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Geology topics

J.K. Ringelman

Publications and source records attributed to J.K. Ringelman.

13 recordsLinked to original sources

Declines of greater and lesser scaup populations: issues, hypotheses, and research needs

The population estimate for greater (Aythya marila) and lesser (Aythya affinis) scaup (combined) has declined dramatically since the early 1980s to record lows in 1998. The 1998 estimate of 3.47 million scaup is far below the goal of 6.3 million set in the North American Waterfowl Management Plan (NAWMP), causing concern among biologists and hunters. We summarize issues of concern, hypotheses for factors contributing to the population decline, and research and management needs recommended by participants of the Scaup Workshop, held in September 1999. We believe that contaminants, lower female survival, and reduced recruitment due to changes in food resources or breeding-ground habitats are primary factors contributing to the decline. These factors are not mutually exclusive but likely interact across seasons. Workshop participants identified seven action items. We need to further delineate where declines in breeding populations have occurred, with a primary focus on the western Canadian boreal forest, where declines appear to be most pronounced. Productivity in various areas and habitats throughout the breeding range needs to be assessed by conducting retrospective analyses of existing data and by intensive field studies at broad and local scales. Annual and seasonal survival rates need to be determined in order to assess the role of harvest or natural mortality. Effects of contaminants on reproduction, female body condition, and behavior must be investigated. Use, distribution, and role of food resources relative to body condition and reproduction need to be examined to better understand seasonal dynamics of nutrient reserves and the role in reproductive success. Affiliations among breeding, migration, and wintering areas must be assessed in order to understand differential exposure to harvest or contaminants, and differential reproductive success and recruitment. Biologists and agencies need to gather and improve information needed to manage greater and lesser scaup separately; this includes monitoring the breeding populations of each species separately, closer examination of existing data to improve surveys and data collection, and re-evaluation of the NAWMP population goal. These complex issues will require extensive cooperation and communication among many agencies and organizations in North America.

Wildlife Society Bulletin

Characteristics of some black duck nest sites in Maine

A standard method for characterizing nest sites and concealment (visibility of orange decoy and percent overhead cover measured by densiometer) was used to obtain characteristics of 36 nest sites of black ducks in Maine, 1978-89. Nest locations were represented by cutover areas (10), islands (6), bogmat (5), emergent meadow (5), emergent wetland (3), stream floodplain (3), hardwood forest (1), conifer forest (1), mixed forest (1) and ephemeral pond (1). Within these locations nests were found in shrub clumps (1), under conifers (6), on hummocks (6), on ericaceous mats (4), under a clump of hardwood trees (4), under woody slash (3), on an emergent herbaceous clump (1), on a boulder (1) and on a muskrat house (1). After excluding 7 nests disturbed by investigators, 22 (76%) of the 29 remaining nests were successful nests. Nests in upland cuts were especially successful (9 of 9) and success was 75 - 100% at most locations, but both nests along stream floodplain were abandoned because of human disturbance. Unsuccessful nests were usually closer to ponds (2.5 times) or streams (6.6 times) and often at land water interfaces, i.e., islands and bogmat. Nests under conifers (5 of 6) and woody slash (3 of 3) usually were successful. The combination of low nesting density and isolation of nests in upland cutover areas (successful nests averaged nearly 3 times farther from roads) seem to influence black duck nest success.

Book chapter

Densities of breeding American black ducks in southcentral Maine: 1958-60 and 1978-80

Numbers of wetlands and amount of surface water (ha) within a 105 km2 area increased substantially between 1958-60 and 1978-80 in southcentral Maine. Conversely, in 1978-80 mean numbers of black duck pairs (11.9/100 ha surface water) and broods (10.8/100 ha surface water) were lower than for 1958-60 (pairs = 23.3; broods = 13.3) for wetlands surveyed in both periods. Differences in pairs are discussed in terms of wetland changes and in terms of differences in numbers of black ducks counted in the Atlantic Flyway mid-winter inventory for comparable years.

Book chapter

Use of Wetland Habitats by Selected Nongame Water Birds in Maine

We examined the use of 87 palustrine and lacustrine wetlands by nongame water birds in central and eastern Maine using 3,527 h of observation (1,501 visits) made during April-August, 1977-85. Wetlands used by 15 species of water birds were distinguished from those not used, according to 20 habitat features. The species were the common loon (Gavia immer) , pied-billed grebe (Podilymbus podiceps), double-crested cormorant (Phalacrocorax auritus), American bittern (Botaurus lentiginosus), great blue heron (Ardea herodias), green-backed heron (Butorides striatus), osprey (Pandion haliaetus), bald eagle (Haliaeetus leucocephalus), northern harrier (Circus cyaneus), Virgima rail (Rallus limicola), sora (Porzana carolina), spotted sandpiper (Actitis macularia), common snipe (Gallinago gallinago), herring gull (Larus argentatus), and belted kingfisher (Ceryle alcyon). Predictive models of habitat use were developed for each species. Water birds were classified by similarity of habitats used, and species use was contrasted by wetland type. Smaller, isolated wetlands were used by fewer (P < 0.05) species than larger wetlands in complexes; many species had large area-requirements (pied-billed grebe, common loon, herring gull, double-crested cormorant, bald eagle) or preferred to use wetlands near other wetlands (common loon, herring gull, great blue heron, spotted sandpiper, osprey, bald eagle). Wetland area contributed more to overall variation in species richness on wetlands than wetland isolation, although on small wetlands (<3.6 ha) isolation was a better predictor of species richness than wetland area. Wetlands with intermediate amounts (33-66%) of emergent vegetation supported more species (P< 0.05) than closed (>66%) or open (<33%) wetlands. Low pH typified wetlands used by large-bodied piscivores (common loon, cormorant, osprey). Other water birds were associated with more densely vegetated, chemically buffered wetlands. Habitat features associated with wetland use by each waterbird species are reported, as are numerical responses of waterbird populations to wetland features and estimates of annual variation in habitat occupancy. Lacustrine wetlands supported a distinct, low diversity community of water birds, including most fish-eating species. Waterbird diversity at forested palustrine wetlands was intermediate between lacustrine communities and more species-rich assemblages at palustrine emergent and scrub-shrub wetlands. Regional variation in wetland characteristics and water bird use was associated with surficial geology, soils, and management practices. Management for nongame water birds in Maine should consider providing emergent and aquatic-bed vegetation with variable cover-to-water ratios, accommodating species-specific habitat needs, focusing on species of restricted distribution and low abundance, and maintaining wetland complexes. Bird use and habitat information from 87 wetlands and models of habitat selection for each species are provided in appendixes.

Fish and Wildlife Research

Population ecology and harvest of the American black duck: a review

1. The purpose of our review was to examine available data on population trends and current status of black ducks and trends in natality and survival and to relate these, where possible, to changes in habitat, predation, disease, contaminants, harvest, and hybridization with mallards. 2. The number of black ducks tallied in the winter survey has declined steadily over the past 30 years at an average rate of about 3%/ year. Reliability and precision of the survey are uncertain; it may not provide an adequate index to the continental population of black ducks. Breeding surveys are incomplete and sporadic, but black ducks have decreased in Ontario and increased in the Maritime Provinces and Quebec. 3. Recent declines in numbers of black ducks tallied in the winter survey are not unusual in magnitude or much different from those that have occurred among several other species of waterfowl. 4. At present, black ducks are not especially scarce relative to numbers of several other ducks in eastern North America. 5. There is no solid evidence of major decreases in quality or quantity of breeding habitat for black ducks in recent years; in some areas, habitat has improved. 6. Natural mortality of black ducks has not been well studied, but does not seem unusually high compared to other dabbling ducks. 7. Harvest rates of black ducks are similar to those of sympatric mallards as determined by banding analyses. 8. There is no strong evidence for direct effects of contaminants on black ducks, but some indirect effects through invertebrate food resources have been detected. 9. Age ratios in black ducks show no trend in the past 18 years. 10. The quality and quantity of wintering habitat for black ducks have decreased substantially in some areas. 11. Disease and other natural mortality that affect black ducks do .not occur in unusually high frequency. 12. A decline in harvest of black ducks has occurred; most of the decline has been in the United States, especially since restrictive regulations were implemented in 1983. 13. Recovery rates of black ducks have declined recently in the U.S., but not in Canada. 14. Survival rates of black ducks are 56-63% for adults and 43% for young. These rates of survival are similar to comparable estimates in sympatric mallards. 15. Long hunting seasons may depress survival in some sex-age classes of black ducks, buteffects of small reductions in survival on population trends are unknown. 16. Available evidence does not support the contention that hunting is either the sole or most important cause of the decline in the winter tally of black duck numbers. 17. Surveys and banding of black ducks should be thoroughly reviewed and maintained or improved as warranted. Obtaining or maintaining a reasonable index to numbers of black ducks is the top priority among survey needs. 18. Experimental manipulation of hunting seasons should be considered to elucidate relationships among regulations, harvest, survival, and population trends. 19. Black ducks and mallards are genetically similar; there is as much genetic differentiation within the 2 species as there is between them. 20. Black duck x mallard hybrids are fertile. Hybrids are difficult to detect by plumage and thus published frequencies (0-13%) of hybrids may be low. 21. Hybridization could be a result of concomitant mallard increases and black duck decreases, or changes in black duck-to-mallard ratios could be from hybridization and genetic swamping of black ducks.

Wildlife Society Bulletin

Survival of female black ducks, Anas rubripes, during the breeding season

The Mayfield method was used to estimate the survival rate of 19 radio-marked, female Black Ducks (Anas rubripes) in southcentral Maine during 1977-80. An overall survival rate of 0.74 was estimated for the 121-day monitoring period that included the pre-laying and laying, incubation, brood rearing, and post-rearing stages. No differences in survival rates were detected among these stages. Two instrumented hens were killed by Red-shouldered Hawks (Buteo lineatus) and a third was killed by an unknown predator. We found no evidence that the attachment of radio transmitters affected hen survival.

Canadian Field-Naturalist

Breeding habitat selection and home range of radio-marked black ducks (Anas rubripes) in Maine

Telemetry techniques were used to monitor the movements and habitat use of 13 female and 7 male black ducks ( Anas rubripes ) in an inland breeding region of south central Maine in 1977–1980. Black ducks preferred persistent emergent, broad-leaved deciduous forested, and broad-leaved deciduous scrub–shrub wetlands over unconsolidated organic bottom, needle-leaved evergreen forested, and broad-leaved evergreen scrub–shrub ponds. Birds also made frequent use of small ephemeral pools and streams throughout the breeding period. Nests were located in several habitats ranging from wetland sites to upland areas 1.5 km from the most frequently used pond. Home range size averaged 119 ha for females and 231 ha for males and did not differ by reproductive stage. Three pairs used only a single pond during the incubation period. Home ranges were linear (linearity index = 2.8), averaging 1956 m long for females and 2755 m for males. Wetlands used most by hens during incubation recesses were not always those located closest to the nest. Radio-marked ducks that returned in subsequent breeding seasons demonstrated fidelity to the previously used home range. Pair bonds of marked birds lasted until day 19 or 20 of incubation for initial nesting attempts.

Maine

Nest and brood attentiveness in female black ducks

Incubation rhythms and brood attentiveness of radio-marked Black Ducks (Anas rubripes) were studied in southcentral Maine during 1977-1980. Recess duration and frequency differed between three females nesting near wetlands (x = 82 min, 2.3 recesses/day) and two nesting at upland sites(x = 183 min, I. I recesses/day), but incubation constancy was similar for all birds (x = 86.7%). A fourth wetland-nesting female apparently responded to the absence of down and concealing cover at the nest site by taking shorter and fewer recesses (x= 34 min, 1.0 recesses/day) than did other wetland-nesters with typical nests. Wetland-nesters took longer recesses with increasing air temperature and following long incubation sessions. After their ducklings were two weeks old, two brood-rearing females began leaving their broods to forage on nearby wetlands. Duration of rearing recess (x = 56 min) and total recess time (x = 94 min/day) were less during this mid-rearing stage than during the latter part of rearing (x = 265 min, 488 min/day). The two hen-brood bonds terminated at 43 and 48 days. We suggest that small endogenous nutrient reserves and the low density of invertebrate foods, typical of Black Duck breeding habitat, were critical factors affecting the evolution of Black Duck incubation behavior. Bouts of inattention during brood rearing may have also evolved in relation to food requirements; by foraging on wetlands away from their broods, females avoid competing with offspring for common food resources.

Condor

Computer simulation models as tools for identifying research needs: A black duck population model

Existing data on the mortality and production rates of the black duck (Anas rubripes) were used to construct a WATFIV computer simulation model. The yearly cycle was divided into 8 phases: hunting, wintering, reproductive, molt, post-molt, and juvenile dispersal mortality, and production from original and renesting attempts. The program computes population changes for sex and age classes during each phase. After completion of a standard simulation run with all variable default values in effect, a sensitivity analysis was conducted by changing each of 50 input variables, 1 at a time, to assess the responsiveness of the model to changes in each variable. Thirteen variables resulted in a substantial change in population level. Adult mortality factors were important during hunting and wintering phases. All production and mortality associated with original nesting attempts were sensitive, as was juvenile dispersal mortality. By identifying those factors which invoke the greatest population change, and providing an indication of the accuracy required in estimating these factors, the model helps to identify those variables which would be most profitable topics for future research.

Transactions of the Northeast Section of the Wildl

Factors affecting waterfowl breeding density and productivity estimates in the Northeast

During 1977-79, information useful for making breeding pair and brood surveys was obtained while studying black duck (Anas rubripes) habitat selection and productivity in south-central Maine. Surveys should be initiated in relation to sunrise and sunset time. Morning versus evening counts, familiarity with the survey area, wetland dynamics of the study area, wetland surface water area, and allotment of relative survey effort are discussed as they affect the conduct and results of brood surveys.

Transactions of the Northeast Section of the Wildl