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Chad Raymond Kluender

Publications and source records attributed to Chad Raymond Kluender.

7 recordsLinked to original sources

Reliability of satellite-based vegetation maps for planning wildfire-fuel treatments in shrub steppe: Inferences from two contrasting national parks

Protecting habitat threatened by increasing wildfire size and frequency requires identifying the spatial intersection of wildfire behavior and ecological conditions that favor positive management outcomes. In the perennial sagebrush steppe of Western North America, invasions by fire-prone annual grasses are a key concern, and management of them requires reliable maps of vegetation cover, fuels, and wildfire behavior. We compared commonly used, publicly available vegetation cover and fuels maps, specifically the Rangeland Analysis Platform (RAP) and LANDFIRE, with field-based assessments at two U.S. National Parks dominated by sagebrush steppe: City of Rocks National Reserve and Craters of the Moon National Monument and Preserve. Plant-community composition and fuels measured at ∼1700 field locations spanning ∼300,000 ha revealed that 1) RAP generally underestimated each vegetation cover type where the cover was actually abundant, and conversely overestimated cover types where they were actually scarce, and 2) there was considerable disagreement in fuel-bed maps derived from LANDFIRE compared to field observations. As a result, there were substantial discrepancies in the spatial patterning of wildfire behavior estimated from the fire-spread model FLAMMAP when parameterized with LANDFIRE compared to field-based fuel-bed maps created from Random Forests models. Reliable maps of vegetation cover and fuel conditions are needed to help guide fuels and invasive species management, especially given recent increases in pre- and post-fire treatments in arid and semiarid landscapes. The costs associated with poorly informed fuel reduction may greatly exceed the costs of field-based vegetation and fuels inventory to inform effective design of vegetative fuels treatments.

Idaho

Patchy response of cheatgrass and nontarget vegetation to indaziflam and imazapic applied after wildfire in sagebrush steppe

Control of nonnative grasses is needed where they are altering fire regimes and degrading rangelands, such as cheatgrass ( Bromus tectorum ) invasion of perennial sagebrush-steppe communities. Aerial broadcast of the pre-emergent and postemergent herbicide imazapic has been used for decades over vast areas to control cheatgrass after fire. Recent small-scale studies indicate that the pre-emergent herbicide indaziflam may provide more enduring cheatgrass control. We evaluated landscape-level vegetation responses to indaziflam sprayed in replicated areas at 66.7 g · ai · ha −1 , with and without imazapic (66.1 g · ai · ha −1 ) over almost 500 ha of sagebrush steppe. Herbicides were strip-sprayed by helicopter in the fall of 2019 in subregions that either 1) had burned in the summer of 2019 and had moderate background cheatgrass invasion, 2) had burned in 2011 and became heavily invaded, or 3) were burned in both 2011 and 2019 and had intermediate invasion. Tarps were temporarily deployed to intercept herbicides and create untreated controls. Overall, indaziflam + imazapic had greater initial control of cheatgrass, but by 2023, both treatments led to similar ∼17 percentage-point reductions in cheatgrass cover. Cheatgrass individuals that “escaped” the herbicide treatment grew exceptionally large and fecund. There were no reductions in cover in any native vegetation type, including biocrusts, and nontarget increases in cover were observed for 1) deep-rooted perennial grasses treated with indaziflam + imazapic in the 2011 burn subregion and 2) the shallow-rooted Sandberg bluegrass ( Poa secunda ) treated with either herbicide in the 2011 or 2011 + 2019 burn subregions. Consideration of burn legacies, pretreatment landscape condition, and evenness of treatment application may improve restoration outcomes and help prioritize management allocation, timing, and treatment expectations.

Idaho

Propensity score matching mitigates risk of faulty inferences in observational studies of effectiveness of restoration trials

Determining effectiveness of restoration treatments is an important requirement of adaptive management, but it can be non-trivial where only portions of large and heterogeneous landscapes of concern can be treated and sampled. Bias and non-randomness in the spatial deployment of treatment and thus sampling is nearly unavoidable in the data available for large-scale management trials, and the biophysical landscape characteristics underlying the bias are key but rare considerations in analyses of treatment effects. Treatment effects from large-scale management trials are typically estimated with multivariable regression (MVR) models. However, this method is unsuited to reliable estimations of treatment effects when treated and untreated areas differ in their underlying biophysical variability. An alternative to conventional regression is to use propensity score (PS) matching, which can limit the differences in confounding variables among treatment groups and assure the data collected or selected for analysis are more consistent with a randomized and unconfounded experiment. Thus, PS is expected to identify treatment effects more accurately. We used data from a large-scale monitoring effort of a megafire to evaluate the efficacy of PS matching in making inferences on treatment effects when treatments are applied non-randomly over a large heterogeneous area. We compared the resulting inference to both traditional MVR methods and to “naïve” methods that do not consider treatment allocation bias. Treatment effects varied between the different statistical methods for controlling selection bias and confounding biophysical factors. The PS-matched model revealed a weaker treatment effect of drill seeding and a greater effect of herbicide spraying on the cover of perennial bunchgrasses when compared to MVR or naïve modelled estimates. The inferences from the PS-matched model are considered more reliable because the treated and untreated plots are more similar in their underlying biophysical characteristics. Synthesis and applications . Failure to consider the non-random and selective deployment of restoration treatments by managers leads to faulty inference on their effectiveness. However, tools such as propensity-score matching can be used to remove the bias from analyses of the outcomes of management trials or to devise sampling plans that efficiently protect against the bias.

Journal of Applied Ecology

Multiple plant-community traits improve predictions of later-stage outcomes of restoration drill seedings: Implications for metrics of success

Success of ecological restoration is often only knowable if treatments meet criteria defined by biotic thresholds, but analytical frameworks to determine metrics of success and their underlying thresholds are needed. Early indicators of longer-term recovery trajectories are particularly critical where re-treatments may be required, such as in harsh climates or where repeated disturbances or invasive pressures prevail. We developed a framework for identifying which biotic traits would provide the best initial indication of longer-term target restoration goals and applied the framework to restoration drill-seedings of deep-rooted perennial bunchgrasses (DRPBGs) used to rehabilitate and restore semiarid rangelands threatened by exotic annual grasses (EAGs, e.g. cheatgrass) and the recurrent wildfire that EAGs cause. Initial traits measured included cover, basal diameter, height, and density (#plants/area) of DRPBGs and cover of EAGs and Sandberg bluegrass ( Poa secunda , POSE, a disturbance-adapted perennial). The longer-term target objective was ≥25 % DRPBG cover and ≤13 % EAG cover by the 5th year following drill-seedings. Measurements were made on 112 plots spanning 113,000 ha in sagebrush steppe on the Soda wildfire scar, in the Northern Great Basin, USA. Traits of DRPBGs tended to be uncorrelated with one another, thus each was informative in describing vegetation condition. Where DRPBG cover was initially >17 %, it tended to become >25 % by the 5th-year post-seeding. In plots that overcame an initial risk of not meeting the target objective (i.e. <17 % initial DRPBG cover), DRPBG tended be large DRPBGs (>22.8 cm height) and plots also had >7 % cover of POSE. Additional “sets” of initial vegetation traits were also predictive of longer-term restoration success or failure. Restoration drill-seeding of DRPBGs is a key but varied-outcome tool for breaking the exotic grass-fire cycle, and, contrary to a conventional tendency to rely on a limited number of mean traits such as % cover, a suite of biotic traits appears necessary to monitor to reliably know if trials are likely to yield success.

Ecological Indicators

A nontarget, disturbance-resilient native species influences post-fire recovery and multiphasic herbicide-seeding outcomes in drylands threatened by exotic annual grasses

Native species that are abundant and persistent across disturbance-succession cycles can affect recovery and restoration of plant communities, especially in drylands. In the sagebrush-steppe deserts of North America, restoring deep-rooted perennial bunchgrasses (DRPBGs) is key to the strategy for breaking an increasingly problematic cycle of wildfire promoted by exotic annual grasses (EAGs) and displacement of perennials by post-fire increases in EAGs. We asked how Sandberg bluegrass ( Poa secunda , POSE)—a common native grass that shares traits with EAGs such as resilience to disturbance and rapid, shallow-rooted, early season growth—(1) recovered after wildfire, (2) responded to different combinations of native-plant seedings of DRPBGs and EAG-targeting herbicides; and (3) in turn, related to DRPBG recovery. In repeated sampling of up to approximately 2,000 plots spanning 113,000 burned hectares, POSE was initially more abundant than DRPBGs until POSE began to decline 5 years post-fire, and POSE was marginally affected or unaffected by restoration treatments targeting DRPBGs or EAGs. In comparison, EAG cover decreased 8–16% where preemergent herbicide was sprayed and DRPBG cover increased 3–9% where they were drill-seeded, and the greatest EAG reductions and DRPBG increases were seen where seeding and herbicides were combined in a time-staggered fashion. Treatments had less target effects where POSE cover was high (>18%), which is also where EAG cover was scarcer and DRPBG cover greater, regardless of post-fire interventions. Consideration of the ecological role of disturbance-resilient species with ruderal characteristics that match traits of key invaders may improve the efficiency and effectiveness of restoration interventions.

Restoration Ecology

Plant community trajectories following livestock exclusion for conservation vary and hinge on initial invasion and soil-biocrust conditions in shrub steppe

Adjustments or complete withdrawal of livestock grazing are among the most common conservation actions in semiarid uplands, but outcomes can vary considerably with ecological context. Invasion by exotic annual grasses and the excessive wildfire they promote are increasing threats to semiarid shrub-steppe, and plant-community response to livestock exclusion in these areas may be complicated by the rapid colonization ability of invaders. We evaluated vegetation-community changes over 14-year interval (2007–2021) in a shrub-steppe landscape where a >100-year history of livestock grazing had been terminated in 1996. Field surveys revealed that bare-soil exposure decreased >20% over the 14 years owing to biomass accumulation, but this was primarily due to large increases in exotic annual “cheatgrass” ( Bromus tectorum , +1.8-fold) and the litter it produces (+1.5-fold). Soil biocrusts increased 11.9% and perennial bunchgrasses increased 3% over the 14 years. These community changes varied at the patch scale and entailed inverse relationships of (1) both cheatgrass and biocrusts to plant-community basal cover, (2) cheatgrass to both biocrusts and perennial grasses, and (3) biocrusts to cheatgrass and litter. The spatiotemporal variability in vegetation constituted changes in plant-community states, according to cluster analysis. The modeled probability of a community transitioning to a cheatgrass state was (1) strongly and positively related to the initial (2007) cover of cheatgrass in hotspots where initial cheatgrass cover was >20%, and (2) negatively related to biocrust cover where initial biocrust cover was >4% of ground area. The decision space for this landscape can be framed as a shifting from acceptance towards resisting further degradation by removing livestock and their trampling of soil surfaces and utilization of perennial herbs. However, cheatgrass appears to be the most impactful agent of change and continued invasion appears imminent. Active restoration may help resist further degradation and direct change towards tolerable conditions.

Colorado

Relationship of greater sage-grouse to natural and assisted recovery of key vegetation types following wildfire: Insights from scat

Megafires are creating severe conservation problems worldwide for wildlife that have obligate dependencies on plant species that are foundational but fire-intolerant. Wildfire-induced loss of native perennials and increases in exotic annual grasses threaten greater sage-grouse (GRSG, Centrocercus urophasianus ) in its sagebrush steppe habitat in western North America. Post-fire restoration using herbicides, seeding, and planting of native perennials such as sagebrush are common, but there are few assessments of GRSG response to the treatments. We measured the presence of GRSG scat and modeled the probability of GRSG presence (PrGRSG-scat ) in relation to variation in plot-level and landscape-level predictors, and land treatments, in an intensive, repeat sampling from 2017-2020 of 113,000-ha area burned in 2015 in the Soda Megafire (Oregon and Idaho, USA). GRSG scat was present in <200 of >8000 observations, as would be expected for a philopatric species (i.e., high fidelity to home site) returning to degraded habitat. PrGRSG-scat was positively associated with sagebrush presence at the plot-level and was positively related to elevation, lower-angle slopes, and proximity to sagebrush seedling outplant islands. The statistical significance of relationships of PrGRSG-scat to restoration treatments was marginal at best, with the largest effect being a positive response of PrGRSG-scat to pre-emergent herbicide sprayed to reduce exotic annual grasses. More time may be required for restored sagebrush steppe to meet GRSG needs or for GRSG to “adopt” the restored vegetation. Moreover, whereas scat is a convenient and non-invasive method to monitor GRSG, its post-fire scarcity weakens the strength of statistical inference on GRSG recovery patterns and response to restoration.

Restoration Ecology