes in two southern California sub-regions, where humans are responsible for more than 95% of all fires, and asked whether these causes exhibited distinct spatial or intra-annual temporal patterns, or resulted in different extents of fire in 10-29-year periods, depending on sub-region. Different ignition causes had distinct spatial patterns and those that burned the most area tended to occur in autumn months. Both the number of fires and area burned varied according to cause of ignition, but the cause of the most numerous fires was not always the cause of the greatest area burned. In both sub-regions, power line ignitions were one of the top two causes of area burned: the other major causes were arson in one sub-region and power equipment in the other. Equipment use also caused the largest number of fires in both sub-regions. These results have important implications for understanding why, where and how ignitions are caused, and in turn, how to develop strategies to prioritise and focus fire prevention efforts. Fire extent has increased tremendously in southern California, and because most fires are caused by humans, ignition reduction offers a potentially powerful management strategy, especially if optimised to reflect the distinct spatial and temporal distributions in different ignition causes.

International Journal of Wildland Fire↗
USGS2015

Different fire-climate relationships on forested and non-forested landscapes in the Sierra Nevada ecoregion

In the California Sierra Nevada region, increased fire activity over the last 50 years has only occurred in the higher-elevation forests on US Forest Service (USFS) lands, and is not characteristic of the lower-elevation grasslands, woodlands and shrublands on state responsibility lands (Cal Fire). Increased fire activity on USFS lands was correlated with warmer and drier springs. Although this is consistent with recent global warming, we found an equally strong relationship between fire activity and climate in the first half of the 20th century. At lower elevations, warmer and drier conditions were not strongly tied to fire activity over the last 90 years, although prior-year precipitation was significant. It is hypothesised that the fire–climate relationship in forests is determined by climatic effects on spring and summer fuel moisture, with hotter and drier springs leading to a longer fire season and more extensive burning. In contrast, future fire activity in the foothills may be more dependent on rainfall patterns and their effect on the herbaceous fuel load. We predict spring and summer warming will have a significant impact on future fire regimes, primarily in higher-elevation forests. Lower elevation ecosystems are likely to be affected as much by global changes that directly involve land-use patterns as by climate change.

California↗
USGS2015

Effect of mastication and other mechanical treatments on fuel structure in chaparral

Mechanical fuel treatments are a common pre-fire strategy for reducing wildfire hazard that alters fuel structure by converting live canopy fuels to a compacted layer of dead surface fuels. Current knowledge concerning their effectiveness, however, comes primarily from forest-dominated ecosystems. Our objectives were to quantify and compare changes in shrub-dominated chaparral following crushing, mastication, re-mastication and mastication-plus-burning treatments, and to assess treatment longevity. Results from analysis of variance (ANOVA) identified significant differences in all fuel components by treatment type, vegetation type and time since treatment. Live woody fuel components of height, cover and mass were positively correlated with time since treatment, whereas downed woody fuel components were negatively correlated. Herbaceous fuels, conversely, were not correlated, and exhibited a 5-fold increase in cover across treatment types in comparison to controls. Average live woody fuel recovery was 50% across all treatment and vegetation types. Differences in recovery between time-since-treatment years 1–8 ranged from 32–65% and exhibited significant positive correlations with time since treatment. These results suggest that treatment effectiveness is short term due to the rapid regrowth of shrubs in these systems and is compromised by the substantial increase in herbaceous fuels. Consequences of not having a full understanding of these treatments are serious and leave concern for their widespread use on chaparral-dominated landscapes.

International Journal of Wildland Fire↗
USGS2014

Influence of fuels, weather and the built environment on the exposure of property to wildfire

Wildfires can pose a significant risk to people and property. Billions of dollars are spent investing in fire management actions in an attempt to reduce the risk of loss. One of the key areas where money is spent is through fuel treatment – either fuel reduction (prescribed fire) or fuel removal (fuel breaks). Individual treatments can influence fire size and the maximum distance travelled from the ignition and presumably risk, but few studies have examined the landscape level effectiveness of these treatments. Here we use a Bayesian Network model to examine the relative influence of the built and natural environment, weather, fuel and fuel treatments in determining the risk posed from wildfire to the wildland-urban interface. Fire size and distance travelled was influenced most strongly by weather, with exposure to fires most sensitive to changes in the built environment and fire parameters. Natural environment variables and fuel load all had minor influences on fire size, distance travelled and exposure of assets. These results suggest that management of fuels provided minimal reductions in risk to assets and adequate planning of the changes in the built environment to cope with the expansion of human populations is going to be vital for managing risk from fire under future climates.

PLoS ONE↗
USGS2014

Evolutionary ecology of resprouting and seeding in fire-prone ecosystems

There are two broad mechanisms by which plant populations persist under recurrent disturbances: resprouting from surviving tissues, and seedling recruitment. Species can have one of these mechanisms or both. However, a coherent framework explaining the differential evolutionary pressures driving these regeneration mechanisms is lacking. We propose a bottom-up approach in addressing this question that considers the relative survivorship of adults and juveniles in an evolutionary context, based on two assumptions. First, resprouting and seeding can be interpreted by analogy with annual versus perennial life histories; that is, if we consider disturbance cycles to be analogous to annual cycles, then resprouting species are analogous to the perennial life history with iteroparous reproduction, and obligate seeding species that survive disturbances solely through seed banks are analogous to the annual life history with semelparous reproduction. Secondly, changes in the selective regimes differentially modify the survival rates of adults and juveniles and thus the relative costs and benefits of resprouting versus seeding. Our approach provides a framework for understanding temporal and spatial variation in resprouting and seeding under crown-fire regimes. It accounts for patterns of coexistence and environmental changes that contribute to the evolution of seeding from resprouting ancestors.

New Phytologist↗