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James I. Kirkland

Publications and source records attributed to James I. Kirkland.

5 recordsLinked to original sources

New information on the braincase of the North American therizinosaurian (Theropoda, Maniraptora) Falcarius utahensis

Many disarticulated bones from multiple individuals of a primitive therizinosaurian, referred to Falcarius utahensis , were found in the paucispecific Crystal Geyser bonebed in the Lower Cretaceous Cedar Mountain Formation of eastern Utah. To date, more than 2000 specimens from this species have been excavated. Included in this collection are two partial braincases, one of which is designated the holotype. Here we describe the braincase morphology of Falcarius utahensis . These specimens help establish the primitive cranial condition for the Therizinosauria and further substantiate intraspecific and contralateral braincase pneumatic variation in theropods. When combined with new observations on the cranial remains of the therizinosaurid Nothronychus mckinleyi derived from computed tomographic (CT) scans, the braincase morphology of Falcarius clarifies several evolutionary trends within the Therizinosauria and establishes a suite of synapomorphies for the Therizinosauridae. Trends within the clade include increased basicranial pneumatization (the development of a basisphenoid bulla and loss of external subcondylar recesses), anterior deflection of the supraoccipital, and the reduction of points of origin of the craniocervical musculature, associated with the loss of discrete basipterygoid processes, probably due to incorporation of these structures into the expanded hyperpneumatic bone. Finally, CT scans reveal a complete, nearly avian, inner ear with bird-like semicircular canals and a long cochlea indicating broad frequency discrimination.

Utah↗

Dinosaur tracks from the Cedar Mountain Formation (Lower Cretaceous), Arches National Park, Utah

The seventh and largest known dinosaur tracksite from the Cedar Mountain Formation is reported from two important stratigraphic levels in the Ruby Ranch Member within the boundaries of Arches National Park. Previous reports of sites with a few isolated tracks are of limited utility in indicating the fauna represented by track makers. The Arches site reveals evidence of several theropod morphotypes, including a possible match for the coelurosaur Nedcolbertia and an apparently didactyl Utahraptor -like dromeosaurid. Sauropod tracks indicate a wide-gauge morphotype (cf. Brontopodus ). Ornithischian tracks suggest the presence of an iguandontid-like ornithopod and a large ankylosaur. Dinosaur track diversity is high in comparison with other early Cretaceous vertebrate ichnofaunas, and it correlates well with faunal lists derived from skeletal remains, thus providing a convincing census of the known fauna.

Utah↗

Velafrons coahuilensis , a new labeosaurine haddrosaurid (Dinosauria: Ornithopoda) from the Late Campanian Cerro del Pueblo formation, Coahuila, Mexico

A new lambeosaurine hadrosaurid, Velafrons coahuilensis , is described as the first lambeosaurine from the Cerro del Pueblo Formation of Coahuila, Mexico, and the first lambeosaurine genus to be named from North America in more than 70 years. Although the holotype specimen is a juvenile individual—as evidenced by its incomplete crest development and relative size compared to other North American lambeosaurines—ontogeny independent autapomor-phies have been identified including quadrate with narrow quadratojugal notch and a postorbital with well developed, dorsally positioned squamosal process. Additionally, this taxon is unique in that the prefrontal is not dorsally deflected and anteroposteriorly expanded as in other lambeosaurine taxa of its size, but rather retains the frontal-prefrontal “clamp” present in smaller individuals of other taxa. Phylogenetic analysis places Velafrons in a polytomy with numerous other fan-crested lambeosaurines. The crest structure of Velafrons more closely resembles that of Corythosaurus and Hypacrosaurus because it possesses an anteriorly projecting nasal process over the dorsal premaxilla process. Biogeo-graphically, Velafrons is one of three distinct hadrosaurids known from approximately 73.5 Ma—two lambeosaurines and one hadrosaurine—all restricted to the southern region of the Western Interior Basin of North America.

Coahuila↗

Probable relationships between the lower Jurassic crocodilomorph trackways Batrachopus and Selenichus: Evidence and implications based on new finds from the St. George area southwestern Utah

The enigmatic ichnogenus Selenichnus (Hitchcock, 1858) from the Lower Jurassic of Massachusetts and Utah may represent poorly preserved, extramorphological examples of Batrachopus . Selenichnus trackways from the St. George area (Utah) are virtually indistinguishable from the type material described by Hitchcock (1858) and Lull (1953) from Massachusetts. However, Selenichnus type specimens from Massachusetts, as well as the material from Utah, suggest a relationship with Batrachopus : The former track morphology appears to transform into the latter as preservational conditions change. The size and gait indicated by the two ichnogenera are similar. The suggested relationship between the two ichnogenera does not alter the fact that there are distinct differences in the morphology of the type specimens. Thus, pending discovery of more material, formal synonymy of the two ichnogenera is not formally proposed.

Utah↗

A new pterosaur tracksite from the Jurassic Summerville formation, near Ferron, Utah

Pterosaur tracks (cf. Pteraichnus ) from the Summerville Formation of the Ferron area of central Utah add to the growing record of Pteraichnus tracksites in the Late Jurassic Summerville Formation and time-equivalent, or near time-equivalent, deposits. The site is typical in revealing high pterosaur track densities, but low ichnodiversity suggesting congregations or “flocks” of many individuals. Footprint length varies from 2.0 to 7.0 cms. The ratio of well-preserved pes:manus tracks is about 1:3.4. This reflects a bias in favor of preservation of manus tracks due to the greater weight-bearing role of the front limbs, as noted in other pterosaur track assemblages. The sample also reveals a number of well-preserved trackways including one suggestive of pes-only progression that might be associated with take off or landing, and another that shows pronounced lengthening of stride indicating acceleration. One well-preserved medium-sized theropod trackway ( Therangospodus ) and other larger theropod track casts (cf. Megalosauripus ) are associated with what otherwise appears to be a nearly monospecific pterosaur track assemblage. However, traces of a fifth pes digit suggest some tracks are of rhamphorynchoid rather than pterodactyloid origin, as usually inferred for Pteraichnus . The tracks occur at several horizons in a thin stratigraphic interval of ripple marked sandstones and siltstones. Overall the assemblage is similar to others found in the same time interval in the Western Interior from central and eastern Utah through central and southern Wyoming, Colorado, northeastern Arizona, and western Oklahoma. This vast “ Pteraichnus ichnofacies,” with associated saurischian tracks, remains the only ichnological evidence of pre-Cretaceous pterosaurs in North America and sheds important light on the vertebrate ecology of the Summerville Formation and contiguous deposits.

Utah↗